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''Myroxylon'' is a genus of
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
native to
Latin America Latin America or * french: Amérique Latine, link=no * ht, Amerik Latin, link=no * pt, América Latina, link=no, name=a, sometimes referred to as LatAm is a large cultural region in the Americas where Romance languages — languages derived f ...
.


History

The first described species in this genus was '' M. balsamum.'' It was originally described in 1753 by
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
as ''Toluifera balsamum'', based on a specimen collected in the province of Cartagena (at the time
Tolú Tolú is a small municipality and town in Sucre Department, northern Colombia by the Caribbean sea. The municipality has an area of 500 km². It is named for the Tolú, one the Pre-Columbian era, pre-Columbian Indigenous peoples of South Amer ...
was located in the province of Cartagena). The genus ''Myroxylon'' was first established by Linnaeus filius in 1781, when he described '' M. peruiferum'' based on a specimen collected by Mutis in South America. Although ''Toluifera'' is prior in term of publication time, ''Myroxylon'' is chosen as the
conserved name A conserved name or ''nomen conservandum'' (plural ''nomina conservanda'', abbreviated as ''nom. cons.'') is a scientific name that has specific nomenclatural protection. That is, the name is retained, even though it violates one or more rules whic ...
and ''Toluifera'' is rejected. The name derives from Greek μύρρα (''myrrha'', "
myrrh Myrrh (; from Semitic, but see '' § Etymology'') is a gum-resin extracted from a number of small, thorny tree species of the genus ''Commiphora''. Myrrh resin has been used throughout history as a perfume, incense and medicine. Myrrh mi ...
") and ξύλον (''xylon'', "
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...
").


Species

Some authors recognize infra-specific taxa based, mainly, in their balsam phytochemistry; while other authors do not recognize such categories. There are reports of differences in composition of balsams obtained from ''M. balsamum'' var. ''balsamum'' (Tolu balsam tree), ''M. balsamum'' var. ''pereirae'' (Peru balsam tree), and ''M. peruiferum'' (quina). It is in the flowering plant family
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
(Leguminosae). There are two species:


Distribution

''Myroxylon'' species grow in
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
(primarily in
El Salvador El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south b ...
) and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
. ''Myroxylon balsamum'' occurs in Central America, and northern and western South America, it is fairly common in tropical forest at 200–690 m altitude. In Peru and Brazil this species is mostly associated with rivers, and sometimes grows on lateritic soil. It is found in remnants of mesophillous forest. At present it is considered as being of least concern (LC) according to CITES classification. ''Myroxylon peruiferum'' is disjunctly distributed in the Americas, from Mexico to northern Argentina and southern Brazil, though it has a wide distribution, it is not abundant within its area of occurrence. It is found in remnants of mesophillous forest and dry habitats at 540–2000 m elevation. It is considered to be Near Threatened (NT), according to CITES classification.


The tree

The trees are large, growing to tall, with
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
long, with 5–13 leaflets. The flowers are white with yellow
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s, produced in
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is a pod long, containing a single
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
. The tree is often called ''Quina'' or ''Balsamo'', ''Tolu'' in
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
, ''Quina quina'' in
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, and sometimes ''Santos Mahogany'' or ''Cabreuva'' in the lumber trade. Members of this genus produce hydroxy
pipecolic acid Pipecolic acid (piperidine-2-carboxylic acid) is an organic compound with the formula HNC5H9CO2H. It is a carboxylic acid derivative of piperidine and, as such, an amino acid, although not one encoded genetically. Like many other α-amino acids, p ...
s in their leaves. The wood is dark brown, with a deep red
heartwood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin tha ...
. Natural oils grant it excellent decay resistance. In fact, it is also resistant to preservative treatment. Its
specific gravity Relative density, or specific gravity, is the ratio of the density (mass of a unit volume) of a substance to the density of a given reference material. Specific gravity for liquids is nearly always measured with respect to water (molecule), wa ...
is 0.74–0.81. With regard to
woodwork Woodworking is the skill of making items from wood, and includes cabinet making (cabinetry and furniture), wood carving, joinery, carpentry, and woodturning. History Along with stone, clay and animal parts, wood was one of the first materials ...
ing, the tree is moderately difficult to work but can be finished with a high natural polish; it tends to cause some tool dulling.


Invasive species

The balsam tree can become a highly invasive species when introduced into tropical countries where it is not native. In
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, it has overgrown several hectares of the
Udawatta Kele Sanctuary Udawattakele Forest Reserve often spelled as Udawatta Kele, is a historic forest reserve on a hill-ridge in the city of Kandy. It is 104 hectares (257 acres) large. During the days of the Kandyan kingdom, Udawattakele was known as "Uda Wasala Watt ...
and is rapidly spreading there. In this
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
n rain forest, ''Myroxylon'' seeds sprout in very high numbers due to tolerating more diverse light conditions than native species and due to the absence of natural enemies such as diseases and insects. This has given rise to dense stands of young trees where no other vegetation can grow, causing severe ecological disruption, ''i.e.'', the disappearance of local, native plant species and consequently of the animals and insects that feed on these. The tree has also been introduced to several Pacific islands such as
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
and to
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, and is a potential ecological threat there.


References


External links

* {{Taxonbar, from=Q589643 Amburaneae Invasive plant species in Sri Lanka Resins Crops originating from the Americas Fabaceae genera