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Moths are a
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
group of insects that includes all members of the order
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
that are not
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
, with moths making up the vast majority of the order. There are thought to be approximately 160,000
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of moth, many of which have yet to be described. Most species of moth are
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
, but there are also
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
and diurnal species.


Differences between butterflies and moths

While the
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
form a
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
group, the moths, comprising the rest of the Lepidoptera, do not. Many attempts have been made to group the superfamilies of the Lepidoptera into natural groups, most of which fail because one of the two groups is not monophyletic:
Microlepidoptera Microlepidoptera (micromoths) is an artificial (i.e., unranked and not monophyletic) grouping of moth families, commonly known as the 'smaller moths' (micro, Lepidoptera). These generally have wingspans of under 20 mm, and are thus harder to ...
and
Macrolepidoptera Macrolepidoptera is a group within the insect order Lepidoptera. Traditionally used for the larger butterflies and moths as opposed to the "microlepidoptera", this group is artificial. However, it seems that by moving some taxa about, a monoph ...
,
Heterocera Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
and
Rhopalocera Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
, Jugatae and Frenatae,
Monotrysia The Monotrysia are a group of moths in the lepidopteran order, not currently considered to be a natural group or clade. Apart from the recently discovered family Andesianidae, most of the group consists of small, relatively understudied species ...
and
Ditrysia The Ditrysia are a natural group or clade of insects in the lepidopteran order containing both butterflies and moths. They are so named because the female has two distinct sexual openings: one for mating, and the other for laying eggs (in contra ...
.Scoble, MJ 1995. The Lepidoptera: Form, function and diversity. Oxford, UK: Oxford University Press; 404 p. Although the rules for distinguishing moths from butterflies are not well established, one very good guiding principle is that butterflies have thin antennae and (with the exception of the family
Hedylidae Hedylidae, the "American moth-butterflies", is a family of insects in the order Lepidoptera, representing the superfamily Hedyloidea. They have traditionally been viewed as an extant sister group of the butterfly superfamily Papilionoidea. In 198 ...
) have small balls or clubs at the end of their antennae. Moth antennae are usually feathery with no ball on the end. The divisions are named by this principle: "club-antennae" (Rhopalocera) or "varied-antennae" (Heterocera). Lepidoptera first evolved during the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
period, but only evolved their characteristic proboscis alongside the rise of angiosperms in the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
period.


Etymology

The modern English word ''moth'' comes from
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
(
cf. The abbreviation ''cf.'' (short for the la, confer/conferatur, both meaning "compare") is used in writing to refer the reader to other material to make a comparison with the topic being discussed. Style guides recommend that ''cf.'' be used onl ...
Northumbrian ) from
Common Germanic Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the reconstructed proto-language of the Germanic branch of the Indo-European languages. Proto-Germanic eventually developed from pre-Proto-Germanic into three Germanic branc ...
(compare
Old Norse Old Norse, Old Nordic, or Old Scandinavian, is a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants of Scandinavia and t ...
,
Dutch Dutch commonly refers to: * Something of, from, or related to the Netherlands * Dutch people () * Dutch language () Dutch may also refer to: Places * Dutch, West Virginia, a community in the United States * Pennsylvania Dutch Country People E ...
, and
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
all meaning 'moth'). Its origins are possibly related to the Old English meaning '
maggot A maggot is the larva of a fly (order Diptera); it is applied in particular to the larvae of Brachycera flies, such as houseflies, cheese flies, and blowflies, rather than larvae of the Nematocera, such as mosquitoes and crane flies. ...
' or from the root of ''
midge A midge is any small fly, including species in several families of non-mosquito Nematoceran Diptera. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid zones. Some mid ...
'' which until the 16th century was used mostly to indicate the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
, usually in reference to devouring clothes.


Caterpillar

Moth larvae, or
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s, make cocoons from which they emerge as fully grown moths with wings. Some moth caterpillars dig holes in the ground, where they live until they are ready to turn into adult moths.


History

Moths evolved long before butterflies; moth
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s have been found that may be 190 million years old. Both types of Lepidoptera are thought to have co-evolved with
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s, mainly because most modern species, both as adults and larvae, feed on flowering plants. One of the earliest known species that is thought to be an ancestor of moths is ''Archaeolepis mane''. Its fossil fragments show scaled wings that are similar to
caddisflies The caddisflies, or order Trichoptera, are a group of insects with aquatic larvae and terrestrial adults. There are approximately 14,500 described species, most of which can be divided into the suborders Integripalpia and Annulipalpia on the b ...
in their veining.


Economics


Significance to humans

Some moths, particularly their
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s, can be major
agricultural Agriculture or farming is the practice of cultivating Plant, plants and livestock. Agriculture was the key development in the rise of Sedentism, sedentary human civilization, whereby farming of Domestication, domesticated species created food ...
pests in many parts of the world. Examples include
corn borer ''Ostrinia'' is a genus of moths in the family Crambidae described by Jacob Hübner in 1825. Several of them, including the European corn borer, are agricultural pests. Species *'' Ostrinia avarialis'' Amsel, 1970 *'' Ostrinia dorsivittata'' (Mo ...
s and
bollworm Bollworm is the common term for a moth larva that attacks the fruiting bodies of certain crops, especially cotton. The most common moths known as bollworms are: * Red or Sudan bollworm, '' Diparopsis castanea'' * Rough bollworm, '' Earias perhuege ...
s. The caterpillar of the
spongy moth ''Lymantria dispar'', also known as the gypsy moth or the spongy moth, is a species of moth in the family Erebidae. ''Lymantria dispar'' is subdivided into several subspecies, with subspecies such as ''L. d. dispar'' and ''L. d. japonica'' bei ...
(''Lymantria dispar'') causes severe damage to forests in the northeastern United States, where it is an
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
. In temperate climates, the
codling moth The codling moth (''Cydia pomonella'') is a member of the Lepidopteran family Tortricidae. They are major pests to agricultural crops, mainly fruits such as apples and pears. Because the larvae are not able to feed on leaves, they are highly d ...
causes extensive damage, especially to fruit farms. In tropical and subtropical climates, the diamondback moth (''
Plutella xylostella The diamondback moth (''Plutella xylostella''), sometimes called the cabbage moth, is a moth species of the family Plutellidae and genus '' Plutella''. The small, grayish-brown moth sometimes has a cream-colored band that forms a diamond along ...
'') is perhaps the most serious pest of brassicaceous crops. Also in
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
, the African sugarcane borer is a major pest of sugarcane,
maize Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. Th ...
, and
sorghum ''Sorghum'' () is a genus of about 25 species of flowering plants in the grass family (Poaceae). Some of these species are grown as cereals for human consumption and some in pastures for animals. One species is grown for grain, while many othe ...
. Several moths in the family
Tineidae Tineidae is a family of moths in the order Lepidoptera described by Pierre André Latreille in 1810. Collectively, they are known as fungus moths or tineid moths. The family contains considerably more than 3,000 species in more than 300 genera. ...
are commonly regarded as pests because their larvae eat
fabric Textile is an umbrella term that includes various fiber-based materials, including fibers, yarns, filaments, threads, different fabric types, etc. At first, the word "textiles" only referred to woven fabrics. However, weaving is not the ...
such as
clothes Clothing (also known as clothes, apparel, and attire) are items worn on the human body, body. Typically, clothing is made of fabrics or textiles, but over time it has included garments made from animal skin and other thin sheets of materials ...
and
blanket A blanket is a swath of soft cloth large enough either to cover or to enfold most of the user's body and thick enough to keep the body warm by trapping radiant body heat that otherwise would be lost through convection. Etymology The term ...
s made from natural
proteinaceous Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
fibers such as
wool Wool is the textile fibre obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have properties similar to animal wool. As ...
or
silk Silk is a natural protein fiber, some forms of which can be woven into textiles. The protein fiber of silk is composed mainly of fibroin and is produced by certain insect larvae to form cocoons. The best-known silk is obtained from the coc ...
.Scott, Thomas (1995)
''Concise Encyclopedia Biology''
. Walter de Gruyter. .
They are less likely to eat mixed materials containing some artificial fibers. There are some reports that they may be repelled by the scent of wood from
juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' () of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere, from the Arcti ...
and
cedar Cedar may refer to: Trees and plants *''Cedrus'', common English name cedar, an Old-World genus of coniferous trees in the plant family Pinaceae *Cedar (plant), a list of trees and plants known as cedar Places United States * Cedar, Arizona * ...
, by
lavender ''Lavandula'' (common name lavender) is a genus of 47 known species of flowering plants in the mint family, Lamiaceae. It is native to the Old World and is found in Cape Verde and the Canary Islands, and from Europe across to northern and easte ...
, or by other natural oils; however, many consider this unlikely to prevent infestation.
Naphthalene Naphthalene is an organic compound with formula . It is the simplest polycyclic aromatic hydrocarbon, and is a white crystalline solid with a characteristic odor that is detectable at concentrations as low as 0.08  ppm by mass. As an aromati ...
(the chemical used in
mothball Mothballs are small balls of chemical pesticide and deodorant, sometimes used when storing clothing and other materials susceptible to damage from mold or moth larvae (especially clothes moths like ''Tineola bisselliella''). Composition Older ...
s) is considered more effective, but there are concerns over its effects on human health. Moth larvae may be killed by freezing the items which they infest for several days at a temperature below . While moths are notorious for eating clothing, most species do not, and some moth adults do not even eat at all. Some, like the
Luna Luna commonly refers to: * Earth's Moon, named "Luna" in Latin * Luna (goddess), the ancient Roman personification of the Moon Luna may also refer to: Places Philippines * Luna, Apayao * Luna, Isabela * Luna, La Union * Luna, San Jose Romania * ...
,
Polyphemus Polyphemus (; grc-gre, Πολύφημος, Polyphēmos, ; la, Polyphēmus ) is the one-eyed giant son of Poseidon and Thoosa in Greek mythology, one of the Cyclopes described in Homer's ''Odyssey''. His name means "abounding in songs and lege ...
,
Atlas An atlas is a collection of maps; it is typically a bundle of maps of Earth or of a region of Earth. Atlases have traditionally been bound into book form, but today many atlases are in multimedia formats. In addition to presenting geographic ...
,
Promethea ''Promethea'' is a comic book series created by Alan Moore, J. H. Williams III and Mick Gray, published by America's Best Comics/WildStorm. It tells the story of Sophie Bangs, a college student from an alternate futuristic New York City in 1999 ...
,
cecropia ''Cecropia'' is a Neotropical genus consisting of 61 recognized species with a highly distinctive lineage of dioecious trees. The genus consists of pioneer trees in the more or less humid parts of the Neotropics, with the majority of the speci ...
, and other large moths do not have mouth parts. This is possible because they live off the food stores from when they were a caterpillar, and only live a short time as an adult (roughly a week for some species). Many species of adult moths do however eat: for instance, many will drink
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
. Some moths are
farm A farm (also called an agricultural holding) is an area of land that is devoted primarily to agricultural processes with the primary objective of producing food and other crops; it is the basic facility in food production. The name is used fo ...
ed for their economic value. The most notable of these is the
silkworm The domestic silk moth (''Bombyx mori''), is an insect from the moth family Bombycidae. It is the closest relative of ''Bombyx mandarina'', the wild silk moth. The silkworm is the larva or caterpillar of a silk moth. It is an economically imp ...
, the larva of the domesticated moth ''
Bombyx mori The domestic silk moth (''Bombyx mori''), is an insect from the moth family Bombycidae. It is the closest relative of ''Bombyx mandarina'', the wild silk moth. The silkworm is the larva or caterpillar of a silk moth. It is an economically imp ...
''. It is farmed for the silk with which it builds its cocoon. , the silk industry produces more than 130 million kilograms of raw silk, worth about 250 million
U.S. dollars The United States dollar (symbol: $; code: USD; also abbreviated US$ or U.S. Dollar, to distinguish it from other dollar-denominated currencies; referred to as the dollar, U.S. dollar, American dollar, or colloquially buck) is the official ...
, each year. Not all silk is produced by ''Bombyx mori''. There are several species of
Saturniidae Saturniidae, commonly known as saturniids, is a family of Lepidoptera with an estimated 2,300 described species. The family contains some of the largest species of moths in the world. Notable members include the emperor moths, royal moths, and gi ...
that also are farmed for their silk, such as the ailanthus moth (''
Samia cynthia ''Samia cynthia'', the ailanthus silkmoth, is a saturniid moth, used to produce silk fabric but not as domesticated as the silkworm, ''Bombyx mori''. The moth has very large wings of , with a quarter-moon shaped spot on both the upper and lower ...
'' group of species), the Chinese oak silkmoth (''
Antheraea pernyi ''Antheraea pernyi'', the Chinese (oak) tussar moth, Chinese tasar moth or temperate tussar moth, is a large moth in the family Saturniidae. The species was first described by Félix Édouard Guérin-Méneville in 1855. '' Antheraea roylei'' is a ...
''), the Assam silkmoth (''
Antheraea assamensis ''Antheraea assamensis'', known as the muga silkworm as a larva and Assam silk moth as an adult, is a moth of the family Saturniidae. The species was first described by Johann Wilhelm Helfer in 1837. It is found in Assam in northeast India where 9 ...
''), and the Japanese silk moth (''
Antheraea yamamai ''Antheraea yamamai'', the Japanese silk moth or Japanese oak silkmoth (Japanese: or ) is a moth of the family Saturniidae. It is endemic to east Asia, but has been imported to Europe for tussar silk production and is now found in southeastern Eu ...
''). The larvae of many species are used as food, particularly in Africa, where they are an important source of nutrition. The mopane worm, the caterpillar of ''
Gonimbrasia belina ''Gonimbrasia belina'' is a species of emperor moth which is native to the warmer parts of southern Africa. Its large edible caterpillar, known as the mopane worm, madora, amacimbi or masontja, feeds primarily but not exclusively on mopane tree ...
'', from the family Saturniidae, is a significant food resource in
southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
. Another saturniid used as food is the cavorting emperor ('' Usta terpsichore''). In one country alone,
Congo Congo or The Congo may refer to either of two countries that border the Congo River in central Africa: * Democratic Republic of the Congo, the larger country to the southeast, capital Kinshasa, formerly known as Zaire, sometimes referred to a ...
, more than 30 species of moth larvae are harvested. Some are sold not only in the local village markets, but are shipped by the ton from one country to another.


Predators and parasites

Nocturnal insectivores often feed on moths; these include some
bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most bi ...
s, some species of
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s and other species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s. Moths also are eaten by some species of
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s,
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
s,
cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
s,
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Do ...
s,
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s, and some
bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Nor ...
s. Moth larvae are vulnerable to being parasitized by
Ichneumonidae The Ichneumonidae, also known as the ichneumon wasps, Darwin wasps, or ichneumonids, are a family (biology), family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 2 ...
. Baculoviruses are parasite double-stranded DNA
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
viruses that are used mostly as
biological control Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also invo ...
agents. They are members of the
Baculoviridae ''Baculoviridae'' is a family of viruses. Arthropods, among the most studied being Lepidoptera, Hymenoptera and Diptera, serve as natural hosts. Currently, 85 species are placed in this family, assigned to four genera. Baculoviruses are known ...
, a family that is restricted to insects. Most baculovirus isolates have been obtained from insects, in particular from Lepidoptera. There is evidence that ultrasound in the range emitted by bats causes flying moths to make evasive maneuvers.
Ultrasonic Ultrasound is sound waves with frequencies higher than the upper audible limit of human hearing. Ultrasound is not different from "normal" (audible) sound in its physical properties, except that humans cannot hear it. This limit varies fr ...
frequencies trigger a reflex action in the
noctuid The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family of moths. They are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly changing, along with the other f ...
moth that causes it to drop a few centimeters or inches in its flight to evade attack, and tiger moths can emit clicks to foil bats' echolocation. The fungus ''
Ophiocordyceps sinensis ''Ophiocordyceps sinensis'' (formerly known as ''Cordyceps sinensis''), known colloquially as caterpillar fungus, is an entomopathogenic fungus (a fungus that grows on insects) in the family Ophiocordycipitaceae. It is mainly found in the mead ...
'' infects the larvae of many different species of moths.


Ecological importance

Moths, like butterflies, bees and other more popularly recognized
pollinating Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
insects, serve an essential role as pollinators for many flowering plants, including species that bees do not visit. Nocturnal moths fly from flower to flower to feed on nectar during the night much as their diurnal relatives do during the day. A study conducted in the UK found moths dusted with pollen from 47 different plant species, including seven species largely ignored by bees. Some studies indicate that certain species of moths, such as those belonging to the families
Erebidae The Erebidae are a family of moths in the superfamily Noctuoidea. The family is among the largest families of moths by species count and contains a wide variety of well-known macromoth groups. The family includes the underwings ('' Catocala'') ...
and
Sphingidae The Sphingidae are a family of moths (Lepidoptera) called sphinx moths, also colloquially known as hawk moths, with many of their caterpillars known as “hornworms”; it includes about 1,450 species. It is best represented in the tropics, bu ...
, may be the key pollinators for some flowering plants in the Himalayan ecosystem. The roles of moths as pollinators have been studied less frequently than those of diurnal pollinators, but recent studies have established that moths are important, but often overlooked, nocturnal pollinators of a wide range of plants. Some researchers say it is likely that many plants thought to be dependent on bees for pollination also rely on moths, which have historically been less observed because they pollinate mainly at night.


Attraction to light

Moths frequently appear to circle artificial lights, although the reason for this behavior (positive
phototaxis Phototaxis is a kind of taxis, or locomotory movement, that occurs when a whole organism moves towards or away from a stimulus of light. This is advantageous for phototrophic organisms as they can orient themselves most efficiently to receive li ...
) is currently unknown. One hypothesis is called celestial or transverse orientation. By maintaining a constant angular relationship to a bright celestial light, such as the moon, they can fly in a straight line. Celestial objects are so far away that, even after travelling great distances, the change in angle between the moth and the light source is negligible; further, the moon will always be in the upper part of the visual field, or on the horizon. When a moth encounters a much closer artificial light and uses it for navigation, the angle changes noticeably after only a short distance, in addition to being often below the horizon. The moth instinctively attempts to correct by turning toward the light, thereby causing airborne moths to come plummeting downward, and resulting in a spiral flight path that gets closer and closer to the light source. Studies have found that
light pollution Light pollution is the presence of unwanted, inappropriate, or excessive use of artificial Visible spectrum, lighting. In a descriptive sense, the term ''light pollution'' refers to the effects of any poorly implemented lighting, during the day ...
caused by increasing use of artificial lights has either led to a severe decline in moth population in some parts of the world or has severely disrupted nocturnal pollination.


Noteworthy moths

*Atlas moth (''
Attacus atlas ''Attacus atlas'', the Atlas moth, is a large Saturniidae, saturniid moth endemic to the forests of Asia. The species was first described by Carl Linnaeus in his 1758 10th edition of Systema Naturae, 10th edition of ''Systema Naturae''. Descri ...
''), one of the largest moths in the world *Herculese moth (''
Coscinocera hercules ''Coscinocera hercules'', the Hercules moth, is a moth of the family Saturniidae, endemic to New Guinea and northern Australia. The species was first described by William Henry Miskin in 1876. Description Adults have a wingspan of about , makin ...
''), largest moth in Australia *White witch moth (''
Thysania agrippina ''Thysania agrippina'' is a species of moth in the family Erebidae. It was described by Maria Sibylla Merian in her 1705 publication ''Metamorphosis insectorum Surinamensium,'' and Pieter Cramer provided the Species description, formal descripti ...
''), the Lepidopteran with the longest wingspan * Madagascan sunset moth (''Chrysiridia rhipheus''), considered to be one of the most impressive and beautiful Lepidoptera *
Death's-head hawkmoth The name death's-head hawkmoth refers to any of three moth species of the genus ''Acherontia'' (''Acherontia atropos'', '' Acherontia styx'' and '' Acherontia lachesis''). The former species is found in Europe and throughout Africa, the latter ...
(''Acherontia'' spp.), is associated with the supernatural and evil and has been featured in art and movies *
Peppered moth The peppered moth (''Biston betularia'') is a temperate species of night-flying moth. It is mostly found in the northern hemisphere in places like Asia, Europe and North America. Peppered moth evolution is an example of population genetics an ...
(''Biston betularia''), the subject of a well-known study in natural selection *Luna moth (''
Actias luna The Luna moth (''Actias luna'') also known as the American moon moth is a Nearctic moth in the family Saturniidae, subfamily Saturniinae, a group commonly known as giant silk moths. It has lime-green colored wings and a white body. The larvae ( ...
'') *
Grease moth ''Aglossa cuprina'', the grease moth, is a snout moth, family Pyralidae, described by Philipp Christoph Zeller in 1872. The grease moth is closely related to the genus '' Pyralis'', and as a result, is usually associated with the meal moth, ''P ...
(''Aglossa cuprina''), known to have fed on the rendered fat of humansBrundage, Adrienne (23 March 2009), Other Arthropods of Forensic Importance, Texas A&M University, Texas A&M University Forensic Entomology Lecture *
Emperor gum moth ''Opodiphthera eucalypti'', the emperor gum moth, is a species of moth in the family Saturniidae native to Australia. This species was formerly placed in the genus ''Antheraea''. Distribution The emperor gum moth may inhabit all states of Austr ...
(''Opodiphthera eucalypti'') *
Polyphemus moth ''Antheraea polyphemus'', the Polyphemus moth, is a North American member of the family Saturniidae, the giant silk moths. It is a tan-colored moth, with an average wingspan of 15 cm (6 in). The most notable feature of the moth is its ...
(''Antheraea polyphemus'') *
Bogong moth The bogong moth (''Agrotis infusa'') is a temperate species of night-flying moth, notable for its biannual long-distance seasonal migrations towards and from the Australian Alps, similar to the diurnal monarch butterfly. During the autumn a ...
(''Agrotis infusa''), known to have been a food source for southeastern
indigenous Australians Indigenous Australians or Australian First Nations are people with familial heritage from, and membership in, the ethnic groups that lived in Australia before British colonisation. They consist of two distinct groups: the Aboriginal peoples ...
*
Ornate moth ''Utetheisa ornatrix'', also called the bella moth, ornate moth or rattlebox moth is a moth of the subfamily Arctiinae. It is aposematically colored ranging from pink, red, orange and yellow to white coloration with black markings arranged in va ...
(''Utetheisa ornatrix''), the subject of numerous behavioral studies regarding sexual selection


Moths of economic significance

*Spongy moth (''
Lymantria dispar ''Lymantria dispar'', also known as the gypsy moth or the spongy moth, is a species of moth in the family Erebidae. ''Lymantria dispar'' is subdivided into several subspecies, with subspecies such as ''L. d. dispar'' and ''L. d. japonica'' bei ...
''), an invasive species pest of hardwood trees in North America *
Winter moth :''In North America, "winter moth" usually denotes the invasive species ''Operophtera brumata'', but may also mean refer to a native species, ''Erannis tiliaria'' (linden looper) or '' Operophtera bruceata'' (bruce spanworm).'' The winter moth (' ...
(''Operophtera brumata''), an invasive species pest of hardwood trees, cranberry and blueberry in northeastern North America * Corn earworm or cotton bollworm (''Helicoverpa zea''), a major agricultural pest *
Indianmeal moth The Indianmeal moth (''Plodia interpunctella''), also spelled Indian meal moth and Indian-meal moth, is a pyraloid moth of the family Pyralidae. Alternative common names are weevil moth, pantry moth, flour moth or grain moth. The almond moth ...
(''Plodia interpunctella''), a major pest of grain and flour *
Codling moth The codling moth (''Cydia pomonella'') is a member of the Lepidopteran family Tortricidae. They are major pests to agricultural crops, mainly fruits such as apples and pears. Because the larvae are not able to feed on leaves, they are highly d ...
(''Cydia pomonella''), a pest mostly of apple, pear and walnut trees *
Light brown apple moth The light brown apple moth (''Epiphyas postvittana'') (often abbreviated to LBAM) is a leafroller moth belonging to the lepidopteran family Tortricidae. Identification Adult moths Light brown apple moth adults are variable in colour and may be ...
(''Epiphyas postvittana''), a highly
polyphagous Feeding is the process by which organisms, typically animals, obtain food. Terminology often uses either the suffixes -vore, -vory, or -vorous from Latin ''vorare'', meaning "to devour", or -phage, -phagy, or -phagous from Greek φαγε ...
pest *Silkworm (''
Bombyx mori The domestic silk moth (''Bombyx mori''), is an insect from the moth family Bombycidae. It is the closest relative of ''Bombyx mandarina'', the wild silk moth. The silkworm is the larva or caterpillar of a silk moth. It is an economically imp ...
''), for its silk *
Wax moth Waxworms are the caterpillar larvae of wax moths, which belong to the family Pyralidae (snout moths). Two closely related species are commercially bred – the lesser wax moth (''Achroia grisella'') and the greater wax moth (''Galleria m ...
s (''
Galleria mellonella ''Galleria mellonella'', the greater wax moth or honeycomb moth, is a moth of the family Pyralidae. ''G. mellonella'' is found throughout the world. It is one of two species of wax moths, with the other being the lesser wax moth. ''G. mellonella' ...
'', ''
Achroia grisella The lesser wax moth (''Achroia grisella'') is a small moth of the snout moth family (Pyralidae) that belongs to the subfamily Galleriinae. The species was first described by Johan Christian Fabricius in 1794. Adults are about 0.5 inches (13  ...
''), pests of bee hives *''
Duponchelia fovealis ''Duponchelia fovealis'' is a species of moth of the family Crambidae described by Philipp Christoph Zeller in 1847. It is endemic to the area surrounding the Mediterranean Sea, and the Canary Islands, but has extended its range to other parts of ...
'', a new invasive pest of vegetables and ornamental plants in the United States


Gallery

File:Micrographia Schem 30.jpg, Diagram of a
plume moth The Pterophoridae or plume moths are a family of Lepidoptera with unusually modified wings. Though they belong to the Apoditrysia like the larger moths and the butterflies, unlike these they are tiny and were formerly included among the assemblag ...
from
Robert Hooke Robert Hooke FRS (; 18 July 16353 March 1703) was an English polymath active as a scientist, natural philosopher and architect, who is credited to be one of two scientists to discover microorganisms in 1665 using a compound microscope that ...
's ''
Micrographia ''Micrographia: or Some Physiological Descriptions of Minute Bodies Made by Magnifying Glasses. With Observations and Inquiries Thereupon.'' is a historically significant book by Robert Hooke about his observations through various lenses. It w ...
'' File:Kerala Leaf Insect.jpg, Leaf-shaped moth (''
Pergesa acteus ''Pergesa'' is a monotypic moth genus in the family Sphingidae first described by Francis Walker in 1856. Its only species, ''Pergesa acteus'', the green pergesa hawkmoth, was described by Pieter Cramer in 1779. Distribution It is found in S ...
'') File:Giant grey moth.png, Giant grey moth (''
Agrius convolvuli ''Agrius convolvuli'', the convolvulus hawk-moth, is a large hawk-moth. It is common throughout Europe, Asia, Africa, Australia and New Zealand, partly as a migrant. In New Zealand, it is also known as the kumara moth, and in the Māori language ...
'') File:Colourful Moth.jpg, Oleander hawk-moth or army green moth (''
Daphnis nerii ''Daphnis nerii'', the oleander hawk-moth or army green moth, is a moth of the family Sphingidae. It was described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. Distribution ''Daphnis nerii'' is a large hawk-moth found in wi ...
'') File:Red spotted moths 1 (3745889925).jpg, Six-spot burnet moths mating (''
Zygaena filipendulae The six-spot burnet (''Zygaena filipendulae'') is a day-flying moth of the family Zygaenidae. Subspecies *''Z. f. altapyrenaica'' Le Charles, 1950 *''Z. f. arctica'' Schneider, 1880 *''Z. f. balcanirosea'' Holik, 1943 *''Z. f. campaniae'' Re ...
'') File:Silk cocoon.jpg, Protective silk (or similar material) case (cocoon) File:Caterpillar-----02.jpg, A caterpillar of
death's-head hawkmoth The name death's-head hawkmoth refers to any of three moth species of the genus ''Acherontia'' (''Acherontia atropos'', '' Acherontia styx'' and '' Acherontia lachesis''). The former species is found in Europe and throughout Africa, the latter ...
File:Joined moths.JPG, Mating pair of ''
Laothoe populi ''Laothoe populi'', the poplar hawk-moth, is a moth of the family Sphingidae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. It is found throughout the Palearctic region and the Near East and ...
'', or poplar hawkmoths, showing two different color variants File:White-lined sphinx moth.JPG,
White-lined sphinx moth ''Hyles lineata'', also known as the white-lined sphinx, is a moth of the family Sphingidae. They are sometimes known as the hummingbird moth because of their bird-like size (2-3 inch wingspan) and flight patterns. As caterpillars, they have a ...
in Colorado, United States File:Мебельная моль.jpg, Closeup of a common clothes moth File:Giant silk moth (Adelowalkeria tristygma).jpg, Giant silk moth (''
Adelowalkeria tristygma ''Adelowalkeria'' is a genus of moths in the family Saturniidae first described by Travassos in 1941. Species *''Adelowalkeria caeca'' Lemaire, 1969 *''Adelowalkeria eugenia'' (Druce, 1904) — Ecuador *''Adelowalkeria flavosignata'' (Walker, 18 ...
'') File:Adult Emperor Moth.jpg, Adult emperor moth (''
Gonimbrasia belina ''Gonimbrasia belina'' is a species of emperor moth which is native to the warmer parts of southern Africa. Its large edible caterpillar, known as the mopane worm, madora, amacimbi or masontja, feeds primarily but not exclusively on mopane tree ...
'') File:A Moth on marble floor.jpg, A moth on a marble floor in
Kolkata Kolkata (, or , ; also known as Calcutta , the official name until 2001) is the capital of the Indian state of West Bengal, on the eastern bank of the Hooghly River west of the border with Bangladesh. It is the primary business, comme ...
, India File:Clothes moth.jpg, Clothes moth, eye File:Female rose-myrtle lappet moth.jpg, Female
rose-myrtle lappet moth ''Trabala vishnou'', the rose-myrtle lappet moth, is a moth of the family Lasiocampidae. It is found in southern Asia, including Pakistan, India, Bangladesh, Thailand, Sri Lanka, Myanmar, Java, China, Japan, Taiwan, Hong Kong, Vietnam and Indone ...
hanging on the wooden door


See also

*
Baculovirus ''Baculoviridae'' is a family of viruses. Arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropod ...
*
Clothing moth Clothes moth or clothing moth is the common name for several species of moth considered to be pests, whose larvae eat animal fibres (hairs), including clothing and other fabrics. These include: * ''Tineola bisselliella'', the common clothes moth ...
*
Comparison of butterflies and moths A common classification of the Lepidoptera involves their differentiation into butterflies and moths. Butterflies are a natural monophyletic group, often given the suborder Rhopalocera, which includes Papilionoidea (true butterflies), Hesperiid ...
*
List of moths This article is a list of lists of some of the 160,000 species of Lepidoptera that are commonly known as moths. By region *Lists of Lepidoptera by region By taxonomy *Taxonomy of the Lepidoptera *List of Sphingidae species: (hawk moths) a fam ...
*
Lepidopterism __NOTOC__ Lepidopterism is an irritant contact dermatitis caused by irritating caterpillar or moth hairs coming into contact with the skin or mucosa. When referring to the cause, moth dermatitis and caterpillar dermatitis are commonly used; Caripi ...
*
Pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds ...


References

{{Authority control Agricultural pest insects Household pest insects * Paraphyletic groups