Morelia Tracyae
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''Simalia tracyae'', the Halmahera python, is a species of python found only on the Indonesian island of
Halmahera Halmahera, formerly known as Jilolo, Gilolo, or Jailolo, is the largest island in the Maluku Islands. It is part of the North Maluku province of Indonesia, and Sofifi, the capital of the province, is located on the west coast of the island. Hal ...
. It belongs to the family
Pythonidae The Pythonidae, commonly known as pythons, are a family of nonvenomous snakes found in Africa, Asia, and Australia. Among its members are some of the largest snakes in the world. Ten genera and 42 species are currently recognized. Distributi ...
and the genus ''
Simalia ''Simalia'' is a genus of snakes in the family Pythonidae. Taxonomy ''Simalia'' , was considered a taxonomic synonym of *''Liasis'' (a genus of non-venomous pythons found in Indonesia, New Guinea and Australia) and *''Morelia'' (a genus of larg ...
''. This snake was previously believed to have belonged to the '' Simalia amesthistina'' species; however, studies in recent years have caused scientists to distinguish between the two types of snakes, resulting in the reclassification of the Halmahera python as its own species.


Taxonomy and etymology

Until recently, ''Morelia tracyae'' was considered to be part of the species ''Morelia amesthistina''. Previously, the scientific community believed that this species, despite the large varieties in appearance and wide range of habitat, was thought simply to have "exceptional dispersal abilities". Harvey, Michael. "Systematics of Pythons of the ''Morelia Amethistina'' Complex (Serpentes: Boidae) with the Description of Three New Species". However, more recent scientific studies have indicated ''Morelia amesthistina's'' lack of mobility between habitats, leading to the realization that this type of snake is "more prone to speciation than other pythons". Hoser, Raymond. "Creationism and Contrived Science: A Review of Recent Python Systematics Papers and the Resolution of Issues of Taxonomy and Nomenclature". David G. Barker, a herpetologist and avid python collector, suspected that the high level of variety of Indonesian pythons indicated the presence of multiple species. In 1995, David G. Barker and Michael B. Harvey, herpetological expert and professor at the University of Texas, conducted a study of various specimens of Indonesian snakes. By examining both live specimens and museum specimens, these scientists observed morphological differences and
phylogenetically In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
analyzed the snakes' molecular DNA sequences. In its molecular sequencing, ''Morelia tracyae'' showed a 6-7% divergence in from all other scrub pythons in the surrounding area. The resultant conclusions of this study led to the identification of three new species: ''M. clastolepis'', ''M. nauta'', and ''M. tracyae''. The Halmhera python has been recognized by the scientific community as a separate species since Barker and Harvey's publication of their data in 2000. The species ''Morelia tracyae'' was named for David G. Barker's wife, Tracy M. Barker, who is also a herpetologist specialized in pythons. In 2014, a phylogenetic analysis of Pythonidae based on a combined analysis of eight nuclear DNA loci and three mitochondrial DNA regions concluded the Halmahera python belonged in genus ''Simalia'' with six other species.


Description

''Simalia tracyae'' is a large species of python, and can range anywhere from in length. Corlett, Richard. "Vertebrate carnivores and predation in the Oriental (Indomalayan) region." Barker, David. "Pythones (Pythonidae)." It is described as “heavy-bodied,” and more closely resembles the boa than any other species of python. The Halmahera python is recognized by its distinct red irises, as compared to the typical brown or gold irises of other closely related species. Other facial features include its rounded snout, black tongue, and pale blueish grey
oral mucosa The oral mucosa is the mucous membrane lining the inside of the mouth. It comprises stratified squamous epithelium, termed "oral epithelium", and an underlying connective tissue termed ''lamina propria''. The oral cavity has sometimes been descri ...
. One way to distinguish the sex of a Halmahera python is by noticing the snout: females have a somewhat shorter snout, which leads to the appearance of having a wider head. ''Simalia tracyae'' is also marked by two or three pairs of enlarged parietal scales on either side of its head. The Halmahera python is marked by a distinct scale pattern; there have never been any patternless specimens observed. There are anywhere from 48-56 dark brown or black bands on the body, with the stripes reaching their darkest point on the snake's back and lighter towards the sides. The stripes are 4-11
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
in length, and are separated by 3-7 dorsal scale lengths. In addition to the dorsal stripes, a well-defined postocular stripe and one to two thick black stripes around the neck are often present. The Halmahera python is unusual because of the pattern's consistency across the species, with very little variation. This snake is also unique in that the adult snake has the same pattern as it did in adolescence; there is little to no change in design as the Halmahera python develops.


Coloration

The Halmahera python changes color over a short period of time; however, scientists are unable to determine what circumstances cause the color change. At the darker end of the color spectrum, ''Simalia tracyae'' could be medium brown, orangish-brown, or russet. These colors could change to be a greenish-straw, grey-tan, yellowish-tan, or tan on the lighter end of the spectrum. Though the base color of the snake changes, the pattern of thick dorsal stripes remains the same. The stripes could be dark brown, charcoal-brown, brownish black, charcoal, or black at the dark end of the color continuum and become a medium greenish brown or medium reddish brown as the python shifts to the lighter end of the continuum. However, if a stripe is pure black it will remain black despite any other color changes. Though no newly hatched Halmahera pythons have been observed, Barker and Harvey of the 2000 python study “would predict that hatchlings are very dark reddish-brown in color.”


Behavior

Though scrub pythons in general are found above the ground in canopies, locals of Halmahera have most often encountered ''Simalia tracyae'' crawling on the ground. O'Shea, Mark. "Boas and Pythons of the World." However, this species certainly has the ability to travel through the trees as it is often found near
fruit bat Megabats constitute the family Pteropodidae of the order Chiroptera (bats). They are also called fruit bats, Old World fruit bats, or—especially the genera ''Acerodon'' and ''Pteropus''—flying foxes. They are the only member of the sup ...
roosts. According to local collectors, the Halmahera python feeds on fruit bats and presumably other small mammals and birds. There have also been reports by natives of Halmahera pythons carrying off large numbers of domestic chickens. Like other pythons, the Halmahera python will swallow its victims whole or defeat larger prey through
constriction Constriction is a method used by various snake species to kill or subdue their prey. Although some species of venomous and mildly venomous snakes do use constriction to subdue their prey, most snakes which use constriction lack venom. The snake i ...
. This snake is non-venomous.


Geographic Range

''Simalia tracyae'' is only found in the northwestern area of Halmahera, Indonesia. Specimens have been seen as far south as Jailolo and have been sighted near Galela and Tobelo in the north. ''Simalia tracyae'' has evolved as a separate species because of its secluded location on Halmahera.


Habitat

This python's main habitat is the evergreen rainforest of northern Halmahera, where it moves equally comfortable on the ground and in the trees.


Captivity

The only clutches obtained in captivity are part of a Conservation Program, so information regarding their reproduction, clutch size, and development are extremely limited. In general, scrub pythons do not tend to do well in captivity, due in part to their reputation for biting, and have been described as “heat-seeking guided missiles tipped with teeth.” Barker, David. "Pythons of the World:Australia."


Citations


References

* * * * * * * {{Taxonbar, from=Q178604 Morelia (snake) Reptiles described in 2000