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''Montifringilla'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
birds in the sparrow family
Passeridae Old World sparrows are a group of small passerine birds forming the family Passeridae. They are also known as true sparrows, a name also used for a particular genus of the family, ''Passer''. They are distinct from both the New World sparrows, i ...
. It is one of three genera containing the snowfinches. As the English and scientific names suggest, these are high-altitude species, found in the mountain ranges of southern Eurasia, from the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to C ...
east to the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
,
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
and western China.


Description

Snowfinches are stocky sparrows in length, with strong conical bills. They have pale brown upperparts, white underparts and extensive white panels in the wings, which transform them in flight. Adults may have black markings on the chin or around the eyes. Sexes are usually very similar, although the male white-winged snowfinch has a distinctive grey head. Young birds are a drabber version of the adult., , They have simple repetitive songs, given from a rock or during the elaborate circling display flight. The call is a simple ''chip'' or similar.


Ecology

Most snowfinches breed above altitudes of , but the white-winged snowfinch can occur from upwards. These hardy birds inhabit bare open mountain
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
. Snowfinches are not migratory but may move to lower altitudes or human habitation in winter, when these highly gregarious birds form large flocks. Snowfinches are primarily ground-feeding seed-eaters, though they also consume small
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s, especially when breeding. They are typically fearless, and will forage around ski resorts, human habitation and rubbish tips. They nest in rock crevices, or more typically in holes left by
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s or (and even more often)
pika A pika ( or ; archaically spelled pica) is a small, mountain-dwelling mammal found in Asia and North America. With short limbs, very round body, an even coat of fur, and no external tail, they resemble their close relative, the rabbit, but wi ...
s (Ochotonidae). The typical
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
is from three to six eggs.


Systematics and taxonomy

The genus ''Montifringilla'' was introduced by the German ornithologist
Christian Ludwig Brehm Christian Ludwig Brehm (24 January 1787 – 23 June 1864) was a German pastor and ornithologist. He was the father of the zoologist Alfred Brehm. Life Brehm was born in Schönau near Gotha on 24 January 1787. He was educated at University ...
in 1828. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
was subsequently designated as the
white-winged snowfinch The white-winged snowfinch (''Montifringilla nivalis''), or snowfinch, is a small passerine bird. Despite its name, it is a sparrow rather than a true finch. Taxonomy In 1760 the French zoologist Mathurin Jacques Brisson included a description o ...
. The name of the genus combines the Latin words ''mons'', ''montis'' "mountain" and ''fringilla'' "finch".


Species and systematics

The genus contains three
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
: Four species of snowfich are now separated in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Pyrgilauda ''Pyrgilauda'' is a genus of passerine birds in the sparrow family Passeridae. They are found in the Himalayas, Tibet and western China. The genus name was introduced by the French naturalist Charles Lucien Bonaparte in 1850. The type species wa ...
'', based on the genetics and the different vocalizations and
ecological Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
preferences. Similarly, the
white-rumped snowfinch The white-rumped snowfinch (''Onychostruthus taczanowskii'') is a species of passerine bird in the sparrow family Passeridae. It is the only member of the genus ''Onychostruthus''. It is sometimes placed in the genus ''Montifringilla''. It is f ...
is placed in a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus ''Onychostruthus''.


Taxonomy

The
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
s of ''Montifringilla'' have a convoluted history. ''Pyrgilauda'' is probably the most confused case among them: It was first established by
Charles Lucien Jules Laurent Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857), was a French naturalist and ornithologist. Lucien and his wife had twelve children, including Cardinal Lucien Bonaparte. Life and career ...
as a junior synonym of ''Pyrrhulauda'', and attributed to the brothers Edouard and
Jules Verreaux Jules Pierre Verreaux (24 August 1807 – 7 September 1873) was a French botanist and ornithologist and a professional collector of and trader in natural history specimens. He was the brother of Édouard Verreaux and nephew of Pierre Antoine Dela ...
. But ''Pyrrhulauda'' – established by Andrew Smith before Bonaparte – actually refers to certain sparrow-larks, today all placed in ''Eremopterix''. J. Verreaux indeed seems to have been the first to formally use ''Pyrgilauda'' for a particular snowfinch species, but that was only in 1871, when he described the small snowfinch. It was widely used for the southern snowfinch species in the late 19th and early 20th century, and continues to be used by some authors today. According to Article 11.6.1. and 50.7. in the third and fourth editions of the ''
International Code of Zoological Nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the ...
'', the genus ''Pyrgilauda'' was thus validly established by Bonaparte as a possible snowfinch genus name – even though he did not intend to – because many authors after J. Verreaux used it for these birds, and not for the sparrow-larks. Bonaparte had correctly used ''Montifrigilla'' for the
white-winged snowfinch The white-winged snowfinch (''Montifringilla nivalis''), or snowfinch, is a small passerine bird. Despite its name, it is a sparrow rather than a true finch. Taxonomy In 1760 the French zoologist Mathurin Jacques Brisson included a description o ...
– the only snowfinch known then – however, and thus ''Pyrgilauda'' had remained an unused "shell" of a
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
until someone placed a species in it. That was done by J. Verreaux, and thus many erroneously attributed authorship of the genus to him, although as per the ICZN ''Code'' Article 69.3. he technically just fixed its
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
as the small snowfinch. Another error occurred in 1982, when G. N. Kasin believed ''Pyrgilauda'' had been established twice, and thought the snowfinch genus name – which he believed to have been established by Verreaux in 1870 – was the
junior homonym In biology, a homonym is a name for a taxon that is identical in spelling to another such name, that belongs to a different taxon. The rule in the International Code of Zoological Nomenclature is that the first such name to be published is the se ...
of Bonaparte's sparrow-lark synonym. He set up ''Stepaniania'' to replace it. But with Bonaparte's name (as co-opted by Verreaux and later authors) being a valid junior
subjective synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnae ...
and not
preoccupied The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linn ...
, ''Stepaniania'' itself becomes a
junior objective synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnae ...
of ''Pyrgilauda''. The name ''Orospiza'' has a similar history to ''Pyrgilauda''. Bonaparte merely remarked it had been used by "some" (''aliquī'') unspecified authors, while many subsequent sources attributed it to
Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kaup ...
. But that is a misunderstanding, and the only technically valid (though synonymous) name for the snowfinch genus proposed by Kaup was ''Chionospina''. Thus, in the case of ''Orospiza'' Bonaparte is again the valid author, though
Thomas Horsfield Thomas Horsfield (May 12, 1773 – July 24, 1859) was an American physician and natural history, naturalist who worked extensively in Indonesia, describing numerous species of plants and animals from the region. He was later a curator of the Eas ...
and
Frederic Moore Frederic Moore FZS (13 May 1830 – 10 May 1907) was a British entomologist and illustrator. He produced six volumes of ''Lepidoptera Indica'' and a catalogue of the birds in the collection of the East India Company. It has been said that Mo ...
were the first to set a type species. As ''M. nivalis'' was still the only known snowfinch in 1858 when they did this, ''Orospiza'' is a junior objective synonym of ''Montifringilla''. For somewhat different reasons, ''Orites'' is a junior objective synonym of ''Montifringilla''. First proposed for the
long-tailed tit The long-tailed tit (''Aegithalos caudatus''), also named long-tailed bushtit, is a common bird found throughout Europe and the Palearctic. The genus name ''Aegithalos'' was a term used by Aristotle for some European tits, including the long-tail ...
(''Aegithalos caudatus'') in Moehring's ''Geslachten der Vogelen'' in 1758, that widely ignored work was suppressed as a source of taxa by the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
in 1987. Consequently, later attempts to re-establish that name as a junior synonym of ''
Aegithalos ''Aegithalos'' is a songbird genus in the family Aegithalidae. Taxonomy The genus ''Aegithalos'' was introduced in 1804 by the French naturalist Johann Hermann to accommodate a single species, the long-tailed tit. The genus name was a term used ...
'' are preempted by the erection of ''Orites'' for the white-winged snowfinch by
Eugen von Keyserling Eugen von Keyserling (22 March 1833 in Pockroy, Lithuania – 4 April 1889 in Dzierżoniów, Silesia) was a Baltic-German arachnologist. He studied in the University of Tartu. He was the author of ''Die Spinnen Amerikas'', and completed ''Die Ara ...
and
Johann Heinrich Blasius Johann Heinrich Blasius (7 October 1809 – 26 May 1870) was a German zoologist. His sons, Rudolf Heinrich Paul Blasius (1842-1907) and August Wilhelm Heinrich Blasius (1845–1912) were ornithologists. Biography In 1836, he was appointed as a ...
in 1840. Meanwhile, Kaup's ''Chionospina'' was misspelled "Chionospiza" by
George Robert Gray George Robert Gray FRS (8 July 1808 – 6 May 1872) was an English zoologist and author, and head of the ornithological section of the British Museum, now the Natural History Museum, in London for forty-one years. He was the younger brother o ...
. As it, too, is a junior objective synonym of ''Montifringilla'',
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museu ...
's subsequent valid establishment of that name is still inconsequential – except for purely taxonomic purposes, as it makes neither Kaup nor Grey the valid author of ''Chionospiza'', but Reichenbach., , And finally, ''Onychostruthus'' had to be established because ''Onychospiza'',
Nikolai Przhevalsky Nikolay Mikhaylovich Przhevalsky (or Prjevalsky;; pl, Nikołaj Przewalski, . – ) was a Russian geographer of Polish descent (he was born in a Polish noble family), and a renowned explorer of Central and East Asia. Although he never reache ...
's 1876 name for the proposed monotypic genus of the
white-rumped snowfinch The white-rumped snowfinch (''Onychostruthus taczanowskii'') is a species of passerine bird in the sparrow family Passeridae. It is the only member of the genus ''Onychostruthus''. It is sometimes placed in the genus ''Montifringilla''. It is f ...
, was preoccupied. It had been given by Rey to the chestnut-eared bunting (today ''Emberiza fucata'') in 1872 already, as an unnecessary correction – and thus junior objective synonym – of Bonaparte's original ''Onychospina''.


Footnotes


References

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External links


Media related to Passeridae
on the Internet Bird Collection {{Taxonbar, from=Q922089 Taxa named by Christian Ludwig Brehm