Mesostigmata is an order of

mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evid ...

s belonging to the

Parasitiformes Parasitiformes are a superorder of Arachnids, constituting one of the two major groups of mites, alongside Acariformes. Parasitiformes has, at times, been classified at the rank of order or suborder. It is uncertain whether Parasitiformes and Ac ...

. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata includes

parasitic Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson ha ...

as well as free-living and

predatory Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill t ...

forms. They can be recognized by the single pair of spiracles positioned laterally on the body. The family with the most described species is

Phytoseiidae The Phytoseiidae are a family of mites which feed on thrips and other mite species. They are often used as a biological control agent for managing mite pests. Because of their usefulness as biological control agents, interest in Phytoseiidae has ...

. Other families of note are

Diplogyniidae Diplogyniidae is a family of parasitic mites belonging to the order Mesostigmata. Many are parasites on beetles but some live on larger animals (e.g., rats). Taxonomy This is a list of the described species. The data is taken from Joel Hallan's ...

Macrochelidae Macrochelidae is a family of mites in the order Mesostigmata. Description Adult females of Macrochelidae have: an undivided dorsal shield bearing at least 28 pairs of setae, a sternal shield with three pairs of setae, a genital shield with one ...

Pachylaelapidae Pachylaelapidae is a family of mites in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata includ ...

, Uropodidae and Veigaiidae. Amongst the best known species are ''

Varroa destructor ''Varroa destructor'', the ''Varroa'' mite is an external parasitic mite that attacks and feeds on the honey bees ''Apis cerana'' and ''Apis mellifera''. The disease caused by the mites is called varroosis. The ''Varroa'' mite can reproduce on ...

'', an economically important parasite of honey bees, as well as the red mite (''Dermanyssus gallinae'') a parasite of poultry, most commonly chickens.

Description

Mesostigmata are mites ranging from 0.12-4 mm long (0.2-4 mm according to another source). They have a pair of stigmatal openings above legs III-IV usually associated with a peritrematal groove. The

gnathosoma The gnathosoma (from Greek , ' = "jaw" and , ' = "body") is the part of the body of the Acari (mites and ticks) comprising the mouth and feeding parts. These are the hypostome, the chelicerae and the pedipalps. It is also called the capitulum (how ...

has a sclerotised ring around the bases of the

chelicerae The chelicerae () are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as " jaws", chelicerae may be shaped as either articulated fangs, or simila ...

(basis capitulum). The palps have 5 (rarely 4) free segments and usually a subdistal palp apotele. The chelicerae are 3-segmented. The subcapitulum usually has a median groove with transverse rows of 1-many denticles. There are usually bifurcate or membranous corniculi present. Except in some

species, a flagellate tritosternum is present. The coxae of the legs are freely articulating with the body. The intercoxal region has sternal and genital shield elements. Adults have a genital opening and either chelicerae modified for sperm transfer (if male) or a sperm-receiving structure (if female). The above description applies to adults. Larvae have six legs, instead of the eight possessed by later stages, and may or may not feed. There are two nymphal stages (protonymph, deutonymph) that usually have lightly sclerotized dorsal, intercoxal and ventral plates.

Ecology

Many Mesostigmata are free-living predators of invertebrates that live in soil and litter, on the soil surface or on plants. There are also some that live in freshwater. Other Mesostigmata are parasites of vertebrates or arthropods, pollen and nectar feeders in flowers, fungus feeders, or saprophages that subsist on dead or decaying organic matter. The soil-dwelling Mesostigmata are not as abundant as oribatids or prostigmatids that also occur in this habitat, but they are still ubiquitous in soil and may be important predators. Larger species tend to be predators of small arthropods or arthropod eggs, whereas smaller species prey on

nematodes The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broa ...

. Size of these mesostigmatans decreases with soil depth:

plant litter Plant litter (also leaf litter, tree litter, soil litter, litterfall or duff) is dead plant material (such as leaves, bark, needles, twigs, and cladodes) that have fallen to the ground. This detritus or dead organic material and its constitue ...

and

humus In classical soil science, humus is the dark organic matter in soil that is formed by the decomposition of plant and animal matter. It is a kind of soil organic matter. It is rich in nutrients and retains moisture in the soil. Humus is the Lati ...

have large species such as '' Veigaia'' ( Veigaiidae), the humus-soil interface has smaller species like '' Dendrolaelaps'' (

Digamasellidae Digamasellidae is a family of mites in the order Mesostigmata. Genera These 11 genera belong to the family Digamasellidae: * ''Dendrolaelaps'' Halbert, 1915 * '' Dendrolaelaspis'' Lindquist, 1975 * ''Dendroseius'' Karg, 1965 * ''Digamasellus'' B ...

) and the mineral soil has the tiny '' Rhodacarellus'' (

Rhodacaridae Rhodacaridae is a family of mites in the order Mesostigmata.Walter, David Evans, edRhodacaridae Species Listing Biology Catalog. Texas A&M University. Retrieved on August 27, 2010. Genera * '' Afrodacarellus'' Hulbutt, 1974 * '' Afrogamasellus ...

). A few species are known from freshwater habitats, such as wet soil, phytotelmata, waterside vegetation and sewage filter-beds. These appear to move by crawling as no species are known to swim. Some species are known to prey on

mosquito Mosquitoes (or mosquitos) are members of a group of almost 3,600 species of small flies within the family Culicidae (from the Latin ''culex'' meaning "gnat"). The word "mosquito" (formed by ''mosca'' and diminutive ''-ito'') is Spanish for "lit ...

eggs and one species was reared on a diet of nematodes. The parasitic Mesostigmata are mostly in superfamily Dermanyssoidea. These include parasites of invertebrates (e.g. Varroidae) and of vertebrates (other families), as well as both ectoparasites (external) and endoparasites (internal).

Phoresy Phoresis or phoresy is a non-permanent, commensalistic interaction in which one organism (a phoront or phoretic) attaches itself to another (the host) solely for the purpose of travel. Phoresis has been observed directly in ticks and mites s ...

, the temporary attachment of a smaller animal to a larger one for travel, is common in the Mesostigmata. For example, the freshwater species are phoretic on flies of families

Tipulidae Crane fly is a common name referring to any member of the insect family Tipulidae. Cylindrotominae, Limoniinae, and Pediciinae have been ranked as subfamilies of Tipulidae by most authors, though occasionally elevated to family rank. In the m ...

Ceratopogonidae Ceratopogonidae is a family of flies commonly known as no-see-ums, or biting midges, generally in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic. Ceratopogonidae are holomet ...

, and Culicidae.

Economic importance

'' (Varroidae) is a major pest of

honey bees A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cos ...

. It harms bees both directly by feeding on fat body tissue, and indirectly by transmitting viruses. Similarly, the red mite (''

Dermanyssus gallinae ''Dermanyssus gallinae'' (also known as the red mite) is a haematophagous ectoparasite of poultry. It has been implicated as a vector of several major pathogenic diseases. Despite its common names, it has a wide range of hosts including sever ...

'') feeds on the blood of birds, including

poultry Poultry () are domesticated birds kept by humans for their eggs, their meat or their feathers. These birds are most typically members of the superorder Galloanserae (fowl), especially the order Galliformes (which includes chickens, quail ...

(

chickens The chicken (''Gallus gallus domesticus'') is a domestication, domesticated junglefowl species, with attributes of wild species such as the grey junglefowl, grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster ...

turkeys The turkey is a large bird in the genus ''Meleagris'', native to North America. There are two extant turkey species: the wild turkey (''Meleagris gallopavo'') of eastern and central North America and the ocellated turkey (''Meleagris ocella ...

ducks Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a for ...

) and wild birds. It reduces animal health, welfare and production. In agriculture, soil-dwelling mesostigmatans are important predators of nematodes,

springtails Springtails (Collembola) form the largest of the three lineages of modern hexapods that are no longer considered insects (the other two are the Protura and Diplura). Although the three orders are sometimes grouped together in a class called Ento ...

and insect larvae, while plant-dwelling mesostigmatans control pests such as spider mites.

Evolution

The oldest known record of the group is an indeterminate

Sejidae Sejidae is a family of mites in the order Mesostigmata. The oldest known record of the group is an indeterminate deutonymph from the mid Cretaceous (Albian-Cenomanian) aged Burmese amber of Myanmar. Species '' Archaeopodella'' Athias-Henriot ...

deutonymph from the mid-

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

(Albian-Cenomanian) aged

Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. ...

of Myanmar.

Taxonomy

Taxonomy to families (Beaulieu 2011). Genus and species counts fluctuate over time. : Order Mesostigmata G. Canestrini, 1891 :: Suborder Monogynaspida Camin & Gorirossi, 1955 ::: Infraorder Gamasina Kramer, 1881 :::: Hyporder Arctacariae Johnston, 1982 ::::: Superfamily Arctacaroidea Evans, 1955 :::::: Family Arctacaridae Evans, 1955 (2 genera, 6 species) :::: Hyporder

Dermanyssiae Dermanyssiae is a "sub-cohort" of mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but ...

Evans & Till, 1979 ::::: Superfamily Ascoidea Voigts & Oudemans, 1905 :::::: Family

Ameroseiidae The family Ameroseiidae is one of the three families of mites under the superfamily Ascoidea. There are about 12 genera and more than 130 described species in Ameroseiidae. The family has a worldwide distribution. Description Ameroseiidae ...

Evans, 1961 (10 genera, 148 species) :::::: Family

Ascidae Ascidae is a family of mites in the order Mesostigmata. Description Ascidae are mites characterised by: seta ''st4'' usually on unsclerotised cuticle, peritrematic shield broadly connected to exopodal shield beside coxa IV, fixed chelicera ...

Voigts & Oudemans, 1905 (17 genera, 338 species) :::::: Family Melicharidae Hirschmann, 1962 (12 genera, 201 species) ::::: Superfamily Dermanyssoidea Kolenati, 1859 :::::: Family Dasyponyssidae Fonseca, 1940 (2 genera, 2 species) :::::: Family Dermanyssidae Kolenati, 1859 (2 genera, 26 species) :::::: Family Entonyssidae Ewing, 1923 (9 genera, 27 species) :::::: Family

Haemogamasidae Haemogamasidae is a family of mites in the order Mesostigmata. Genera * '' Acanthochela'' Ewing, 1933 * '' Brevisterna'' Keegan, 1949 * '' Euhaemogamasus'' Ewing, 1933 * ''Eulaelaps'' Berlese, 1903 * ''Haemogamasus ''Haemogamasus'' is a genus ...

Oudemans, 1926 (5 genera, 78 species) :::::: Family Halarachnidae Oudemans, 1906 (7 genera, 43 species) :::::: Family

Hystrichonyssidae ''Hystrichonyssus turneri'' is a species of mite placed in its own family, Hystrichonyssidae, in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, wit ...

Keegan, Yunker & Baker, 1960 (1 genus, 1 species) :::::: Family Iphiopsididae Kramer, 1886 (14 genera, 68 species) :::::: Family

Ixodorhynchidae Ixodorhynchidae is a small family of mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, ...

Ewing, 1923 (6 genera, 43 species) :::::: Family

Laelapidae The Laelapidae are a family (biology), family of mites in the order Mesostigmata. The family is also referred to in the literature as Laelaptidae, which may be the correct spelling. Description Laelapidae have a shield covering all or most of ...

Berlese, 1892 (90 genera, 1316 species) :::::: Family

Larvamimidae ''Larvamima'' is a genus of mites placed in its own family, Larvamimidae, in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species ...

Elzinga, 1993 (6 genera, 43 species) :::::: Family Macronyssidae Oudemans, 1936 (34 genera, 233 species) :::::: Family

Manitherionyssidae ''Manitherionyssus heterotarsus'' is a species of mite placed in its own family, Manitherionyssidae, in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitifor ...

Radovsky & Yunker, 1971 (1 genus, 1 species) :::::: Family

Omentolaelapidae ''Omentolaelaps mehelyae'' is a species of mite placed in its own family, Omentolaelapidae, in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with ...

Fain, 1961 (1 genus, 1 species) :::::: Family

Rhinonyssidae Rhinonyssidae is a family of mites in the order Mesostigmata. There are about 16 genera and at least 460 described species in Rhinonyssidae. The mites of this family are obligate parasites of avian respiratory systems, living in nasal passagewa ...

Trouessart, 1895 (8 genera, 510 species) :::::: Family Spelaeorhynchidae Oudemans, 1902 (1 genus, 7 species) :::::: Family

Spinturnicidae Spinturnicidae is a family of mites in the order Mesostigmata. The mites are highly specialized parasites of wing or tail membrane or bats. Some species infest eyelids and eye canthi. The species of Spinturnicidae are found in bat habitats thro ...

Oudemans, 1901 (12 genera, 101 species) :::::: Family

Varroidae ''Varroa'' is a genus of parasitic mesostigmatan mites associated with honey bees, placed in its own family, Varroidae. The genus was named for Marcus Terentius Varro, a Roman scholar and beekeeper. The condition of a honeybee colony being infe ...

Delfinado & Baker, 1974 (2 genera, 6 species) ::::: Superfamily Eviphidoidea Berlese, 1913 :::::: Family Eviphididae Berlese, 1913 (19 genera, 108 species) :::::: Family

Leptolaelapidae Leptolaelapidae is a family of mites in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata includ ...

Karg, 1978 (12 genera, 48 species) :::::: Family

Vitzthum, 1930 (20 genera, 470 species) :::::: Family

Berlese, 1913 (26 genera, 199 species) :::::: Family Parholaspididae Evans, 1956 (12 genera, 96 species) ::::: Superfamily Phytoseioidea Berlese, 1916 :::::: Family Blattisociidae Garman, 1948 (11 genera, 369 species) :::::: Family Otopheidomenidae Treat, 1955 (10 genera, 28 species) :::::: Family

Berlese, 1916 (90 genera, 2300 species) :::::: Family Podocinidae Berlese, 1913 (2 genera, 25 species) ::::: Superfamily

Rhodacaroidea Rhodacaroidea is a superfamily of mites in the order Mesostigmata. There are 6 families with more than 900 described species in Rhodacaroidea, found worldwide. These mites inhabit soil and litter, including rodent and nests, moss and lichen, te ...

Oudemans, 1902 :::::: Family

Evans, 1957 (13 genera, 261 species) :::::: Family Halolaelapidae Karg, 1965 (4 genera, 80 species) :::::: Family

Laelaptonyssidae ''Starkovia'' is a genus of mites in the family Laelaptonyssidae. There are about eight described species in ''Starkovia'', some of which were transferred from the genus ''Laelaptonyssus'' when it was determined to be a synonym of ''Starkovia''. ...

Womersley, 1956 (1 genus, 6 species) :::::: Family

Ologamasidae Ologamasidae is a family of mites in the order Mesostigmata. There are more than 40 genera and 470 described species in Ologamasidae. Genera These 44 genera belong to the family Ologamasidae: * '' Acugamasus'' Lee, 1970 * '' Acuphis'' Karg, 19 ...

Ryke, 1962 (45 genera, 452 species) :::::: Family

Oudemans, 1902 (15 genera, 148 species) :::::: Family Teranyssidae Halliday, 2006 (15 genera, 148 species) ::::: Superfamily Veigaioidea Oudemans, 1939 :::::: Family Veigaiidae Oudemans, 1939 (4 genera, 95 species) :::: Hyporder Epicriiae Kramer, 1885 ::::: Superfamily Epicrioidea Berlese, 1885 :::::: Family Epicriidae Berlese, 1885 (4 genera, 48 species) ::::: Superfamily Heatherelloidea Walter, 1997 :::::: Family

Heatherellidae ''Heatherella'' is a genus of mites placed in its own family, Heatherellidae, in the order Mesostigmata. ''Heatherella'' contains two recognized species: * '' Heatherella acanthocharis'' Walter, 1997 * '' Heatherella callimaulos'' Walter, 199 ...

Walter, 1997 (1 genus, 2 species) ::::: Superfamily Zerconoidea G. Canestrini, 1891 :::::: Family Coprozerconidae Moraza & Lindquist, 1999 (1 genus, 1 species) :::::: Family

Zerconidae Zerconidae is a family of mites in the order Mesostigmata. Species '' Acoesejus'' Selinick, 1941 *''Acoesejus muricatus'' (C.L.Koch, 1839) '' Aleksozercon'' A. D. Petrova, 1978 *''Aleksozercon zachvatkini'' A. D. Petrova, 1978 '' Amerozercon' ...

G. Canestrini, 1891 (36 genera, 390, species) :::: Hyporder Parasitiae Evans & Till, 1979 ::::: Superfamily Parasitoidea Oudemans, 1901 :::::: Family Parasitidae Oudemans, 1901 (35 genera, 426 species) ::: Infraorder

Uropodina Uropodina is an infraorder of mites in the order Mesostigmata. Description In Uropodina, the sternal and ventral shields are usually fused, the hypostomal setae are situated in a row, the legs are relatively short and with reduced setae, the ...

Kramer, 1881 ::::: Family Dithinozerconidae Ainscough, 1979 :::: Superfamily Diarthrophalloidea Trägårdh, 1946 ::::: Family Diarthrophallidae Trägårdh, 1946 (22 genera, 63 species) :::: Superfamily Microgynioidea Trägårdh, 1942 ::::: Family

Microgyniidae Microgyniidae is a small family of mites in the order Mesostigmata. Species Microgyniidae contains two genera, with four recognized species: * Genus ''Microsejus'' Trägårdh, 1942 ** ''Microsejus camini'' Trägårdh, 1942 * Genus ''Microgynium ...

Trägårdh, 1942 (2 genera, 4 species) ::::: Family Nothogynidae Walter & Krantz, 1999 (1 genus, 2 species) :::: Superfamily Thinozerconoidea Halbert, 1915 ::::: Family Protodinychidae Evans, 1957 (1 genus, 3 species) ::::: Family

Thinozerconidae ''Thinozercon michaeli'' is a species of mite, placed in its own family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other rel ...

Halbert, 1915 (1 genus, 1 species) :::: Superfamily Uropodoidea Kramer, 1881 ::::: Family Baloghjkaszabiidae Hirschmann, 1979 (1 genus, 3 species) ::::: Family Brasiluropodidae Hirschmann, 1979 (2 genera, 18 species) ::::: Family Cillibidae Trägårdh, 1944 (2 genera, 19 species) ::::: Family Clausiadinychidae Hirschmann, 1979 (1 genus, 4 species) ::::: Family Cyllibulidae Hirschmann, 1979 (1 genus, 32 species) ::::: Family

Deraiophoridae ''Deraiophorus'' is a genus of mites in the order Mesostigmata, placed in its own family, Deriaphoridae. Species * '' Deraiophorus adriaticus'' Hirschmann & Zirngiebl-Nicol, 1972 * '' Deraiophorus aokii'' Hiramatsu, 1982 * '' Deraiophorus au ...

Trägårdh, 1952 (1 genus, 36 species) ::::: Family Dinychidae Berlese, 1916 (1 genus, 34 species) ::::: Family Discourellidae Baker & Wharton, 1952 (1 genus, 76 species) ::::: Family Eutrachytidae Trägårdh, 1944 (1 genus, 36 species) ::::: Family Hutufeideriidae Hirschmann, 1979 (1 genus, 9 species) ::::: Family Kaszabjbaloghiidae Hirschmann, 1979 (1 genus, 6 species) ::::: Family

Macrodinychidae ''Macrodinychus'' is the sole genus in the monotypic taxon, monotypic Macrodinychidae, a family of mites in the order Mesostigmata. Species Macrodinychidae contains one genus, with 22 recognized species: * Genus ''Macrodinychus'' Berlese, 1917 ...

Hirschmann, 1979 (4 genera, 22 species) ::::: Family

Metagynuridae ''Metagynella'' is a genus of mites, placed in its own family, Metagynuridae in the order Mesostigmata. Species * '' Metagynella africana'' Ryke, 1958 * '' Metagynella applicata'' (Vitzthum, 1921) * '' Metagynella baloghi'' Hirschmann, 1975 * ...

Balogh, 1943 (2 genera, 17 species) ::::: Family Nenteriidae Hirschmann, 1979 (2 genera, 128 species) ::::: Family Oplitidae Johnston, 1968 (8 genera, 163 species) ::::: Family Phymatodiscidae Hirschmann, 1979 (1 genus, 10 species) ::::: Family Polyaspididae Berlese, 1913 (1 genus, 16 species) ::::: Family Prodinychidae Berlese, 1917 (3 genera, 16 species) ::::: Family Rotundabaloghiidae Hirschmann, 1979 (4 genera, 165 species) ::::: Family Tetrasejaspidae Hirschmann, 1979 (1 genus, 15 species) ::::: Family

Trachytidae Trachytidae is a family of mites in the order Mesostigmata. Genera In 1960, Daniel G. Maiello, PhD in entomology, was doing research about mites Laelaps multispinosus. He discovered a related unnamed family, similar to the family Trachytidae. * ...

Trägårdh, 1938 (7 genera, 108 species) ::::: Family

Trachyuropodidae Trachyuropodidae is a family of mites in the order Mesostigmata. Species * ''Capitodiscus'' Vitzthum, 1931 ** ''Capitodiscus venusta'' (Berlese, 1884) * ''Cephalouropoda'' Berlese, 1903 ** ''Cephalouropoda berlesiana'' (Berlese, 1887) * ''Crini ...

Berlese, 1917 (17 genera, 99 species) ::::: Family

Trematuridae Trematuridae is a family of mites in the order Mesostigmata. Genera * '' Ipiduropoda'' Sellnick, 1952 * '' Trematuroides'' Cooreman, 1960 * '' Trichofrondosa'' W. Hirschmann, 1986 * '' Trichoobscura'' W. Hirschmann, 1986 * '' Trichouropoda'' B ...

Berlese, 1917 (13 genera, 401 species) ::::: Family Trichocyllibidae Hirschmann, 1979 (5 genera, 57 species) ::::: Family Trichouropodellidae Hirschmann, 1979 (1 genus, 11 species) ::::: Family Trigonuropodidae Hirschmann, 1979 (1 genus, 87 species) ::::: Family Uroactiniidae Hirschmann & Zirngiebl-Nicol, 1964 (3 genera, 67 species) ::::: Family Urodiaspididae Trägårdh, 1944 (3 genera, 26 species) ::::: Family

Urodinychidae Urodinychidae is a family of mites in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata includes ...

Berlese, 1917 (13 genera, 267 species) ::::: Family Uropodidae Kramer, 1881 (9 genera, 261 species) :: Suborder Sejida Kramer, 1885 ::: Superfamily Heterozerconoidea Berlese, 1892 :::: Family

Discozerconidae Discozerconidae is a small family of mites in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata ...

Berlese, 1910 (2 genera, 3 species) :::: Family

Heterozerconidae Heterozerconidae is a small family of mites in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata ...

Berlese, 1892 (7 genera, 13 species) ::: Superfamily Sejoidea Berlese, 1885 :::: Family

Ichthyostomatogasteridae Ichthyostomatogasteridae is a family of mites belonging to the order Mesostigmata. Genera: * '' Asternolaelaps'' Berlese, 1923 * '' Japanasternolaelaps'' Hirschmann & Hiramatsu, 1984 * ''Archaeopodella ''Archaeopodella'' is a genus of mites in ...

Sellnick, 1953 (3 genera, 10 species) :::: Family

Berlese, 1885 (5 genera, 46 species) (5 genera, 46 species) :::: Family Uropodellidae Camin, 1955 (1 genus, 6 species) :: Suborder Trigynaspida Camin & Gorirossi, 1955 ::: Infraorder Antennophorina Camin & Gorirossi, 1955 :::: Superfamily Aenictequoidea Kethley, 1977 ::::: Family Aenictequidae Kethley, 1977 (1 genus, 1 species) ::::: Family Euphysalozerconidae Kim, 2008 (1 genus, 1 species) ::::: Family

Messoracaridae Messoracaridae is a family of mites in the order Mesostigmata. Taxonomy * Genus '' Leptantennus'' Berlese, 1916 ** '' Leptantennus pendulipes'' Berlese, 1916 * Genus '' Messoracarus'' Silvestri, 1912 ** '' Messoracarus emarginatus'' (Banks, 19 ...

Kethley, 1977 (2 genera, 3 species) ::::: Family Ptochacaridae Kethley, 1977 (1 genus, 3 species) :::: Superfamily Antennophoroidea Berlese, 1892 ::::: Family

Antennophoridae Antennophoridae is a family of mites in the order Mesostigmata. Genera Antennophoridae contains five genera, with nine recognized species. '' Antennophorus grandis'' is an ectoparasite on ants of the species ''Lasius flavus The yellow meadow ...

Berlese, 1892 (6 genera, 19 species) :::: Superfamily Celaenopsoidea Berlese, 1892 ::::: Family

Celaenopsidae Celaenopsidae is a family of mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but gene ...

Berlese, 1892 (7 genera, 14 species) ::::: Family

Costacaridae Costacaridae is a family of mites in the order Mesostigmata. Species Costacaridae contains one genus, with one recognized species: * Genus ''Costacarus'' P. E. Hunter, 1993 ** '' Costacarus reyesi'' P. E. Hunter, 1993 References Mesosti ...

Hunter, 1993 (1 genus, 1 species) ::::: Family

Trägårdh, 1941 (42 genera, 85 species) ::::: Family Euzerconidae Trägårdh, 1938 (12 genera, 24 species) ::::: Family Megacelaenopsidae Funk, 1975 (2 genera, 2 species) ::::: Family Neotenogyniidae Kethley, 1974 (1 genus, 1 species) ::::: Family Schizogyniidae Trägårdh, 1950 (6 genera, 10 species) ::::: Family

Triplogyniidae Triplogyniidae is a family of mites in the order Mesostigmata. Species Triplogyniidae contains two genera, with ten recognized species: * Genus ''Funkotriplogynium'' A. Kumar-Datta, 1985 ** ''Funkotriplogynium bengalensis'' (Pramanik & Raychaud ...

Funk, 1977 (2 genera, 11 species) :::: Superfamily Fedrizzioidea Trägårdh, 1937 ::::: Family

Fedrizziidae Fedrizziidae is a family of mites in the order Mesostigmata. Species Fedrizziidae contains three genera, with 31 recognized species: * Genus '' Fedrizzia'' G.Canestrini, 1884 ** '' Fedrizzia abradoalves'' Seeman, 2007 ** '' Fedrizzia bornemis ...

Trägårdh, 1937 (3 genera, 34 species) ::::: Family Klinckowstroemiidae Camin & Gorirossi, 1955 (4 genera, 36 species) :::: Superfamily Megisthanoidea Berlese, 1914 ::::: Family

Hoplomegistidae Hoplomegistidae is a family of mites in the order Mesostigmata. Species Hoplomegistidae contains one genus, with two recognized species: * Genus ''Stenosternum'' Kramer, 1898 ** ''Stenosternum armiger'' (Berlese, 1888) ** ''Stenosternum bipilos ...

Camin & Gorirossi, 1955 (1 genus, 7 species) ::::: Family Megisthanidae Berlese, 1914 (1 genus, 30 species) :::: Superfamily Paramegistoidea Trägårdh, 1946 ::::: Family Paramegistidae Trägårdh, 1946 (5 genera, 30 species) :::: Superfamily Parantennuloidea Willmann, 1941 ::::: Family Parantennulidae Willmann, 1941 (3 genera, 5 species) ::::: Family Philodanidae Kethley, 1977 (2 genera, 2 species) ::::: Family

Promegistidae Promegistidae is a family of mites in the order Mesostigmata. Species Promegistidae contains one genus, with one recognized species: * Genus '' Promegistus'' Womersley, 1958 ** '' Promegistus armstrongi'' Womersley, 1958 References Meso ...

Kethley, 1977 (1 genus, 1 species) ::: Infraorder Cercomegistina Camin & Gorirossi, 1955 :::: Superfamily Cercomegistoidea Trägårdh, 1937 ::::: Family

Asternoseiidae Asternoseiidae is a family of mites in the order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the ...

Vale, 1954 (2 genera, 3 species) ::::: Family

Cercomegistidae Cercomegistidae is a family of mites in the order Mesostigmata. Species Cercomegistidae contains four genera, with six recognized species: * Genus '' Celaenogamasus'' Berlese, 1901 ** '' Celaenogamasus hirtellus'' Berlese, 1901 * Genus '' Cer ...

Trägårdh, 1937 (5 genera, 13 species) ::::: Family

Davacaridae Davacaridae is a family of mites in the order Mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata includes p ...

Kethley, 1977 (2 genera, 4 species) ::::: Family Pyrosejidae Lindquist & Moraza, 1993 (2 genera, 3 species) ::::: Family Saltiseiidae Walter, 2000 (1 genus, 1 species) ::::: Family Seiodidae Kethley, 1977 (1 genus, 1 species) ; Other :'' Meliponopus palpifer'' Fain & Flechtmann, 1985 has not yet been placed into a family.

References

* * NCBI
Mesostigmata

External links

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''Featured Creatures'' website {{Taxonbar, from=Q1240844 Arachnid orders