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Monkey is a common name that may refer to most mammals of the
infraorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
Simiiformes The simians, anthropoids, or higher primates are an infraorder (Simiiformes ) of primates containing all animals traditionally called monkeys and apes. More precisely, they consist of the parvorders New World monkeys (Platyrrhini) and Catarrhi ...
, also known as the simians. Traditionally, all animals in the group now known as simians are counted as monkeys except the
ape Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its siste ...
s, which constitutes an incomplete
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
grouping; however, in the broader sense based on
cladistics Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
, apes (Hominoidea) are also included, making the terms ''monkeys'' and ''simians'' synonyms in regards to their scope. In 1812, Geoffroy grouped the
apes Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its sister g ...
and the
Cercopithecidae Old World monkey is the common English name for a family of primates known taxonomically as the Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons ...
group of monkeys together and established the name
Catarrhini The parvorder Catarrhini , catarrhine monkeys, Old World anthropoids, or Old World monkeys, consisting of the Cercopithecoidea and apes (Hominoidea). In 1812, Geoffroy grouped those two groups together and established the name Catarrhini, "Old ...
, "Old World monkeys", ("''singes de l'Ancien Monde''" in
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
). The extant sister of the Catarrhini in the monkey ("singes") group is the Platyrrhini (New World monkeys). Some nine million years before the divergence between the Cercopithecidae and the apes, the Platyrrhini emerged within "monkeys" by migration to South America likely by ocean. Apes are thus deep in the tree of extant and extinct monkeys, and any of the apes is distinctly closer related to the Cercopithecidae than the Platyrrhini are. Many monkey species are tree-dwelling (
arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
), although there are species that live primarily on the ground, such as
baboon Baboons are primates comprising the genus ''Papio'', one of the 23 genera of Old World monkeys. There are six species of baboon: the hamadryas baboon, the Guinea baboon, the olive baboon, the yellow baboon, the Kinda baboon and the chacma ba ...
s. Most species are mainly active during the day ( diurnal). Monkeys are generally considered to be intelligent, especially the Old World monkeys. Within suborder
Haplorhini Haplorhini (), the haplorhines (Greek for "simple-nosed") or the "dry-nosed" primates, is a suborder of primates containing the tarsiers and the simians (Simiiformes or anthropoids), as sister of the Strepsirrhini ("moist-nosed"). The name is some ...
, the simians are a sister group to the
tarsier Tarsiers ( ) are haplorhine primates of the family Tarsiidae, which is itself the lone extant family within the infraorder Tarsiiformes. Although the group was once more widespread, all of its species living today are found in Maritime Southeast ...
s – the two members diverged some 70 million years ago. New World monkeys and catarrhine monkeys emerged within the simians roughly 35 million years ago. Old World monkeys and apes emerged within the catarrhine monkeys about 25 million years ago. Extinct basal simians such as ''
Aegyptopithecus ''Aegyptopithecus'' ("Egyptian ape", from Greek ''Αίγυπτος'' "Egypt" and ''πίθηκος'' "ape") is an early fossil catarrhine that predates the divergence between hominoids (apes) and cercopithecids (Old World monkeys). It is known ...
'' or ''
Parapithecus ''Parapithecus'' is an extinct genus of primate that lived during the Late Eocene- Earliest Oligocene in what is now Egypt. Its members are considered to be basal anthropoids and the genus is closely related to ''Apidium The genus ''Apidium'' ...
'' (35–32 million years ago) are also considered monkeys by primatologists.
Lemur Lemurs ( ) (from Latin ''lemures'' – ghosts or spirits) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea (), divided into 8 Family (biology), families and consisting of 15 genera and around 100 exist ...
s,
lorises Loris is the common name for the strepsirrhine mammals of the subfamily Lorinae (sometimes spelled Lorisinae) in the family Lorisidae. ''Loris'' is one genus in this subfamily and includes the slender lorises, ''Nycticebus'' is the genus contain ...
, and
galago Galagos , also known as bush babies, or ''nagapies'' (meaning "night monkeys" in Afrikaans), are small nocturnal primates native to continental, sub-Sahara Africa, and make up the family Galagidae (also sometimes called Galagonidae). They ar ...
s are not monkeys, but
strepsirrhine Strepsirrhini or Strepsirhini (; ) is a suborder of primates that includes the lemuriform primates, which consist of the lemurs of Madagascar, galagos ("bushbabies") and pottos from Africa, and the lorises from India and southeast Asia. Collec ...
primates (suborder Strepsirrhini). The simians'
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
, the tarsiers, are also haplorhine primates; however, they are also not monkeys. Apes emerged within monkeys as sister of the Cercopithecidae in the Catarrhini, so cladistically they are monkeys as well. However, there has been resistance to directly designate apes (and thus humans) as monkeys, so "Old World monkey" may be taken to mean either the Cercopithecoidea (not including apes) or the Catarrhini (including apes). That apes are monkeys was already realized by
Georges-Louis Leclerc, Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French naturalist, mathematician, cosmologist, and encyclopédiste. His works influenced the next two generations of naturalists, including two prominent Fr ...
in the 18th century.
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
placed this group in 1758 together with the tarsiers, in a single genus "''
Simia In his '' Systema Naturae'' of 1758, Carl Linnaeus divided the Order Primates within ''Mammalia'' into four genera: ''Homo'', ''Simia'', ''Lemur'', and ''Vespertilio''. His ''Vespertilio'' included all bats, and has since been moved from Primates ...
''" (sans ''
Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus ''Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely relate ...
''), an ensemble now recognised as the Haplorhini. Monkeys, including apes, can be distinguished from other primates by having only two pectoral nipples, a pendulous penis, and a lack of sensory
whiskers Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to touch, sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are c ...
.


Historical and modern terminology

According to the '' Online Etymology Dictionary'', the word "monkey" may originate in a
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
version of the ''
Reynard the Fox Reynard the Fox is a literary cycle of medieval allegorical Dutch, English, French and German fables. The first extant versions of the cycle date from the second half of the 12th century. The genre was popular throughout the Late Middle Ages, as ...
'' fable, published circa 1580. In this version of the fable, a character named Moneke is the son of Martin the Ape. In English, no clear distinction was originally made between "ape" and "monkey"; thus the 1911 ''
Encyclopædia Britannica The (Latin for "British Encyclopædia") is a general knowledge English-language encyclopaedia. It is published by Encyclopædia Britannica, Inc.; the company has existed since the 18th century, although it has changed ownership various time ...
'' entry for "ape" notes that it is either a synonym for "monkey" or is used to mean a tailless humanlike primate. Colloquially, the terms "monkey" and "ape" are widely used interchangeably. Also, a few monkey species have the word "ape" in their common name, such as the
Barbary ape The Barbary macaque (''Macaca sylvanus''), also known as Barbary ape, is a macaque species native to the Atlas Mountains of Algeria, Libya, Tunisia and Morocco, along with a small introduced population in Gibraltar. It is the type species of the ...
. Later in the first half of the 20th century, the idea developed that there were trends in
primate evolution The evolutionary history of the primates can be traced back 57-85/90 million years. One of the oldest known primate-like mammal species, ''Plesiadapis'', came from North America; another, ''Archicebus'', came from China. Other similar basal prima ...
and that the living members of the order could be arranged in a series, leading through "monkeys" and "apes" to humans. Monkeys thus constituted a "
grade Grade most commonly refers to: * Grade (education), a measurement of a student's performance * Grade, the number of the year a student has reached in a given educational stage * Grade (slope), the steepness of a slope Grade or grading may also ref ...
" on the path to humans and were distinguished from "apes". Scientific classifications are now more often based on
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
groups, that is groups consisting of ''all'' the descendants of a common ancestor. The New World monkeys and the Old World monkeys are each monophyletic groups, but their combination was not, since it excluded hominoids (apes and humans). Thus, the term "monkey" no longer referred to a recognized scientific
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
. The smallest accepted taxon which contains all the monkeys is the infraorder
Simiiformes The simians, anthropoids, or higher primates are an infraorder (Simiiformes ) of primates containing all animals traditionally called monkeys and apes. More precisely, they consist of the parvorders New World monkeys (Platyrrhini) and Catarrhi ...
, or simians. However this also contains the hominoids, so that monkeys are, in terms of currently recognized taxa, non-hominoid simians. Colloquially and pop-culturally, the term is ambiguous and sometimes monkey includes non-human hominoids. In addition, frequent arguments are made for a monophyletic usage of the word "monkey" from the perspective that usage should reflect cladistics. A group of monkeys may be commonly referred to as a tribe or a troop. Two separate groups of primates are referred to as "monkeys":
New World monkey New World monkeys are the five families of primates that are found in the tropical regions of Mexico, Central and South America: Callitrichidae, Cebidae, Aotidae, Pitheciidae, and Atelidae. The five families are ranked together as the Ceboidea ( ...
s (platyrrhines) from South and Central America and Old World monkeys (
catarrhines The parvorder Catarrhini , catarrhine monkeys, Old World anthropoids, or Old World monkeys, consisting of the Cercopithecoidea and apes (Hominoidea). In 1812, Geoffroy grouped those two groups together and established the name Catarrhini, "Old W ...
in the superfamily Cercopithecoidea) from Africa and Asia.
Apes Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its sister g ...
(hominoids)—consisting of
gibbon Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical rainforest from eastern Bangladesh to Northeast India ...
s,
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...
s,
gorilla Gorillas are herbivorous, predominantly ground-dwelling great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four or fi ...
s,
chimpanzees The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative the ...
and
bonobos The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee and less often the dwarf chimpanzee or gracile chimpanzee, is an endangered great ape and one of the two species making up the genus '' Pan,'' the other being the comm ...
, and
human Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
s—are also catarrhines but were classically distinguished from monkeys. Tailless monkeys may be called "apes", incorrectly according to modern usage; thus the tailless
Barbary macaque The Barbary macaque (''Macaca sylvanus''), also known as Barbary ape, is a macaque species native to the Atlas Mountains of Algeria, Libya, Tunisia and Morocco, along with a small introduced population in Gibraltar. It is the type species of the ...
is historically called the "Barbary ape".


Description

As apes have emerged in the monkey group as sister of the old world monkeys, characteristics that describe monkeys are generally shared by apes as well. Williams et al. outlined evolutionary features, including in stem groupings, contrasted against the other primates such as the
tarsier Tarsiers ( ) are haplorhine primates of the family Tarsiidae, which is itself the lone extant family within the infraorder Tarsiiformes. Although the group was once more widespread, all of its species living today are found in Maritime Southeast ...
s and the
lemuriformes Lemuriformes is an infraorder of primate that falls under the suborder Strepsirrhini. It includes the lemurs of Madagascar, as well as the galagos and lorisids of Africa and Asia, although a popular alternative taxonomy places the lorisoids i ...
. Monkeys range in size from the
pygmy marmoset Pygmy marmosets are two species of small New World monkeys in the genus ''Cebuella''. They are native to rainforests of the western Amazon Basin in South America. These primates are notable for being the smallest monkeys in the world, at just ov ...
, which can be as small as with a tail and just over in weight, to the male
mandrill The mandrill (''Mandrillus sphinx'') is a large Old World monkey native to west central Africa. It is one of the most colorful mammals in the world, with red and blue skin on its face and posterior. The species is sexually dimorphic, as males ...
, almost long and weighing up to . Some are
arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
(living in trees) while others live on the
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
; diets differ among the various species but may contain any of the following: fruit, leaves, seeds, nuts, flowers, eggs and small animals (including insects and spiders). Some characteristics are shared among the groups; most New World monkeys have long tails, with those in the Atelidae family being
prehensile Prehensility is the quality of an appendage or organ (anatomy), organ that has Adaptation (biology), adapted for grasping or holding. The word is derived from the Latin term ''prehendere'', meaning "to grasp". The ability to grasp is likely der ...
, while Old World monkeys have non-prehensile tails or no visible tail at all. Old World monkeys have
trichromatic Trichromacy or trichromatism is the possessing of three independent channels for conveying color information, derived from the three different types of cone cells in the eye. Organisms with trichromacy are called trichromats. The normal expl ...
color vision Color vision, a feature of visual perception, is an ability to perceive differences between light composed of different wavelengths (i.e., different spectral power distributions) independently of light intensity. Color perception is a part of ...
like that of humans, while New World monkeys may be trichromatic, dichromatic, or—as in the
owl monkey Night monkeys, also known as owl monkeys or douroucoulis (), are nocturnal New World monkeys of the genus ''Aotus'', the only member of the family Aotidae (). The genus comprises eleven species which are found across Panama and much of South Ame ...
s and
greater galago The greater galagos or thick-tailed bushbabies are three species of strepsirrhine primates. They are classified in the genus ''Otolemur'' in the family Galagidae. Historical classification and species discovery The diversity of galago species h ...
s—
monochromatic A monochrome or monochromatic image, object or color scheme, palette is composed of one color (or lightness, values of one color). Images using only Tint, shade and tone, shades of grey are called grayscale (typically digital) or Black and wh ...
. Although both the New and Old World monkeys, like the apes, have forward-facing eyes, the faces of Old World and New World monkeys look very different, though again, each group shares some features such as the types of noses, cheeks and rumps.


Classification

The following list shows where the various monkey families (bolded) are placed in the classification of living (extant) primates. * Order
Primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
s ** Suborder
Strepsirrhini Strepsirrhini or Strepsirhini (; ) is a Order (biology), suborder of primates that includes the Lemuriformes, lemuriform primates, which consist of the lemurs of Fauna of Madagascar, Madagascar, galagos ("bushbabies") and pottos from Fauna of A ...
:
lemur Lemurs ( ) (from Latin ''lemures'' – ghosts or spirits) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea (), divided into 8 Family (biology), families and consisting of 15 genera and around 100 exist ...
s,
loris Loris is the common name for the strepsirrhine mammals of the subfamily Lorinae (sometimes spelled Lorisinae) in the family Lorisidae. ''Loris'' is one genus in this subfamily and includes the slender lorises, ''Nycticebus'' is the genus contain ...
es, and
galago Galagos , also known as bush babies, or ''nagapies'' (meaning "night monkeys" in Afrikaans), are small nocturnal primates native to continental, sub-Sahara Africa, and make up the family Galagidae (also sometimes called Galagonidae). They ar ...
s ** Suborder
Haplorhini Haplorhini (), the haplorhines (Greek for "simple-nosed") or the "dry-nosed" primates, is a suborder of primates containing the tarsiers and the simians (Simiiformes or anthropoids), as sister of the Strepsirrhini ("moist-nosed"). The name is some ...
:
tarsier Tarsiers ( ) are haplorhine primates of the family Tarsiidae, which is itself the lone extant family within the infraorder Tarsiiformes. Although the group was once more widespread, all of its species living today are found in Maritime Southeast ...
s, monkeys, and
ape Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its siste ...
s *** Infraorder
Tarsiiformes Tarsiiformes are a group of primates that once ranged across Europe, northern Africa, Asia, and North America, but whose extant species are all found in the islands of Southeast Asia. Tarsiers (family Tarsiidae) are the only living members of ...
**** Family
Tarsiidae Tarsiers ( ) are haplorhine primates of the family Tarsiidae, which is itself the lone extant family within the infraorder Tarsiiformes. Although the group was once more widespread, all of its species living today are found in Maritime Southeast ...
: tarsiers *** Infraorder
Simiiformes The simians, anthropoids, or higher primates are an infraorder (Simiiformes ) of primates containing all animals traditionally called monkeys and apes. More precisely, they consist of the parvorders New World monkeys (Platyrrhini) and Catarrhi ...
: simians **** Parvorder Platyrrhini: New World monkeys ***** Family
Callitrichidae The Callitrichidae (also called Arctopitheci or Hapalidae) are a family of New World monkeys, including marmosets, tamarins, and lion tamarins. At times, this group of animals has been regarded as a subfamily, called the Callitrichinae, of the fa ...
:
marmoset The marmosets (), also known as zaris or sagoin, are 22 New World monkey species of the genera ''Callithrix'', ''Cebuella'', ''Callibella'', and ''Mico''. All four genera are part of the biological family Callitrichidae. The term "marmoset" is ...
s and
tamarin The tamarins are squirrel-sized New World monkeys from the family Callitrichidae in the genus ''Saguinus''. They are the first offshoot in the Callitrichidae tree, and therefore are the sister group of a clade formed by the lion tamarins, Goel ...
s (42 species) ***** Family
Cebidae The Cebidae are one of the five families of New World monkeys now recognised. Extant members are the capuchin and squirrel monkeys. These species are found throughout tropical and subtropical South and Central America. Characteristics Cebid mo ...
:
capuchins Capuchin can refer to: *Order of Friars Minor Capuchin, an order of Roman Catholic friars *Capuchin Poor Clares, an order of Roman Catholic contemplative religious sisters *Capuchin monkey, primates of the genus ''Cebus'' and ''Sapajus'', named af ...
and
squirrel monkey Squirrel monkeys are New World monkeys of the genus ''Saimiri''. ''Saimiri'' is the only genus in the subfamily Saimirinae. The name of the genus is of Tupi origin (''sai-mirím'' or ''çai-mbirín'', with ''sai'' meaning 'monkey' and ''mirím'' ...
s (14 species) ***** Family
Aotidae Night monkeys, also known as owl monkeys or douroucoulis (), are nocturnal New World monkeys of the genus ''Aotus'', the only member of the family Aotidae (). The genus comprises eleven species which are found across Panama and much of South Ame ...
:
night monkey Night monkeys, also known as owl monkeys or douroucoulis (), are nocturnal New World monkeys of the genus ''Aotus'', the only member of the family Aotidae (). The genus comprises eleven species which are found across Panama and much of South Ame ...
s (11 species) ***** Family
Pitheciidae The Pitheciidae () are one of the five families of New World monkeys now recognised. Formerly, they were included in the family Atelidae. The family includes the titis, saki monkeys and uakaris. Most species are native to the Amazon region ...
:
titi The titis, or titi monkeys, are New World monkeys of the subfamily Callicebinae, which contains three extant genera: ''Cheracebus'', ''Callicebus'', and ''Plecturocebus.'' This subfamily also contains the extinct genera ''Miocallicebus, Homuncul ...
s, sakis, and
uakari Uakari (, ) is the common name for the New World monkeys of the genus ''Cacajao''. Both the English and scientific names are believed to have originated from indigenous languages. The uakaris are unusual among New World monkeys in that the tail ...
s (41 species) ***** Family
Atelidae The Atelidae are one of the five families of New World monkeys now recognised. It was formerly included in the family Cebidae. Atelids are generally larger monkeys; the family includes the howler, spider, woolly, and woolly spider monkeys (t ...
: howler,
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
, and
woolly monkey The woolly monkeys are the genus ''Lagothrix'' of New World monkeys, usually placed in the family Atelidae. Both species in this genus originate from the rainforests of South America. They have prehensile tails and live in relatively large soci ...
s (24 species) **** Parvorder
Catarrhini The parvorder Catarrhini , catarrhine monkeys, Old World anthropoids, or Old World monkeys, consisting of the Cercopithecoidea and apes (Hominoidea). In 1812, Geoffroy grouped those two groups together and established the name Catarrhini, "Old ...
***** Superfamily
Cercopithecoidea Old World monkey is the common English name for a family of primates known taxonomically as the Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons ...
****** Family
Cercopithecidae Old World monkey is the common English name for a family of primates known taxonomically as the Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons ...
: Old World monkeys (135 species) ***** Superfamily
Hominoidea Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its sister g ...
: apes ****** Family
Hylobatidae Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical rainforest from eastern Bangladesh to Northeast India ...
:
gibbon Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical rainforest from eastern Bangladesh to Northeast India ...
s ("lesser apes") (20 species) ****** Family
Hominidae The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the ea ...
: great apes (including
human Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
s,
gorilla Gorillas are herbivorous, predominantly ground-dwelling great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four or fi ...
s,
chimpanzee The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative th ...
s, and
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...
s) (8 species)


Cladogram with extinct families

Below is a
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
with some extinct monkey families. Generally, extinct non-hominoid simians, including early catarrhines are discussed as monkeys as well as simians or anthropoids, which
cladistically Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived chara ...
means that Hominoidea are monkeys as well, restoring monkeys as a single grouping. It is indicated approximately how many million years ago (Mya) the clades diverged into newer clades. It is thought the New World monkeys started as a drifted "Old World monkey" group from the Old World (probably Africa) to the New World (South America).


Relationship with humans

The many species of monkey have varied relationships with humans. Some are kept as pets, others used as
model organism A model organism (often shortened to model) is a non-human species that is extensively studied to understand particular biological phenomena, with the expectation that discoveries made in the model organism will provide insight into the workin ...
s in laboratories or in space missions. They may be killed in
monkey drive A monkey drive is an operation where large numbers of wild monkeys are rounded up and killed in order to protect agriculture such as crops, planted rice, banana and citrus fruit trees. Monkey drives have been reported in Sierra Leone, where they ...
s (when they threaten agriculture) or used as
service animal Various definitions exist for a service animal. The Americans with Disabilities Act defines the term as "dogs that are individually trained to do work or perform tasks for people with disabilities". Dogs are the most common service animals, havi ...
s for the disabled. In some areas, some species of monkey are considered agricultural
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
, and can cause extensive damage to commercial and subsistence crops. This can have important implications for the conservation of endangered species, which may be subject to persecution. In some instances farmers' perceptions of the damage may exceed the actual damage. Monkeys that have become habituated to human presence in tourist locations may also be considered pests, attacking tourists.


As service animals for disabled people

Some organizations train
capuchin monkey The capuchin monkeys () are New World monkeys of the subfamily Cebinae. They are readily identified as the " organ grinder" monkey, and have been used in many movies and television shows. The range of capuchin monkeys includes some tropical fores ...
s as
service animal Various definitions exist for a service animal. The Americans with Disabilities Act defines the term as "dogs that are individually trained to do work or perform tasks for people with disabilities". Dogs are the most common service animals, havi ...
s to assist
quadriplegic Tetraplegia, also known as quadriplegia, is defined as the dysfunction or loss of motor and/or sensory function in the cervical area of the spinal cord. A loss of motor function can present as either weakness or paralysis leading to partial or t ...
s and other people with severe spinal cord injuries or
mobility impairment A physical disability is a limitation on a person's physical functioning, mobility, dexterity or stamina. Other physical disabilities include impairments which limit other facets of daily living, such as respiratory disorders, blindness, epilepsy ...
s. After being
socialized In sociology, socialization or socialisation (see spelling differences) is the process of internalizing the norms and ideologies of society. Socialization encompasses both learning and teaching and is thus "the means by which social and cultur ...
in a human home as infants, the monkeys undergo extensive training before being placed with disabled people. Around the house, the monkeys assist with daily tasks such as feeding, fetching, manipulating objects, and personal care.
Helper monkey Various definitions exist for a service animal. The Americans with Disabilities Act defines the term as "dogs that are individually trained to do work or perform tasks for people with disabilities". Dogs are the most common service animals, hav ...
s are usually trained in schools by private organizations, taking seven years to train, and are able to serve 25–30 years (two to three times longer than a
guide dog Guide dogs (colloquially known in the US as seeing-eye dogs) are assistance dogs trained to lead blind or visually impaired people around obstacles. Although dogs can be trained to navigate various obstacles, they are red–green colour blin ...
). In 2010, the
U.S. federal government The federal government of the United States (U.S. federal government or U.S. government) is the Federation#Federal governments, national government of the United States, a federal republic located primarily in North America, composed of 50 ...
revised its definition of service animal under the
Americans with Disabilities Act The Americans with Disabilities Act of 1990 or ADA () is a civil rights law that prohibits discrimination based on disability. It affords similar protections against discrimination to Americans with disabilities as the Civil Rights Act of 1964 ...
(ADA). Non-human primates are no longer recognized as service animals under the ADA. The
American Veterinary Medical Association The American Veterinary Medical Association (AVMA), founded in 1863, is a not-for-profit association representing more than 99,500 veterinarians in the US. The AVMA provides information resources, continuing education opportunities, publicatio ...
does not support the use of non-human primates as assistance animals because of
animal welfare Animal welfare is the well-being of non-human animals. Formal standards of animal welfare vary between contexts, but are debated mostly by animal welfare groups, legislators, and academics. Animal welfare science uses measures such as longevity ...
concerns, the potential for serious injury to people, and risks that primates may transfer dangerous diseases to humans.


In experiments

The most common monkey species found in animal research are the
grivet The grivet (''Chlorocebus aethiops'') is an Old World monkey with long white tufts of hair along the sides of its face. Some authorities consider this and all of the members of the genus ''Chlorocebus'' to be a single species, ''Cercopithecus ae ...
, the
rhesus macaque The rhesus macaque (''Macaca mulatta''), colloquially rhesus monkey, is a species of Old World monkey. There are between six and nine recognised subspecies that are split between two groups, the Chinese-derived and the Indian-derived. Generally b ...
, and the
crab-eating macaque The crab-eating macaque (''Macaca fascicularis''), also known as the long-tailed macaque and referred to as the cynomolgus monkey in laboratories, is a cercopithecine primate native to Southeast Asia. A species of macaque, the crab-eating macaqu ...
, which are either wild-caught or purpose-bred. They are used primarily because of their relative ease of handling, their fast reproductive cycle (compared to apes) and their psychological and physical similarity to
human Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
s. Worldwide, it is thought that between 100,000 and 200,000 non-human primates are used in research each year, 64.7% of which are Old World monkeys, and 5.5% New World monkeys. This number makes a very small fraction of all animals used in research. Between 1994 and 2004 the United States has used an average of 54,000 non-human primates, while around 10,000 non-human primates were used in the
European Union The European Union (EU) is a supranational political and economic union of member states that are located primarily in Europe. The union has a total area of and an estimated total population of about 447million. The EU has often been des ...
in 2002.


In space

A number of countries have used monkeys as part of their space exploration programmes, including the United States and France. The first monkey in space was Albert II, who flew in the US-launched
V-2 rocket The V-2 (german: Vergeltungswaffe 2, lit=Retaliation Weapon 2), with the technical name ''Aggregat 4'' (A-4), was the world’s first long-range guided ballistic missile. The missile, powered by a liquid-propellant rocket engine, was developed ...
on June 14, 1949.


As food

Monkey brains Monkey brains is a supposed dish consisting of, at least partially, the brain of some species of monkey or ape. While animal brains have been consumed in various cuisines (e.g. eggs and brains or fried brain sandwiches), there is debate abou ...
are eaten as a delicacy in parts of
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...
, Africa and China. Monkeys are sometimes eaten in parts of Africa, where they can be sold as "
bushmeat Bushmeat is meat from wildlife species that are hunted for human consumption, most often referring to the meat of game in Africa. Bushmeat represents a primary source of animal protein and a cash-earning commodity for inhabitants of humid tropi ...
". In traditional
Islamic dietary laws Islamic dietary laws are dietary laws that Muslims follow. Islamic jurisprudence specifies which foods are '' '' (, "lawful") and which are '' '' (, "unlawful"). The dietary laws are found in the Quran, the holy book of Islam, as well as in coll ...
, the eating of monkeys is forbidden.


Literature

Sun Wukong The Monkey King, also known as Sun Wukong ( zh, t=孫悟空, s=孙悟空, first=t) in Mandarin Chinese, is a legendary mythical figure best known as one of the main characters in the 16th-century Chinese novel ''Journey to the West'' ( zh, ...
(the "Monkey King"), a character who figures prominently in
Chinese mythology Chinese mythology () is mythology that has been passed down in oral form or recorded in literature in the geographic area now known as Greater China. Chinese mythology includes many varied myths from regional and cultural traditions. Much of t ...
, is the protagonist in the classic comic Chinese novel ''
Journey to the West ''Journey to the West'' () is a Chinese novel published in the 16th century during the Ming dynasty and attributed to Wu Cheng'en. It is regarded as one of the greatest Classic Chinese Novels, and has been described as arguably the most popu ...
''. Monkeys are prevalent in numerous books, television programs, and movies. The
television series A television show – or simply TV show – is any content produced for viewing on a television set which can be broadcast via over-the-air, satellite television, satellite, or cable television, cable, excluding breaking news, television adverti ...
''
Monkey Monkey is a common name that may refer to most mammals of the infraorder Simiiformes, also known as the simians. Traditionally, all animals in the group now known as simians are counted as monkeys except the apes, which constitutes an incomple ...
'' and the literary characters
Monsieur Eek ''Monsieur Eek'' is a short novel by American playwright David Ives, intended for ages 9–12. It was first published September 1, 2001 by HarperCollins. Set in 1609, it is about a chimpanzee who gets arrested for being a French spy.
and
Curious George Curious George is a fictional monkey who is the title character of a series of popular children's picture books written by Margret and H. A. Rey. Various media, including films and TV shows, have been based upon the original book series. Geor ...
are all examples. Informally, "monkey" may refer to apes, particularly chimpanzees, gibbons, and gorillas. Author
Terry Pratchett Sir Terence David John Pratchett (28 April 1948 – 12 March 2015) was an English humourist, satirist, and author of fantasy novels, especially comical works. He is best known for his ''Discworld'' series of 41 novels. Pratchett's first nov ...
alludes to this difference in usage in his ''
Discworld ''Discworld'' is a comic fantasy"Humorous Fantasy" in David Pringle, ed., ''The Ultimate Encyclopedia of Fantasy'' (pp.31-33). London, Carlton,2006. book series written by the English author Terry Pratchett, set on the Discworld, a flat pla ...
'' novels, in which the
Librarian A librarian is a person who works professionally in a library providing access to information, and sometimes social or technical programming, or instruction on information literacy to users. The role of the librarian has changed much over time, ...
of the
Unseen University The Unseen University (UU) is a school of wizardry in Terry Pratchett's ''Discworld'' series of fantasy novels. Located in the fictional city of Ankh-Morpork, the UU is staffed by a faculty composed of mostly indolent and inept old wizards. The ...
is an
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...
who gets very violent if referred to as a monkey. Another example is the use of Simians in Chinese poetry. The
winged monkeys Winged monkeys are fictional characters created by American author L. Frank Baum in his children's novel ''The Wonderful Wizard of Oz'' (1900). They are jungle monkeys with bird-like feathered wings. They are most notably remembered from the famo ...
are prominent characters in
L. Frank Baum Lyman Frank Baum (; May 15, 1856 – May 6, 1919) was an American author best known for his children's books, particularly ''The Wonderful Wizard of Oz'' and its sequels. He wrote 14 novels in the ''Oz'' series, plus 41 other novels (not includ ...
's '' Wizard of Oz'' books and in the 1939 film based on Baum's 1900 novel ''
The Wonderful Wizard of Oz ''The Wonderful Wizard of Oz'' is a children's novel written by author L. Frank Baum and illustrated by W. W. Denslow. It is the first novel in the Oz series of books. A Kansas farm girl named Dorothy ends up in the magical Land of Oz after s ...
''.


Religion and worship

Monkey is the symbol of fourth ''
Tirthankara In Jainism, a ''Tirthankara'' (Sanskrit: '; English: literally a 'ford-maker') is a saviour and spiritual teacher of the ''dharma'' (righteous path). The word ''tirthankara'' signifies the founder of a '' tirtha'', which is a fordable passag ...
'' in
Jainism Jainism ( ), also known as Jain Dharma, is an Indian religions, Indian religion. Jainism traces its spiritual ideas and history through the succession of twenty-four tirthankaras (supreme preachers of ''Dharma''), with the first in the current ...
,
Abhinandananatha Abhinandananatha or Abhinandana Swami was the fourth ''Tirthankara'' of the present age ( Avasarpini). He is said to have lived for 50 lakh ''purva''. He was born to King Sanvara and Queen Siddhartha at Ayodhya in the Ikshvaku clan. His birth d ...
.
Hanuman Hanuman (; sa, हनà¥à¤®à¤¾à¤¨, ), also called Anjaneya (), is a Hindu god and a divine '' vanara'' companion of the god Rama. Hanuman is one of the central characters of the Hindu epic ''Ramayana''. He is an ardent devotee of Rama and on ...
, a prominent deity in
Hinduism Hinduism () is an Indian religion or '' dharma'', a religious and universal order or way of life by which followers abide. As a religion, it is the world's third-largest, with over 1.2–1.35 billion followers, or 15–16% of the global p ...
, is a human-like monkey god who is believed to bestow courage, strength and longevity to the person who thinks about him or
Rama Rama (; ), Ram, Raman or Ramar, also known as Ramachandra (; , ), is a major deity in Hinduism. He is the seventh and one of the most popular '' avatars'' of Vishnu. In Rama-centric traditions of Hinduism, he is considered the Supreme Bein ...
. In
Buddhism Buddhism ( , ), also known as Buddha Dharma and Dharmavinaya (), is an Indian religion or philosophical tradition based on teachings attributed to the Buddha. It originated in northern India as a -movement in the 5th century BCE, and gra ...
, the monkey is an early incarnation of Buddha but may also represent trickery and ugliness. The
Chinese Buddhist Chinese Buddhism or Han Buddhism ( zh, s=汉传佛教, t=漢傳佛教, p=Hànchuán Fójiào) is a Chinese form of Mahayana, Mahayana Buddhism which has shaped Chinese culture in a wide variety of areas including Chinese art, art, politics, Ch ...
"
mind monkey The mind is the set of faculties responsible for all mental phenomena. Often the term is also identified with the phenomena themselves. These faculties include thought, imagination, memory, will, and sensation. They are responsible for vario ...
" metaphor refers to the unsettled, restless state of human mind. Monkey is also one of the Three Senseless Creatures, symbolizing greed, with the tiger representing anger and the deer lovesickness. The ''Sanzaru,'' or
three wise monkeys The three wise monkeys are a Japanese pictorial maxim, embodying the proverbial principle "see no evil, hear no evil, speak no evil". The three monkeys are * Mizaru, who sees no evil, covering his eyes * Kikazaru, who hears no evil, covering ...
, are revered in Japanese folklore; together they embody the proverbial principle to "see no evil, hear no evil, speak no evil". The Moche people of ancient Peru worshipped nature. They placed emphasis on animals and often depicted monkeys in their art. The Tzeltal people of Mexico worshipped monkeys as incarnations of their dead ancestors.


Zodiac

The
Monkey Monkey is a common name that may refer to most mammals of the infraorder Simiiformes, also known as the simians. Traditionally, all animals in the group now known as simians are counted as monkeys except the apes, which constitutes an incomple ...
(猴) is the ninth in the twelve-year cycle of animals which appear in the
Chinese zodiac The Chinese zodiac is a traditional classification scheme based on the lunar calendar that assigns an animal and its reputed attributes to each year in a repeating twelve-year cycle. Originating from China, the zodiac and its variations remain ...
related to the
Chinese calendar The traditional Chinese calendar (also known as the Agricultural Calendar ¾²æ›†; 农历; ''Nónglì''; 'farming calendar' Former Calendar ˆŠæ›†; 旧历; ''Jiùlì'' Traditional Calendar €æ›†; è€åŽ†; ''LÇŽolì'', is a lunisolar calendar ...
. .


See also

*
List of New World monkey species New World monkeys are all simian primates. While they are endemic to South and Central America, their ancestors rafted over or traversed via land bridge from Africa across the Atlantic Ocean when it was much narrower than at present. Taxonom ...
*
List of Old World monkey species Old World monkeys are all simian primates. They are more closely related to the apes than they are to the New World monkeys. Taxonomic classification * Infraorder Simian, Simiiformes ** Parvorder Catarrhini *** Superfamily Old World monkey, Cer ...
*
List of individual monkeys This annotated list of individual monkeys includes monkeys who are in some way famous or notable. The list does not include notable apes, or fictional primates. Monkey actors * Binx - (white-headed capuchin) Appeared in '' Ace Ventura: Pet De ...
*
List of fictional primates This list of fictional primates is a subsidiary to the list of fictional animals. The list is restricted to notable non-human primate characters from the world of fiction including chimpanzees, gorillas, orangutans, bonobos, gibbons, monkeys, lemu ...
*
List of primates List of primates contains the extant species in the order Primates and currently contains 16 families and 72 genera. For extinct species see the list of fossil primates. Suborder Strepsirrhini Infraorder Lemuriformes Superfamily Lem ...
*
List of primates by population This is a list of primate species by estimated global population. This list is not comprehensive, as not all primates have had their numbers quantified. See also * Lists of organisms by population *Lists of mammals by population *Human popul ...
*
International Primate Day International Primate Day, September 1, is an annual educational observance event organized since 2005 largely by British-based Animal Defenders International (ADI) and supported annually by various primate-oriented advocacy organizations, speaks ...
*
Monkey Day Monkey Day is an unofficial international holiday celebrated on December 14. The holiday was created and popularized in 2000 by controversial artists Casey Sorrow and Eric Millikin when they were art students at Michigan State University. Monke ...
*
Signifying monkey The signifying monkey is a character of African-American folklore that derives from the trickster figure of Yoruba mythology, Esu Elegbara. This character was transported with Africans to the Americas under the names of Exu, Echu-Elegua, Papa L ...


Notes


References


Literature cited

*


Further reading


"How to Avoid Monkey Bites and Attacks in Southeast Asia"
by Gregory Rodgers, Trip Savvy, 21 Dec 2018

by Anniina Jokinen, Luminarium: Anthology of English Literature
"The Impossible Housing and Handling Conditions of Monkeys in Research Laboratories"
by Viktor Reinhardt, International Primate Protection League, August 2001

an article by veterinarian Lianne McLeod on About.com
Helping Hands: Monkey helpers for the disabled
a U.S. national non-profit organization based in Boston Massachusetts that places specially trained capuchin monkeys with people who are paralyzed or who live with other severe mobility impairments {{Authority control Extant Eocene first appearances Paraphyletic groups