Mohanamico
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''Mohanamico'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
New World monkey New World monkeys are the five families of primates that are found in the tropical regions of Mexico, Central and South America: Callitrichidae, Cebidae, Aotidae, Pitheciidae, and Atelidae. The five families are ranked together as the Ceboidea ( ...
s from the
Middle Miocene The Middle Miocene is a sub-epoch of the Miocene Epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0.05 Ma to 11.608 ± 0.005 Ma (million y ...
(
Laventan The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes ...
in the
South American land mammal age The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Mya (unit), Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referre ...
s; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of
La Venta La Venta is a pre-Columbian archaeological site of the Olmec civilization located in the present-day Mexican state of Tabasco. Some of the artifacts have been moved to the museum "Parque - Museo de La Venta", which is in nearby Villahermosa, ...
in the Honda Group of
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''M. hershkovitzi''. Due to the relatively few material found of ''Mohanamico'', the placement of the genus is not certain and four possible families have been proposed by different authors,
Atelidae The Atelidae are one of the five family (biology), families of New World monkeys now recognised. It was formerly included in the family Cebidae. Atelids are generally larger monkeys; the family includes the howler monkey, howler, spider monkey, ...
,
Callitrichidae The Callitrichidae (also called Arctopitheci or Hapalidae) are a family of New World monkeys, including marmosets, tamarins, and lion tamarins. At times, this group of animals has been regarded as a subfamily, called the Callitrichinae, of the fa ...
,
Pitheciidae The Pitheciidae () are one of the five families of New World monkeys now recognised. Formerly, they were included in the family Atelidae. The family includes the titis, saki monkeys and uakaris. Most species are native to the Amazon region ...
or
Aotidae Night monkeys, also known as owl monkeys or douroucoulis (), are nocturnal New World monkeys of the genus ''Aotus'', the only member of the family Aotidae (). The genus comprises eleven species which are found across Panama and much of South Ame ...
.


Etymology

''Mohanimico hershkovitzi'' is named after the river god Mohan of the
Magdalena River The Magdalena River ( es, Río Magdalena, ; less commonly ) is the main river of Colombia, flowing northward about through the western half of the country. It takes its name from the biblical figure Mary Magdalene. It is navigable through much of ...
, in which valley the fossils were found and to honor
Philip Hershkovitz Philip Hershkovitz (12 October 1909 – 15 February 1997) was an American mammalogy, mammalogist. Born in Pittsburgh, he attended the Universities of Pittsburgh and Michigan and lived in South America collecting mammals. In 1947, he was appointed ...
for his contributions to the study of Colombian and other South American primates.Luchterhand et al., 1986, p.1757


Description

Fossils of ''Mohanamico'' were discovered in the "Monkey Beds" of the Honda Group, that has been dated to the
Laventan The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes ...
, about 12.5 Ma.Wheeler, 2010, p.133 ''Mohanamico'' was about the size of the living
squirrel monkey Squirrel monkeys are New World monkeys of the genus ''Saimiri''. ''Saimiri'' is the only genus in the subfamily Saimirinae. The name of the genus is of Tupi origin (''sai-mirím'' or ''çai-mbirín'', with ''sai'' meaning 'monkey' and ''mirím'' ...
''
Saimiri sciureus The Guianan squirrel monkey (''Saimiri sciureus'') is a species of squirrel monkey from Guiana, Venezuela and Brazil. ''S. sciureus'' formerly applied to Humboldt's squirrel monkey and Collins' squirrel monkey, but genetic research in 2009 an ...
''. Its molars are low-crowned and the molar crests are not pronounced suggesting a frugivorous diet like '' Aotus''. The lateral incisor is large and high-crowned, which foreshadows living
Pitheciidae The Pitheciidae () are one of the five families of New World monkeys now recognised. Formerly, they were included in the family Atelidae. The family includes the titis, saki monkeys and uakaris. Most species are native to the Amazon region ...
. The
canines Canine may refer to: Zoology and anatomy * a dog-like Canid animal in the subfamily Caninae ** ''Canis'', a genus including dogs, wolves, coyotes, and jackals ** Dog, the domestic dog * Canine tooth, in mammalian oral anatomy People with the surn ...
and Pz were large and sharp like ''
Callimico The Goeldi's marmoset or Goeldi's monkey (''Callimico goeldii'') is a small, South American New World monkey that lives in the upper Amazon basin region of Bolivia, Brazil, Colombia, and Peru. It is the only species classified in the genus ''Call ...
''. Analysis of the mandible and teeth suggest that ''Mohanamico'' is a primitive member of the Pitheciidae. Some similarities with ''Callimico'' and ''
Saguinus The tamarins are squirrel-sized New World monkeys from the family Callitrichidae in the genus ''Saguinus''. They are the first offshoot in the Callitrichidae tree, and therefore are the sister group of a clade formed by the lion tamarins, Goeld ...
'' are also noted raising the possibility that pithecines and callitrichids are monophyletic.Luchterhand et al., 1986, p.1753 The estimated weight of ''Mohanamico'' was ,Silvestro, 2017, p.14 similar in size to ''
Aotus dindensis ''Aotus dindensis'' is an extinct species of New World monkeys in the genus '' Aotus'' from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La ...
'' found in the same location.Gebo et al., 1990, p.738 The
Argentinian Argentines (mistakenly translated Argentineans in the past; in Spanish (masculine) or ( feminine)) are people identified with the country of Argentina. This connection may be residential, legal, historical or cultural. For most Argentines, ...
genus ''
Homunculus A homunculus ( , , ; "little person") is a representation of a small human being, originally depicted as small statues made out of clay. Popularized in sixteenth-century alchemy and nineteenth-century fiction, it has historically referred to the ...
'' had smaller molars than ''Mohanamico''.Luchterhand et al., 1986, p.1756 Some authors place ''Mohanamico'' in the
Callitrichidae The Callitrichidae (also called Arctopitheci or Hapalidae) are a family of New World monkeys, including marmosets, tamarins, and lion tamarins. At times, this group of animals has been regarded as a subfamily, called the Callitrichinae, of the fa ...
,Tejedor, 2013, p.29 possibly related to the ''
Callimico The Goeldi's marmoset or Goeldi's monkey (''Callimico goeldii'') is a small, South American New World monkey that lives in the upper Amazon basin region of Bolivia, Brazil, Colombia, and Peru. It is the only species classified in the genus ''Call ...
'' clade, based especially on its taller incisors and canines, large p2, and broader and longer trigonid in proportion to the talonid.Pérez et al., 2013, p.4 Other authors dispute this and consider the genus more likely a
Pitheciidae The Pitheciidae () are one of the five families of New World monkeys now recognised. Formerly, they were included in the family Atelidae. The family includes the titis, saki monkeys and uakaris. Most species are native to the Amazon region ...
,Wheeler, 2010, p.134 or possibly an
Aotidae Night monkeys, also known as owl monkeys or douroucoulis (), are nocturnal New World monkeys of the genus ''Aotus'', the only member of the family Aotidae (). The genus comprises eleven species which are found across Panama and much of South Ame ...
.Takai et al., 2001, p.303 It is also shown to be a stem atelid. If placed within the Pitheciidae, the primate would be at the base of the evolutionary radiation because of the large
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
s and the structure of its canines and
premolar The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mouth ...
s.Defler, 2004, p.33


Habitat

The Honda Group, and more precisely the "Monkey Beds", are the richest site for fossil primates in South America.Rosenberger & Hartwig, 2001, p.3 It has been argued that the monkeys of the Honda Group were living in habitat that was in contact with the
Amazon Amazon most often refers to: * Amazons, a tribe of female warriors in Greek mythology * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon River, in South America * Amazon (company), an American multinational technology c ...
and
Orinoco Basin The Orinoco Basin is the part of South America drained by the Orinoco river and its tributaries. The Orinoco watershed covers an area of about 990000 km2, making it the third largest in South America, covering most of Venezuela and eastern p ...
s, and that La Venta itself was probably seasonally dry forest.Lynch Alfaro et al., 2015, p.520 From the same level as where ''Mohanamico'' has been found, also fossils of ''
Cebupithecia ''Cebupithecia'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of ...
'', ''
Micodon ''Micodon'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of Colomb ...
'', ''
Saimiri annectens ''Saimiri annectens'', originally described as ''Laventiana annectens'' and later as ''Neosaimiri annectens'', is an extinct species of New World monkey in the genus ''Saimiri'' (squirrel monkeys) from the Middle Miocene (Laventan in the South Am ...
'', ''
Saimiri fieldsi ''Saimiri fieldsi'' is an extinct species of New World monkey in the genus ''squirrel monkey, Saimiri'' (squirrel monkeys) from the Miocene, Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been ...
'' and '' Stirtonia'' have been uncovered.Setoguchi et al., 1986, p.762 From the same location, a fossil specimen of the fish tambaqui (''Colossoma macropomum'') was recovered.


See also

*
List of primates of Colombia The primates of Colombia include 41 extant species in 13 genera and five families. Additionally, 12 fossil species in 10 genera and five families have been identified in Colombia, mainly at the La Venta (Colombia), La Venta Lagerstätte of the Hond ...
* ''
Canaanimico ''Canaanimico'' is an extinct genus of medium-sized New World monkeys from the Late Oligocene (approximately 26.5 Ma, Deseadan in the SALMA classification) fossiliferous fluvio-lacustrine Chambira Formation of the Ucayali Basin in Amazonian ...
'' * ''
Lagonimico ''Lagonimico'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of Col ...
'' * ''
Patasola The Patasola or "one leg" is one of many legends in South American folklore about female monsters from the jungle, appearing to male hunters or loggers in the middle of the wilderness when they think about women. The Patasola appears in the form ...
''


References


Bibliography

* * * * * * * * * *


Further reading

* * * {{Taxonbar, from=Q10331497 Prehistoric primate genera Prehistoric monkeys †Mohanamico Miocene genus first appearances Miocene extinctions Miocene primates of South America Laventan Neogene Colombia Fossils of Colombia Honda Group, Colombia Fossil taxa described in 1986