Milleporidae

Fire corals (''Millepora'') are a genus of colonial marine organisms that exhibit physical characteristics similar to that of coral. The name coral is somewhat misleading, as fire corals are not true corals but are instead more closely related to ''Hydra'' and other hydrozoans, making them hydrocorals. They make up the only genus in the monotypic family Milleporidae.

Distinguishing characteristics

Fire corals have a bright, yellow-green to brown skeletal covering, and are widely distributed in tropical and subtropical waters. They appear in small, brush-like growths on rocks and coral. Divers often mistake fire coral for seaweed, and accidental contact is common. Upon contact, an intense pain can be felt, lasting from two days to two weeks. Occasional relapses of post-treatment inflammation are common. Prominent side effects can include skin irritation, stinging or burning pain, erythema (skin redness), fever, and/or urticarian (hives) lesions. These side effects are due to venom released from the nematocyte, as venom is part of the defense mechanism of the fire coral. Despite its mild to moderate potential for pain, the venom is nonlethal to humans. The very small nematocysts on fire corals contain tentacles, protruding from numerous surface pores (similar to jellyfish stingers). In addition, fire corals have a sharp, calcified external skeleton that can scrape the skin.

Fire coral has several common growth forms; these include branching, plate, and encrusting. Branching fire coral adopts a calcareous structure which branches off into rounded, finger-like tips. Plate-growing fire coral forms a shape similar to that of fellow cnidarian lettuce corals - erect, thin sheets, which group together to form a colony. In encrusting fire coral, growth takes place on the surface structure of calcareous coral or gorgonian structures.

The gonophores in the family Milleporidae arise from the coenosarc (the hollow living tubes of the upright branching individuals of a colony), within chambers embedded entirely in the coenosteum (the calcareous mass forming the skeleton of a compound coral).

Distribution and habitat

Fire corals are found on reefs in tropical and subtropical waters, such as the Indian Ocean, Pacific Ocean, and Atlantic Ocean and the Caribbean Sea.Veron, J.E.N. (2000) ''Corals of the World.'' Vol. 3. Australian Institute of Marine Sciences, Townsville, Australia. The highest densities of fire coral are found in the marine waters of South and Southeast Asia; from India in the west to as far east as French Polynesia; south to coastal northern Australia, and north to Okinawa and extreme southern Japan. They are common in all coastal regions of the Arabian Peninsula and the Persian Gulf, as well as off of South Africa, and up the entire eastern side of the African continent; Madagascar, Seychelles, Mauritius, the Maldives, Cape Verde and the Galápagos Islands support known populations of fire coral. Additionally, they are found throughout the Caribbean, along Brazil’s Atlantic coast (near the equator), and both the Pacific and Atlantic coasts of Panamá. Fire corals form extensive outcrops on projecting parts of reefs where the tidal currents are strongest. They are also abundant on upper reef slopes and in lagoons, and occur down to deep.

Biology

The polyps of fire corals are near microscopic size and are mostly embedded in the skeleton and connected by a network of minute canals. All that is visible on the smooth surface are pores of two sizes: gastropores and dactylopores. In fact, Millepora means ‘thousand pores’. Dactylozooids have long fine hairs that protrude from the skeleton. The hairs possess clusters of stinging cells and capture prey, which is then engulfed by gastrozooid