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:''Common names: Müller's black-headed snake, Mueller's two-headed snake, Muller's snake.'' ''Micrelaps muelleri'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
rear-fanged snake in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Lamprophiidae The Lamprophiidae are a family of snakes found throughout much of Africa, including the Seychelles. There are 89 species as of July 2022. Biology Lamprophiids are a very diverse group of snakes. Many are terrestrial but some are fossorial (e.g. ...
. The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabia (including the Arabian Peninsula and Bahrain), Asia Minor (Asian part of Turkey except Hatay Province), East Thrace (Europ ...
.


Taxonomy

There are no subspecies of ''M. muelleri'' that are recognized as being valid. ''M. muelleri'' is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
for the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
'' Micrelaps''.


Etymology

The specific name, ''muelleri'', is in honor of Swiss
herpetologist Herpetology (from Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is the branch of zoology concerned with the study of amphibians (including frogs, toads, salamanders, newts, and caecilians ( gymnophiona)) and rep ...
Fritz Müller Johann Friedrich Theodor Müller (31 March 1822 – 21 May 1897), better known as Fritz Müller, and also as Müller-Desterro, was a German biologist who emigrated to southern Brazil, where he lived in and near the German community of Blumenau, ...
.Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Micrelaps muelleri'', p. 184).


Geographic range

''Micrelaps muelleri'' is found in
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
, Palestine,
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
,
Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to Lebanon–Syria border, the north and east and Israel to Blue ...
, and Syria.


Description

''Micrelaps muelleri'' is black, with whitish zigzag-edged rings, which may be narrower or wider than the spaces between them, mostly interrupted ventrally. Adults may attain a total length of , which includes a tail long. The
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...
are smooth, without apical pits, and arranged in 15 rows at midbody. The
ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
number 251–275. The
anal plate Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involving s ...
is divided. The
subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail.Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . These scales may be either single or divided (pair ...
, which are also divided, number 26–32. The head is very flattened. The rostral is nearly twice as broad as deep, just visible from above. The
internasals In snakes, the internasal scales are those on top of the head between the scales that surround the nostrils. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: Comstock Publishing Associates. (7 ...
are a little broader than long, shorter than the prefrontals. The
frontal Front may refer to: Arts, entertainment, and media Films * ''The Front'' (1943 film), a 1943 Soviet drama film * ''The Front'', 1976 film Music * The Front (band), an American rock band signed to Columbia Records and active in the 1980s and e ...
is small, not broader than the supraocular, 1⅔ times as long as broad, hardly as long as its distance from the rostral, half as long as the parietals. Each supraocular is as long as broad. A small postocular is in contact with the first temporal. The temporals are arranged 1+1 or 1+2. There are seven
upper labials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ...
, the third and fourth entering the eye. There are three or four lower labials in contact with the anterior
chin shield Chin shields or chinshields, genials scales on a snake are scales found on the underside of the snake's head towards the anterior and touching the lower labial scales. Chin shields to the front of the snake (towards the snout) are called anterior c ...
. The anterior chin shields are as long as the posterior chin shields. Boulenger GA (1896). ''Catalogue of the Snakes in the British Museum (Natural History). Volume III., Containing the Colubridæ (Opisthoglyphæ and Proteroglyphæ), Amblycephalidæ, and Viperidæ''. London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiv + 727 pp. + Plates I-XXV. (''Micrelaps muelleri'', p. 249).


Reproduction

''Micrelaps muelleri'' is oviparous.


Habitat

The preferred
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s of ''M. muelleri'' include
Mediterranean-type shrubby vegetation Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in ...
, and arable land.


Conservation status

''Micrelaps muelleri'' is classified as "
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
" (LC) according to the IUCN Red List of Threatened Species. Although its range is relatively restricted, there are not believed to be major threats to it. It is present in some protected areas.


References


Further reading

Böttger O (1880). "''Die Reptilien und Amphibien von Syrien, Palaestina und Cypern'' ". ''Bericht über die Senckenbergische naturforschende Gesellschaft in Frankurt am Main'' 1880: 132-219. (''Micrelaps muelleri'', new species, pp. 137–138). (In
German German(s) may refer to: * Germany (of or related to) ** Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...
and
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
). {{Taxonbar, from=Q1517893 Atractaspididae Snakes of Asia Snakes of Jordan Reptiles of Syria Taxa named by Oskar Boettger Reptiles described in 1880 Taxonomy articles created by Polbot