The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last
era
An era is a span of time defined for the purposes of chronology or historiography, as in the regnal eras in the history of a given monarchy, a calendar era used for a given calendar, or the geological eras defined for the history of Earth.
Compa ...
of Earth's
geological history, lasting from about , comprising the
Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
,
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
and
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
Periods. It is characterized by the dominance of
archosaurian
Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avi ...
reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s, like the
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s; an abundance of
conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
s and
fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s; a hot
greenhouse
A greenhouse (also called a glasshouse, or, if with sufficient heating, a hothouse) is a structure with walls and roof made chiefly of Transparent ceramics, transparent material, such as glass, in which plants requiring regulated climatic condit ...
climate; and the
tectonic
Tectonics (; ) are the processes that control the structure and properties of the Earth's crust and its evolution through time. These include the processes of mountain building, the growth and behavior of the strong, old cores of continents k ...
break-up of
Pangaea. The Mesozoic is the middle of the three eras since
complex life evolved: the
Paleozoic
The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon.
The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838
by combining the Greek words ''palaiós'' (, "old") and ' ...
, the Mesozoic, and the
Cenozoic
The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
.
The era began in the wake of the
Permian–Triassic extinction event
The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as ...
, the largest well-documented
mass extinction
An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It ...
in Earth's history, and ended with the
Cretaceous–Paleogene extinction event
The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
, another mass extinction whose victims included the
non-avian dinosaurs,
pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
,
mosasaurs
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the ...
, and
plesiosaurs. The Mesozoic was a time of significant tectonic, climatic, and evolutionary activity. The era witnessed the gradual rifting of the supercontinent Pangaea into separate landmasses that would move into their current positions during the next era. The climate of the Mesozoic was varied, alternating between warming and cooling periods. Overall, however, the Earth was hotter than it is today. Dinosaurs first appeared in the Mid-Triassic, and became the dominant terrestrial vertebrates in the Late Triassic or Early Jurassic, occupying this position for about 150 or 135 million years until their demise at the end of the Cretaceous. Archaic
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s appeared in the Jurassic, having
evolved
Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...
from a branch of
theropod dinosaurs, then
true toothless birds appeared in the Cretaceous. The first
mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s also appeared during the Mesozoic, but would remain small—less than 15 kg (33 lb)—until the Cenozoic. The
flowering plants
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
appeared in the early Cretaceous Period and would rapidly diversify throughout the end of the era, replacing conifers and other gymnosperms as the dominant group of plants.
Naming
The phrase "Age of Reptiles" was introduced by the 19th century
paleontologist
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
Gideon Mantell who viewed it as dominated by
diapsids such as ''
Iguanodon'', ''
Megalosaurus'', ''
Plesiosaurus'', and ''
Pterodactylus''.
The current name was proposed in 1840 by the British geologist
John Phillips (1800–1874). "Mesozoic" literally means 'middle life', deriving from the
Greek prefix ( 'between') and ( 'animal, living being').
In this way, the Mesozoic is comparable to the Cenozoic () and Paleozoic ('old life') Eras as well as the
Proterozoic
The Proterozoic () is a geological eon spanning the time interval from 2500 to 538.8million years ago. It is the most recent part of the Precambrian "supereon". It is also the longest eon of the Earth's geologic time scale, and it is subdivided ...
('earlier life') Eon.
The Mesozoic Era was originally described as the "secondary" era, following the "primary" (
Paleozoic
The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon.
The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838
by combining the Greek words ''palaiós'' (, "old") and ' ...
), and preceding the
Tertiary
Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago.
The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
.
Geologic periods
Following the Paleozoic, the Mesozoic extended roughly 186 million years, from when the
Cenozoic
The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
Era began. This time frame is separated into three geologic
periods. From oldest to youngest:
*
Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
()
*
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
()
*
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
()
The lower boundary of the Mesozoic is set by the
Permian–Triassic extinction event
The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as ...
, during which it has been estimated that up to 90-96% of marine species became
extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
although those approximations have been brought into question with some paleontologists estimating the actual numbers as low as 81%.
It is also known as the "Great Dying" because it is considered the largest mass extinction in the Earth's history. The upper boundary of the Mesozoic is set at the
Cretaceous–Paleogene extinction event
The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
(or K–Pg extinction event), which may have been caused by an asteroid impactor that created
Chicxulub Crater
The Chicxulub crater () is an impact crater buried underneath the Yucatán Peninsula in Mexico. Its center is offshore near the community of Chicxulub, after which it is named. It was formed slightly over 66 million years ago when a large ast ...
on the
Yucatán Peninsula. Towards the Late Cretaceous, large volcanic eruptions are also believed to have contributed to the Cretaceous–Paleogene extinction event. Approximately 50% of all genera became extinct, including all of the non-
avian
Avian may refer to:
*Birds or Aves, winged animals
*Avian (given name) (russian: Авиа́н, link=no), a male forename
Aviation
*Avro Avian, a series of light aircraft made by Avro in the 1920s and 1930s
*Avian Limited, a hang glider manufacture ...
dinosaurs.
Triassic
The Triassic ranges roughly from 252 million to 201 million years ago, preceding the Jurassic Period. The period is bracketed between the Permian–Triassic extinction event and the
Triassic–Jurassic extinction event, two of the "
big five", and it is divided into three major epochs: Early, Middle, and Late Triassic.
The Early Triassic, about 252 to 247 million years ago, was dominated by deserts in the interior of the Pangaea supercontinent. The Earth had just witnessed a massive die-off in which 95% of all life became extinct, and the most common vertebrate life on land were ''
Lystrosaurus
''Lystrosaurus'' (; 'shovel lizard'; proper Greek is λίστρον ''lístron'' ‘tool for leveling or smoothing, shovel, spade, hoe’) is an extinct genus of herbivorous dicynodont therapsids from the late Permian and Early Triassic epochs (a ...
'',
labyrinthodont
"Labyrinthodontia" (Greek, 'maze-toothed') is an informal grouping of extinct predatory amphibians which were major components of ecosystems in the late Paleozoic and early Mesozoic eras (about 390 to 150 million years ago). Traditionally consid ...
s, and ''
Euparkeria'' along with many other creatures that managed to survive the Permian extinction.
Temnospondyl
Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order (biology), order of small to giant tetrapods—often considered Labyrinthodontia, primitive amphi ...
s reached peak diversity during the early Triassic.
![Sellosaurus](https://upload.wikimedia.org/wikipedia/commons/9/99/Sellosaurus.jpg)
The Middle Triassic, from 247 to 237 million years ago, featured the beginnings of the breakup of Pangaea and the opening of the
Tethys Ocean. Ecosystems had recovered from the Permian extinction. Algae, sponge, corals, and crustaceans all had recovered, and new aquatic reptiles evolved, such as
ichthyosaur
Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, altho ...
s and
nothosaur
Nothosaurs (order Nothosauroidea) were Triassic marine sauropterygian reptiles that may have lived like seals of today, catching food in water but coming ashore on rocks and beaches. They averaged about in length, with a long body and tail.F. v. ...
s. On land, pine forests flourished, as did groups of insects like mosquitoes and fruit flies. Reptiles began to get bigger and bigger, and the first crocodilians and dinosaurs evolved, which sparked competition with the large amphibians that had previously ruled the freshwater world, respectively mammal-like reptiles on land.
Following the bloom of the Middle Triassic, the Late Triassic, from 237 to 201 million years ago, featured frequent heat spells and moderate precipitation (10–20 inches per year). The recent warming led to a boom of dinosaurian evolution on land as those one began to separate from each other (Nyasasaurus from 243 to 210 million years ago, approximately 235–30 ma, some of them separated into Sauropodomorphs, Theropods and Herrerasaurids), as well as the first
pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
s. During the Late Triassic, some advanced
cynodont
The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s gave rise to the first
Mammaliaformes
Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest Extinction, extinct relatives; the group adaptive radiation, radiated from earlier probainognathian cynodonts. It is defined as the clade origin ...
. All this climatic change, however, resulted in a large die-out known as the Triassic–Jurassic extinction event, in which many
archosaurs (excluding pterosaurs, dinosaurs and
crocodylomorph
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction.
During Mesozoic and early Cenozoic times, cro ...
s), most
synapsids, and almost all large amphibians became extinct, as well as 34% of marine life, in the Earth's fourth mass extinction event. The cause is debatable;
flood basalt eruptions at the
Central Atlantic magmatic province is cited as one possible cause.
Jurassic
![Sericipterus NT](https://upload.wikimedia.org/wikipedia/commons/9/9b/Sericipterus_NT.jpg)
The Jurassic ranges from 200 million years to 145 million years ago and features three major epochs: The Early Jurassic, the Middle Jurassic, and the Late Jurassic.
The Early Jurassic spans from 200 to 175 million years ago.
The climate was tropical and much more humid than the Triassic, as a result of the large seas appearing between the land masses. In the oceans,
plesiosaurs, ichthyosaurs and
ammonites
Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
were abundant. On land, dinosaurs and other archosaurs staked their claim as the dominant race, with
theropods such as ''
Dilophosaurus'' at the top of the food chain. The first true crocodiles evolved, pushing the large amphibians to near extinction. All-in-all, archosaurs rose to rule the world. Meanwhile, the first true mammals evolved, remaining relatively small but spreading widely; the Jurassic ''
Castorocauda
''Castorocauda'' is an extinct, semi-aquatic, superficially otter-like genus of docodont mammaliaforms with one species, ''C. lutrasimilis''. It is part of the Yanliao Biota, found in the Daohugou Beds of Inner Mongolia, China dating to the M ...
'', for example, had adaptations for swimming, digging and catching fish. ''
Fruitafossor
''Fruitafossor'' was a termite-eating mammal endemic to North America during the Late Jurassic epoch (around 150 mya).
The description is based on a complete skeleton of a chipmunk-sized animal.
It was discovered on March 31, 2005, in Fruita, ...
'', from the late Jurassic Period about 150 million years ago, was about the size of a chipmunk, and its teeth, forelimbs and back suggest that it dug open the nests of social insects (probably
termite
Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
s, as ants had not yet appeared) ; ''
Volaticotherium
''Volaticotherium antiquum'' (meaning "ancient gliding beast") is an extinct, gliding, insectivorous mammal that lived in Asia during the Jurassic period, around 164 mya. It is the only member of the genus ''Volaticotherium''.
The discovery of ...
'' was able to glide for short distances, like modern
flying squirrel
Flying squirrels (scientifically known as Pteromyini or Petauristini) are a tribe of 50 species of squirrels in the family Sciuridae. Despite their name, they are not in fact capable of full flight in the same way as birds or bats, but they ar ...
s. The first
multituberculates like ''
Rugosodon'' evolved.
The Middle Jurassic spans from 175 to 163 million years ago.
During this epoch, dinosaurs flourished as huge herds of sauropods, such as ''
Brachiosaurus
''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154to 150million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in th ...
'' and ''
Diplodocus
''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek διπ ...
'', filled the fern prairies, chased by many new predators such as ''
Allosaurus''.
Conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
forests made up a large portion of the forests. In the oceans, plesiosaurs were quite common, and ichthyosaurs flourished. This epoch was the peak of the reptiles.
![Stegosaurus BW](https://upload.wikimedia.org/wikipedia/commons/7/70/Stegosaurus_BW.jpg)
The Late Jurassic spans from 163 to 145 million years ago.
During this epoch, the first
avialans, like ''
Archaeopteryx
''Archaeopteryx'' (; ), sometimes referred to by its German name, "" ( ''Primeval Bird''), is a genus of bird-like dinosaurs. The name derives from the ancient Greek (''archaīos''), meaning "ancient", and (''ptéryx''), meaning "feather" ...
'', evolved from small
coelurosaurian
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrann ...
dinosaurs. The increase in sea levels opened up the Atlantic seaway, which has grown continually larger until today. The further separation of the continents gave opportunity for the diversification of new dinosaurs.
Cretaceous
The Cretaceous is the longest period of the Mesozoic, but has only two epochs: Early and Late Cretaceous.
![Knight Tylosaurus](https://upload.wikimedia.org/wikipedia/commons/f/f7/Knight_Tylosaurus.jpg)
The Early Cretaceous spans from 145 to 100 million years ago.
The Early Cretaceous saw the expansion of seaways and a decline in diversity of sauropods, stegosaurs, and other high-browsing groups, with sauropods particularly scarce in North America.
Seasons came back into effect and the poles got seasonally colder, but some dinosaurs still inhabited the polar forests year round, such as ''
Leaellynasaura'' and ''
Muttaburrasaurus
''Muttaburrasaurus'' was a genus of herbivorous iguanodontian ornithopod dinosaur, which lived in what is now northeastern Australia sometime between 110 and 103 million years agoHoltz, Thomas R. Jr. (2012) ''Dinosaurs: The Most Complete, Up-to-D ...
''. The poles were too cold for crocodiles, and became the last stronghold for large amphibians like ''
Koolasuchus
''Koolasuchus'' is an extinct genus of brachyopoid temnospondyl in the family Chigutisauridae. Fossils have been found from Victoria, Australia and date back 120 Ma to the Aptian stage of the Early Cretaceous. ''Koolasuchus'' is the youngest kno ...
''. Pterosaurs got larger as genera like ''
Tapejara'' and ''
Ornithocheirus'' evolved. Mammals continued to expand their range:
eutriconodont
Eutriconodonta is an order of early mammals. Eutriconodonts existed in Asia, Africa, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 as a replacement name for the ...
s produced fairly large,
wolverine
The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for "gluttony, glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is ...
-like predators like ''
Repenomamus
''Repenomamus'' (Latin: "reptile" (reptilis), "mammal" (mammalis)) is a genus of opossum- to badger-sized gobiconodontid mammal containing two species, ''Repenomamus robustus'' and ''Repenomamus giganticus''. Both species are known from fossils f ...
'' and ''
Gobiconodon'', early
theria
Theria (; Greek: , wild beast) is a subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the placental mammals) and the metatherians (including the marsupials) but excludes the egg-laying monotremes.
Ch ...
ns began to expand into
metatheria
Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well ...
ns and
eutherians, and
cimolodont
Cimolodonta is a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order (biology), order Multituberculata. They probably lived something of a rodent-like existence u ...
multituberculates went on to become common in the fossil record.
The Late Cretaceous spans from 100 to 66 million years ago. The Late Cretaceous featured a cooling trend that would continue in the
Cenozoic
The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
Era. Eventually, tropics were restricted to the equator and areas beyond the tropic lines experienced extreme seasonal changes in weather. Dinosaurs still thrived, as new taxa such as ''
Tyrannosaurus
''Tyrannosaurus'' is a genus of large theropoda, theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosa ...
'', ''
Ankylosaurus
''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the n ...
'', ''
Triceratops
''Triceratops'' ( ; ) is a genus of herbivore, herbivorous Chasmosaurinae, chasmosaurine Ceratopsidae, ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous Period (geology), period, about 68 m ...
'' and
hadrosaurs dominated the food web. In the oceans,
mosasaurs
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the ...
ruled, filling the role of the ichthyosaurs, which, after declining, had disappeared in the
Cenomanian-Turonian boundary event. Though
pliosaurs had gone extinct in the same event, long-necked plesiosaurs such as ''
Elasmosaurus
''Elasmosaurus'' (;) is a genus of plesiosaur that lived in North America during the Campanian stage of the Late Cretaceous period, about 80.5million years ago. The first specimen was discovered in 1867 near Fort Wallace, Kansas, US, and was se ...
'' continued to thrive. Flowering plants, possibly appearing as far back as the Triassic, became truly dominant for the first time. Pterosaurs in the Late Cretaceous declined for poorly understood reasons, though this might be due to tendencies of the fossil record, as their diversity seems to be much higher than previously thought. Birds became increasingly common and diversified into a variety of
enantiornithe
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cl ...
and
ornithurine
Ornithurae (meaning "bird tails" in Greek) is a natural group which includes the common ancestor of ''Ichthyornis'', ''Hesperornis'', and all modern birds as well as all other descendants of that common ancestor.
Classification
Ernst Haeckel coi ...
forms. Though mostly small, marine
hesperornithes became relatively large and flightless, adapted to life in the open sea. Metatherians and primitive eutherian also became common and even produced large and specialised genera like ''
Didelphodon
''Didelphodon'' (from ''is''/nowiki>.html" ;"title="/nowiki>''is''/nowiki>">/nowiki>''is''/nowiki> "opossum" plus "tooth") is a genus of stagodont metatherians from the Late Cretaceous of North America.
Description
Although perhaps little la ...
'' and ''
Schowalteria
''Schowalteria'' is a genus of extinct mammal from the Cretaceous of Canada. It is the earliest known representative of Taeniodonta, a specialised lineage of non-placental eutherian mammals otherwise found in Paleocene and Eocene deposits. It is ...
''. Still, the dominant mammals were multituberculates,
cimolodont
Cimolodonta is a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order (biology), order Multituberculata. They probably lived something of a rodent-like existence u ...
s in the north and
gondwanatheres in the south. At the end of the Cretaceous, the
Deccan traps
The Deccan Traps is a large igneous province of west-central India (17–24°N, 73–74°E). It is one of the largest volcanic features on Earth, taking the form of a large shield volcano. It consists of numerous layers of solidified flood ...
and other volcanic eruptions were poisoning the atmosphere. As this continued, it is thought that a large meteor smashed into earth 66 million years ago, creating the Chicxulub Crater in an event known as the
K-Pg Extinction (formerly K-T), the fifth and most recent mass extinction event, in which 75% of life became extinct, including all non-avian dinosaurs.
Paleogeography and tectonics
Compared to the vigorous convergent plate
mountain-building of the late Paleozoic, Mesozoic tectonic deformation was comparatively mild. The sole major Mesozoic orogeny occurred in what is now the
Arctic
The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
, creating the
Innuitian orogeny
The Innuitian orogeny, sometimes called the Ellesmere orogeny, was a major tectonic (mountain building) episode responsible for the formation of a series of mountain ranges in the Canadian Arctic and Northernmost Greenland. The episode started wit ...
, the
Brooks Range
The Brooks Range ( Gwich'in: ''Gwazhał'') is a mountain range in far northern North America stretching some from west to east across northern Alaska into Canada's Yukon Territory. Reaching a peak elevation of on Mount Isto, the range is believ ...
, the
Verkhoyansk and
Cherskiy Range
The Chersky Range (, ) is a chain of mountains in northeastern Siberia between the Yana River and the Indigirka River. Administratively the area of the range belongs to the Sakha Republic, although a small section in the east is within Magadan ...
s in Siberia, and the
Khingan Mountains
The Greater Khingan Range or Da Hinggan Range (; IPA: ), is a -long volcanic mountain range in the Inner Mongolia region of Northeast China.
It was originally called the Xianbei Mountains, which later became the name of the northern branch of th ...
in Manchuria.
This orogeny was related to the opening of the
Arctic Ocean
The Arctic Ocean is the smallest and shallowest of the world's five major oceans. It spans an area of approximately and is known as the coldest of all the oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, a ...
and
suturing
A surgical suture, also known as a stitch or stitches, is a medical device used to hold body tissues together and approximate wound edges after an injury or surgery. Application generally involves using a needle with an attached length of threa ...
of the
North China
North China, or Huabei () is a List of regions of China, geographical region of China, consisting of the provinces of Beijing, Tianjin, Hebei, Shanxi and Inner Mongolia. Part of the larger region of Northern China (''Beifang''), it lies north ...
and Siberian cratons to Asia. In contrast, the era featured the dramatic rifting of the supercontinent Pangaea, which gradually split into a northern continent,
Laurasia
Laurasia () was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around ( Mya), the other being Gondwana. It separated from Gondwana (beginning in the late Triassic period) during the breakup of Pan ...
, and a southern continent,
Gondwana
Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
. This created the
passive continental margin
A continental margin is the outer edge of continental crust abutting oceanic crust under coastal waters. It is one of the three major zones of the ocean floor, the other two being deep-ocean basins and mid-ocean ridges. The continental margin ...
that characterizes most of the
Atlantic
The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
coastline (such as along the
U.S. East Coast
The East Coast of the United States, also known as the Eastern Seaboard, the Atlantic Coast, and the Atlantic Seaboard, is the coastline along which the Eastern United States meets the Atlantic Ocean, North Atlantic Ocean. The eastern seaboard ...
) today.
By the end of the era, the continents had rifted into nearly their present forms, though not their present positions.
Laurasia
Laurasia () was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around ( Mya), the other being Gondwana. It separated from Gondwana (beginning in the late Triassic period) during the breakup of Pan ...
became
North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
and
Eurasia
Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
, while
Gondwana
Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
split into
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
,
Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
,
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
and the
Indian subcontinent
The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
, which collided with the
Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
n plate during the Cenozoic, giving rise to the
Himalaya
The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
s.
Climate
The Triassic was generally dry, a trend that began in the late
Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
, and highly seasonal, especially in the interior of Pangaea. Low sea levels may have also exacerbated temperature extremes. With its high
specific heat capacity
In thermodynamics, the specific heat capacity (symbol ) of a substance is the heat capacity of a sample of the substance divided by the mass of the sample, also sometimes referred to as massic heat capacity. Informally, it is the amount of heat t ...
, water acts as a temperature-stabilizing heat reservoir, and land areas near large bodies of water—especially oceans—experience less variation in temperature. Because much of Pangaea's land was distant from its shores, temperatures fluctuated greatly, and the interior probably included expansive
desert
A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
s. Abundant
red beds
Red beds (or redbeds) are sedimentary rocks, typically consisting of sandstone, siltstone, and shale, that are predominantly red in color due to the presence of ferric oxides. Frequently, these red-colored sedimentary strata locally contain ...
and evaporites such as
halite
Halite (), commonly known as rock salt, is a type of salt, the mineral (natural) form of sodium chloride ( Na Cl). Halite forms isometric crystals. The mineral is typically colorless or white, but may also be light blue, dark blue, purple, p ...
support these conclusions, but some evidence suggests the generally dry climate of was punctuated by episodes of increased rainfall.
The most important humid episodes were the
Carnian Pluvial Event
The Carnian pluvial episode (CPE), often called the Carnian pluvial event, was an interval of major change in global climate synchronous with significant changes in Earth's biota both in the sea and on land. It occurred during the latter part of ...
and one in the
Rhaetian
The Rhaetian is the latest age of the Triassic Period (in geochronology) or the uppermost stage of the Triassic System (in chronostratigraphy). It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest age ...
, a few million years before the Triassic–Jurassic extinction event.
Sea levels began to rise during the Jurassic, probably caused by an increase in
seafloor spreading. The formation of new crust beneath the surface displaced ocean waters by as much as above today's sea level, flooding coastal areas. Furthermore, Pangaea began to rift into smaller divisions, creating new shoreline around the Tethys Ocean. Temperatures continued to increase, then began to stabilize.
Humidity
Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present.
Humidity depe ...
also increased with the proximity of water, and deserts retreated.
The climate of the Cretaceous is less certain and more widely disputed. Probably, higher levels of
carbon dioxide
Carbon dioxide (chemical formula ) is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature. In the air, carbon dioxide is transpar ...
in the
atmosphere
An atmosphere () is a layer of gas or layers of gases that envelop a planet, and is held in place by the gravity of the planetary body. A planet retains an atmosphere when the gravity is great and the temperature of the atmosphere is low. A s ...
are thought to have almost eliminated the north–south
temperature gradient
A temperature gradient is a physical quantity that describes in which direction and at what rate the temperature changes the most rapidly around a particular location. The temperature gradient is a dimensional quantity expressed in units of degree ...
: temperatures were about the same across the planet, and about 10°
C higher than today. The circulation of
oxygen
Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements as wel ...
to the deep ocean may also have been disrupted, preventing the
decomposition
Decomposition or rot is the process by which dead organic substances are broken down into simpler organic or inorganic matter such as carbon dioxide, water, simple sugars and mineral salts. The process is a part of the nutrient cycle and is e ...
of large volumes of organic matter, which was eventually
deposited as "
black shale
Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4) and tiny fragments (silt-sized particles) of other minerals, especiall ...
".
Different studies have come to different conclusions about the amount of oxygen in the atmosphere during different parts of the Mesozoic, with some concluding oxygen levels were lower than the current level (about 21%) throughout the Mesozoic, some concluding they were lower in the Triassic and part of the Jurassic but higher in the Cretaceous, and some concluding they were higher throughout most or all of the Triassic, Jurassic and Cretaceous.
Life
Flora
![ProspectPemulwuypine](https://upload.wikimedia.org/wikipedia/commons/9/97/ProspectPemulwuypine.jpg)
The dominant land plant species of the time were
gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνό ...
s, which are vascular, cone-bearing, non-flowering plants such as conifers that produce seeds without a coating. This contrasts with the earth's current flora, in which the dominant land plants in terms of number of species are
angiosperm
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s. The earliest members of the genus ''
Ginkgo'' first appeared during the Middle Jurassic. This genus is represented today by a single species, ''
Ginkgo biloba
''Ginkgo biloba'', commonly known as ginkgo or gingko ( ), also known as the maidenhair tree, is a species of tree native to China. It is the last living species in the order Ginkgoales, which first appeared over 290 million years ago. Fossils ...
''. The
extant
Extant is the opposite of the word extinct. It may refer to:
* Extant hereditary titles
* Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English
* Extant taxon, a taxon which is not extinct, ...
genus ''
Sequoia'' is believed to have evolved in the Mesozoic.
Bennettitales, an extinct group of gymnosperms with foliage superficially resembling that of cycads gained a global distribution during the Late Triassic, and represented one of the most common groups of Mesozoic seed plants.
Flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s radiated during the early Cretaceous, first in the
tropics
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referred to ...
, but the even temperature gradient allowed them to spread toward the poles throughout the period. By the end of the Cretaceous, angiosperms dominated tree floras in many areas, although some evidence suggests that
biomass
Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms bi ...
was still dominated by
cycad
Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...
s and
fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s until after the Cretaceous–Paleogene extinction. Some plant species had distributions that were markedly different from succeeding periods; for example, the
Schizeales
Schizaeales is an order of ferns (class Polypodiopsida).
Description
While the three clades of Schizaeales are all well-distinguished from one another by numerous morphological characters, members of the order all have dimorphic fertile and s ...
, a fern order, were skewed to the Northern Hemisphere in the Mesozoic, but are now better represented in the Southern Hemisphere.
[C.Michael Hogan. 2010]
''Fern''. Encyclopedia of Earth. National council for Science and the Environment
. Washington, DC
Fauna
![Europasaurus holgeri Scene 2](https://upload.wikimedia.org/wikipedia/commons/1/1c/Europasaurus_holgeri_Scene_2.jpg)
The extinction of nearly all animal species at the end of the
Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
Period allowed for the
radiation
In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or through a material medium. This includes:
* ''electromagnetic radiation'', such as radio waves, microwaves, infrared, visi ...
of many new lifeforms. In particular, the extinction of the large
herbivorous pareiasaur
Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, ...
s and
carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
gorgonopsians left those
ecological niche
In ecology, a niche is the match of a species to a specific environmental condition.
Three variants of ecological niche are described by
It describes how an organism or population responds to the distribution of resources and competitors (for ...
s empty. Some were filled by the surviving
cynodont
The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s and
dicynodont
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typicall ...
s, the latter of which subsequently became extinct.
Recent research indicates that it took much longer for the reestablishment of complex ecosystems with high biodiversity, complex food webs, and specialized animals in a variety of niches, beginning in the mid-Triassic 4 million to 6 million years after the extinction,
and not fully proliferated until 30 million years after the extinction.
Animal life was then dominated by various archosaurs:
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s, pterosaurs, and aquatic reptiles such as ichthyosaurs, plesiosaurs, and
mosasaur
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on th ...
s.
The climatic changes of the late Jurassic and Cretaceous favored further adaptive radiation. The Jurassic was the height of archosaur diversity, and the first
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s and
eutherian mammals also appeared. Some have argued that
insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s diversified in
symbiosis
Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
with angiosperms, because insect
anatomy
Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
, especially the mouth parts, seems particularly well-suited for flowering plants. However, all major insect mouth parts preceded angiosperms, and insect diversification actually slowed when they arrived, so their anatomy originally must have been suited for some other purpose.
See also
*
References
*''British Mesozoic Fossils'', 1983, The Natural History Museum, London.
External links
Mesozoic (chronostratigraphy scale)
{{Authority control
Geological eras