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Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alpine tundra gets lower until it reaches sea level, and alpine tundra merges with polar tundra. The high elevation causes an adverse climate, which is too cold and windy to support tree growth. Alpine tundra transitions to sub-alpine forests below the tree line; stunted forests occurring at the forest-tundra ecotone are known as '' Krummholz''. With increasing elevation it ends at the
snow line The climatic snow line is the boundary between a snow-covered and snow-free surface. The actual snow line may adjust seasonally, and be either significantly higher in elevation, or lower. The permanent snow line is the level above which snow wil ...
where snow and ice persist through summer. Alpine tundra occurs in mountains worldwide. The flora of the alpine tundra is characterized by dwarf shrubs close to the ground. The cold climate of the alpine tundra is caused by adiabatic cooling of air, and is similar to
polar climate The polar climate regions are characterized by a lack of warm summers but with varying winters. Every month in a polar climate has an average temperature of less than . Regions with polar climate cover more than 20% of the Earth's area. Most of ...
.


Geography

Alpine tundra occurs at high enough altitude at any latitude. Portions of montane grasslands and shrublands ecoregions worldwide include alpine tundra. Large regions of alpine tundra occur in the
North American Cordillera The North American Cordillera, sometimes also called the Western Cordillera of North America, the Western Cordillera or the Pacific Cordillera, is the North American portion of the American Cordillera, the mountain chain system (cordillera) alon ...
and parts of the northern Appalachian Mountains in North America, the Alps and Pyrenees of Europe, the Himalaya and
Karakoram The Karakoram is a mountain range in Kashmir region spanning the borders of Pakistan, China, and India, with the northwest extremity of the range extending to Afghanistan and Tajikistan. Most of the Karakoram mountain range falls under the ...
of Asia, the Andes of South America, the Eastern Rift mountains of Africa, and the
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman ...
of New Zealand. Alpine tundra occupies high-mountain summits, slopes, and ridges above timberline. Aspect plays a role as well; the treeline often occurs at higher elevations on warmer equator-facing slopes. Because the alpine zone is present only on mountains, much of the landscape is rugged and broken, with rocky, snowcapped peaks, cliffs, and talus slopes, but also contains areas of gently rolling to almost flat topography. Averaging over many locations and local microclimates, the treeline rises when moving 1 degree south from 70 to 50°N, and per degree from 50 to 30°N. Between 30°N and 20°S, the treeline is roughly constant, between .


Climate

Alpine climate is the average weather ( climate) for the alpine tundra. The climate becomes colder at high elevations—this characteristic is described by the
lapse rate The lapse rate is the rate at which an atmospheric variable, normally temperature in Earth's atmosphere, falls with altitude. ''Lapse rate'' arises from the word ''lapse'', in the sense of a gradual fall. In dry air, the adiabatic lapse rate is ...
of air: air tends to get colder as it rises, since it expands. The dry adiabatic lapse rate is 10 °C per km (5.5 °F per 1000 ft) of elevation or altitude. Therefore, moving up on a mountain is roughly equivalent to moving 80 kilometers (45 miles or 0.75° of latitude) towards the pole. This relationship is only approximate, however, since local factors such as proximity to oceans can drastically modify the climate. In the alpine tundra, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snowpack, or associated lack of available moisture). Typical high-elevation growing seasons range from 45 to 90 days, with average summer temperatures near . Growing season temperatures frequently fall below freezing, and frost occurs throughout the growing season in many areas. Precipitation occurs mainly as winter snow, but soil water availability is highly variable with season, location, and topography. For example, snowfields commonly accumulate on the lee sides of ridges while ridgelines may remain nearly snow free due to redistribution by wind. Some alpine habitats may be up to 70% snow free in winter. High winds are common in alpine ecosystems, and can cause significant soil erosion and be physically and physiologically detrimental to plants. Also, wind coupled with high solar radiation can promote extremely high rates of
evaporation Evaporation is a type of vaporization that occurs on the surface of a liquid as it changes into the gas phase. High concentration of the evaporating substance in the surrounding gas significantly slows down evaporation, such as when humidi ...
and transpiration.


Quantifying the climate

There have been several attempts at quantifying what constitutes an alpine climate. Climatologist Wladimir Köppen demonstrated a relationship between the Arctic and Antarctic tree lines and the 10 °C summer isotherm; i.e., places where the average temperature in the warmest calendar month of the year is below 10 °C cannot support forests. See Köppen climate classification for more information. Otto Nordenskjöld theorized that winter conditions also play a role: His formula is ''W'' = 9 − 0.1 ''C'', where ''W'' is the average temperature in the warmest month and ''C'' the average of the coldest month, both in degrees Celsius (this would mean, for example, that if a particular location had an average temperature of in its coldest month, the warmest month would need to average or higher for trees to be able to survive there). In 1947, Holdridge improved on these schemes, by defining
biotemperature The Holdridge life zones system is a global bioclimatic scheme for the classification of land areas. It was first published by Leslie Holdridge in 1947, and updated in 1967. It is a relatively simple system based on few empirical data, giving obj ...
: the mean annual temperature, where all temperatures below 0 °C are treated as 0 °C (because it makes no difference to plant life, being dormant). If the mean biotemperature is between , Holdridge quantifies the climate as alpine.


Flora

Since the habitat of alpine vegetation is subject to intense radiation, wind, cold, snow, and ice, it grows close to the ground and consists mainly of perennial grasses, sedges, and forbs. Perennial herbs (including grasses, sedges, and low woody or semi-woody shrubs) dominate the alpine landscape; they have much more root and
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
biomass than that of shoots, leaves, and flowers. The roots and rhizomes not only function in water and nutrient absorption but also play a very important role in over-winter carbohydrate storage. Annual plants are rare in this ecosystem and usually are only a few inches tall, with weak root systems. Other common plant life-forms include prostrate shrubs;
tussock Tussock may refer to: * Tussock grass, a group of species in the family Poaceae *Floating island * Lymantriinae, called tussock moths or tussocks See also * Hassock (disambiguation) Hassock may refer to: * Kneeler, a cushion or a piece of f ...
-forming
graminoid In botany and ecology, graminoid refers to a herbaceous plant with a grass-like morphology, i.e. elongated culms with long, blade-like leaves. They are contrasted to forbs, herbaceous plants without grass-like features. The plants most ofte ...
s; cushion plants; and cryptogams, such as bryophytes and
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. Relative to lower elevation areas in the same region, alpine regions have a high rate of endemism and a high diversity of plant species. This
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
diversity can be attributed to geographical isolation, climate changes, glaciation, microhabitat differentiation, and different histories of migration or evolution or both. These phenomena contribute to plant diversity by introducing new flora and favoring adaptations, both of new species and the dispersal of pre-existing species. Plants have adapted to the harsh alpine environment. Cushion plants, looking like ground-hugging clumps of moss, escape the strong winds blowing a few inches above them. Many flowering plants of the alpine tundra have dense hairs on stems and leaves to provide wind protection or red-colored pigments capable of converting the sun's light rays into heat. Some plants take two or more years to form flower buds, which survive the winter below the surface and then open and produce fruit with seeds in the few weeks of summer. Alpine areas are unique because of the severity and complexity of their environmental conditions. Very small changes in topography – as small as 1 foot (0.3 m) or less – may mean the difference between a windswept area or an area of snow accumulation, changing the potential
productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proces ...
and plant community drastically. Between these extremes of drought versus saturation, several intermediate environments may exist all within a few yards of each other, depending on topography, substrate, and climate. Alpine vegetation generally occurs in a mosaic of small patches with widely differing environmental conditions. Vegetation types vary from cushion and rosette plants on the ridges and in the rock crannies; to herbaceous and grassy vegetation along the slopes; dwarf shrubs with grasses and forbs below the melting snowdrifts; and sedges, grasses, low shrubs, and mosses in the bogs and along the brooks. Alpine meadows form where sediments from the weathering of rocks has produced soils well-developed enough to support grasses and sedges. Non-flowering
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.algal cells can photosynthesize at any temperature above , and the outer fungal layers can absorb more than their own weight in water. The adaptations for survival of drying winds and cold may make tundra vegetation seem very hardy, but in some respects the tundra is very fragile. Repeated footsteps often destroy tundra plants, allowing exposed soil to blow away; recovery may take hundreds of years.


Fauna

Because alpine tundra is located in various widely separated regions of the Earth, there is no animal species common to all areas of alpine tundra. Some animals of alpine tundra environments include the kea, marmot, mountain goat, bighorn sheep,
chinchilla Chinchillas are either of two species (''Chinchilla chinchilla'' and ''Chinchilla lanigera'') of crepuscular rodents of the parvorder Caviomorpha. They are slightly larger and more robust than ground squirrels, and are native to the Andes mount ...
, Himalayan tahr,
yak The domestic yak (''Bos grunniens''), also known as the Tartary ox, grunting ox or hairy cattle, is a species of long-haired domesticated cattle found throughout the Himalayan region of the Indian subcontinent, the Tibetan Plateau, Kachin Sta ...
, snow leopard, and pika.


See also

* Montane grasslands and shrublands *
Urals montane tundra and taiga The Urals montane tundra and taiga ecoregion (WWF ID: PA0610) covers the main ridge of the Ural Mountains (both sides) - a 2,000 km (north-south) by 300 km (west-east) region. The region is on the divide between European and Asian ecoreg ...
* Polonyna (montane meadow) * Montane ecosystems * Tundra


References


External links

*{{Commons-inline Climate by mountain range Tundra Montane grasslands and shrublands Mountain meteorology Natural regions Tundra