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Male (
symbol A symbol is a mark, sign, or word that indicates, signifies, or is understood as representing an idea, object, or relationship. Symbols allow people to go beyond what is known or seen by creating linkages between otherwise very different conc ...
:
♂ A gender symbol is a pictogram or glyph used to represent sex and gender, for example in biology and medicine, in genealogy, or in the sociological fields of gender politics, LGBT subculture and identity politics. In his books (1767) and ...
) is the
sex Sex is the trait that determines whether a sexually reproducing animal or plant produces male or female gametes. Male plants and animals produce smaller mobile gametes (spermatozoa, sperm, pollen), while females produce larger ones ( ova, of ...
of an
organism In biology, an organism () is any life, living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy (biology), taxonomy into groups such as Multicellular o ...
that produces the
gamete A gamete (; , ultimately ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. In species that produce ...
(sex cell) known as sperm, which fuses with the larger
female Female (symbol: ♀) is the sex of an organism that produces the large non-motile ova (egg cells), the type of gamete (sex cell) that fuses with the male gamete during sexual reproduction. A female has larger gametes than a male. Females a ...
gamete, or
ovum The egg cell, or ovum (plural ova), is the female reproductive cell, or gamete, in most anisogamous organisms (organisms that reproduce sexually with a larger, female gamete and a smaller, male one). The term is used when the female gamete i ...
, in the process of
fertilization Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Pro ...
. A male organism cannot reproduce sexually without access to at least one ovum from a female, but some organisms can reproduce both sexually and asexually. Most male mammals, including male humans, have a
Y chromosome The Y chromosome is one of two sex chromosomes ( allosomes) in therian mammals, including humans, and many other animals. The other is the X chromosome. Y is normally the sex-determining chromosome in many species, since it is the presence or a ...
, which codes for the production of larger amounts of
testosterone Testosterone is the primary sex hormone and anabolic steroid in males. In humans, testosterone plays a key role in the development of male reproductive tissues such as testes and prostate, as well as promoting secondary sexual characteris ...
to develop
male reproductive organs The male reproductive system consists of a number of sex organs that play a role in the process of human reproduction. These organs are located on the outside of the body and within the pelvis. The main male sex organs are the penis and the test ...
. Not all species share a common
sex-determination system A sex-determination system is a biological system that determines the development of sexual characteristics in an organism. Most organisms that create their offspring using sexual reproduction have two sexes. In some species there are hermap ...
. In most
animal Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motilit ...
s, including
humans Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
, sex is determined genetically; however, species such as '' Cymothoa exigua'' change sex depending on the number of females present in the vicinity. In humans, the word ''male'' can also be used to refer to
gender Gender is the range of characteristics pertaining to femininity and masculinity and differentiating between them. Depending on the context, this may include sex-based social structures (i.e. gender roles) and gender identity. Most cultures us ...
in the social sense of gender role or gender identity.


Overview

The existence of separate sexes has evolved independently at different times and in different lineages, an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
. The repeated pattern is sexual reproduction in isogamous species with two or more
mating type Mating types are the microorganism equivalent to sexes in multicellular lifeforms and are thought to be the ancestor to distinct sexes. They also occur in macro-organisms such as fungi. Definition Mating types are the microorganism equivalent to s ...
s with gametes of identical form and behavior (but different at the molecular level) to
anisogamous Different forms of anisogamy: A) anisogamy of motile cells, B) 283x283px Anisogamy is a form of sexual reproduction">egg cell">oogamy (egg cell and sperm cell), C) anisogamy of non-motile cells (egg cell and spermatia).">283x283px Anisogamy is ...
species with
gamete A gamete (; , ultimately ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. In species that produce ...
s of male and
female Female (symbol: ♀) is the sex of an organism that produces the large non-motile ova (egg cells), the type of gamete (sex cell) that fuses with the male gamete during sexual reproduction. A female has larger gametes than a male. Females a ...
types to
oogamous Oogamy is an extreme form of anisogamy where the gametes differ in both size and form. In oogamy the large female gamete (also known as ovum) is immobile, while the small male gamete (also known as sperm) is mobile. Oogamy is a common form of ...
species in which the female gamete is very much larger than the male and has no ability to move. There is a good argument that this pattern was driven by the physical constraints on the mechanisms by which two gametes get together as required for
sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote th ...
. Accordingly, sex is defined across species by the type of gametes produced (i.e.: spermatozoa vs. ova) and differences between males and females in one lineage are not always predictive of differences in another. Male/female dimorphism between organisms or reproductive organs of different sexes is not limited to animals; male gametes are produced by
chytrid Chytridiomycota are a division of zoosporic organisms in the kingdom Fungi, informally known as chytrids. The name is derived from the Ancient Greek ('), meaning "little pot", describing the structure containing unreleased zoöspores. Chytri ...
s, diatoms and land
plant Plants are predominantly Photosynthesis, photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all curr ...
s, among others. In land plants, ''female'' and ''male'' designate not only the female and male gamete-producing organisms and structures but also the structures of the sporophytes that give rise to male and female plants.


Evolution

The evolution of anisogamy led to the evolution of male and female function. Before the evolution of anisogamy,
mating types In biology, mating is the pairing of either opposite-sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. '' Copulation'' is the union of the sex organs of two sexually reprodu ...
in a species were isogamous: the same size and both could move, catalogued only as "+" or "-" types. In anisogamy, the mating type is called a gamete. The male gamete is smaller than the female gamete, and usually mobile. Anisogamy remains poorly understood, as there is no fossil record of its emergence. Numerous theories exist as to why anisogamy emerged. Many share a common thread, in that larger female gametes are more likely to survive, and that smaller male gametes are more likely to find other gametes because they can travel faster. Current models often fail to account for why isogamy remains in a few species. Anisogamy appears to have evolved multiple times from isogamy; for example female
Volvocales Chlamydomonadales, also known as Volvocales, are an order of flagellated or pseudociliated green algae, specifically of the Chlorophyceae.See the NCBIbr>webpage on Chlamydomonadales Data extracted from the Chlamydomonadales can form planar or ...
(a type of green algae) evolved from the plus
mating type Mating types are the microorganism equivalent to sexes in multicellular lifeforms and are thought to be the ancestor to distinct sexes. They also occur in macro-organisms such as fungi. Definition Mating types are the microorganism equivalent to s ...
. Although sexual evolution emerged at least 1.2 billion years ago, the lack of anisogamous fossil records make it hard to pinpoint when males evolved. One theory suggests male evolved from the dominant
mating type Mating types are the microorganism equivalent to sexes in multicellular lifeforms and are thought to be the ancestor to distinct sexes. They also occur in macro-organisms such as fungi. Definition Mating types are the microorganism equivalent to s ...
(called mating type minus).


Symbol and usage


Symbol

A common
symbol A symbol is a mark, sign, or word that indicates, signifies, or is understood as representing an idea, object, or relationship. Symbols allow people to go beyond what is known or seen by creating linkages between otherwise very different conc ...
used to represent the male sex is the Mars symbol ♂, a circle with an arrow pointing
northeast The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...
. The
Unicode Unicode, formally The Unicode Standard,The formal version reference is is an information technology standard for the consistent encoding, representation, and handling of text expressed in most of the world's writing systems. The standard, ...
code-point is: : The symbol is identical to the planetary symbol of
Mars Mars is the fourth planet from the Sun and the second-smallest planet in the Solar System, only being larger than Mercury. In the English language, Mars is named for the Roman god of war. Mars is a terrestrial planet with a thin atmos ...
. It was first used to denote sex by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in 1751. The symbol is sometimes seen as a stylized representation of the shield and spear of the
Roman god Roman mythology is the body of myths of ancient Rome as represented in the literature and visual arts of the Romans. One of a wide variety of genres of Roman folklore, ''Roman mythology'' may also refer to the modern study of these representa ...
Mars Mars is the fourth planet from the Sun and the second-smallest planet in the Solar System, only being larger than Mercury. In the English language, Mars is named for the Roman god of war. Mars is a terrestrial planet with a thin atmos ...
. According to Stearn, however, this derivation is "fanciful" and all the histori