Malabar Banded Swallowtail
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''Papilio liomedon'', the Malabar banded swallowtail, is a member of the
swallowtail butterfly Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the large ...
family found in southern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
. Earlier considered a subspecies of the banded swallowtail (''
Papilio demolion ''Papilio demolion'', the banded swallowtail, is a species of swallowtail butterfly belonging to the family Papilionidae. Subspecies Subspecies include: *''Papilio demolion demolion'' Cramer, 1776 *''Papilio demolion energetes'' Fruhstorfer, 190 ...
'') of
southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, it is now considered a distinct species.


Description

It is similar to ''
Papilio demolion ''Papilio demolion'', the banded swallowtail, is a species of swallowtail butterfly belonging to the family Papilionidae. Subspecies Subspecies include: *''Papilio demolion demolion'' Cramer, 1776 *''Papilio demolion energetes'' Fruhstorfer, 190 ...
'' but distinguishable chiefly by the pale greenish-yellow band that crosses the wings starting from the middle and not from just before the middle of the dorsal margin of the hindwing, also this band is composed entirely of separate spots on the forewing.


Range

Western Ghats and hills of southern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
. It is common (May to August) in
Thenmala Thenmala is a tourist destination in the eastern side of Kollam district in Kerala, India that is home to the first eco-tourism centre in India. The word 'Thenmala' means "'Honey hills" in Malayalam language. The town is 66 km away from Ko ...
,
Kollam district Kollam district (), (formerly Quilon district) is one of 14 districts of the state of Kerala, India. The district has a cross-section of Kerala's natural attributes; it is endowed with a long coastline, a major Laccadive Sea seaport and an in ...
, south Kerala.


Status

The
IUCN Red Data Book The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
records the Malabar banded swallowtail as uncommon and not threatened as a species. However a survey in the early 1990s by
Harish Gaonkar Harish (Honnayya) S. Gaonkar (born 1946) is an Indian specialist on butterflies who contributed to the Zoological Museum at the University of Copenhagen, Denmark and wrote a 1996 compilation of butterflies of Western Ghats, South India catalog ...
showed the butterfly to be rare but distributed from Kerala to Goa. The butterfly was considered to be common in Karwar in the past. It is not to be found in Maharashtra and Gujarat. It is protected by law in India.


Habitat

Generally found in the semi-evergreen and evergreen tropical forests of the Western Ghats where it flies mainly during the
monsoon A monsoon () is traditionally a seasonal reversing wind accompanied by corresponding changes in precipitation but is now used to describe seasonal changes in atmospheric circulation and precipitation associated with annual latitudinal oscil ...
months.


Habits

The males are fond of heavy jungle especially on hilltops. The females fly lower in even thicker jungle. The butterflies visit flowers such as ''
Lantana ''Lantana'' () is a genus of about 150 species of perennial flowering plants in the verbena family, Verbenaceae. They are native to tropical regions of the Americas and Africa but exist as an introduced species in numerous areas, especially in ...
'', ''
Stachytarpheta ''Stachytarpheta'' is a plant genus in the verbena family (Verbenaceae). The flowers are rich in nectar and popular with many butterflies, such as the South Asian crimson rose (''Atrophaneura hector''), Malabar banded swallowtail (''Papilio lio ...
'' and ''
Clerodendrum ''Clerodendrum'' is a genus of flowering plants formerly placed in the family Verbenaceae, but now considered to belong to the Lamiaceae (mint) family. Its common names include glorybower, bagflower and bleeding-heart. It is currently classified ...
''.Susanth, C. (2005
Biology of Malabar Banded Swallowtail ''Papilio liomedon'' Moore.
''Journal of the Bombay Natural History Society'' 102(1): 33-37


Life cycle

There are two to three broods a year. Recorded in
Kodagu Kodagu (also known by its former name Coorg) is an administrative district in the Karnataka state of India. Before 1956, it was an administratively separate Coorg State, at which point it was merged into an enlarged Mysore State. It occupies ...
(Coorg) as having broods from September to October, November to December, and, from April to May. Recorded in Karnataka in July and in September. Males appear to outnumber the females. The life-history of this insect is given by Messrs. Davidson and Aitken: One of these gentlemen watched a female, ''P. liomedon'', laying its eggs on a tender shoot of a small jungle tree or shrub (''
Acronychia ''Acronychia'' is a genus of about fifty species of plants in the rue family Rutaceae. The leaves are simple or pinnate, and the flowers bisexual with four sepals, four petals and eight stamens. They have a broad distribution including in India, ...
laurifolia''). There were "ten eggs, laid one on top of the other." Of the caterpillars which emerged five days after the eggs were laid, "five passed successfully through all dangers and became beautiful specimens, one female and four males. (This is one of the butterflies of which we rarely find females.) All through their lives these larvae continued gregarious, dispersing occasionally to feed, but always returning to rest side by side on the upper surface of a leaf. The following dates may be interesting. Eggs laid 2 August, hatched 7 August; skins cast (and eaten) 12 August; again 17 August; again 20 to 22 August. The most advanced cast its skin again on the 28th August, became a pupa on the 2nd of September, and emerged on the 15th of September. The others followed within two days. At first the larvae were of an oily yellow colour and bore many pairs of spiny points, but these disappeared with age and after the last moult there were only the short fleshy processes on the 2nd and last segment which characterise the group, and one additional curved pair on the ninth segment." "The colour after the last moult was a clear slaty-blue, changing eventually to a greenish tint, with light brown markings very much the same as those which characterise the rest of the group. The pupa was more abruptly bent back from the middle of the thorax than that of ''P. erithonius'' (i. e. ''P. demoleus'') and adorned on the thorax with a sword-shaped horn, fully three-eighths of an inch long, and always bent either to the right or the left. The colour was brown or green and yellow according to situation."


Eggs

The female lays her eggs one over the other in a stick of ten to sixteen. The orange eggs gradually fade to yellow. The pale black of the larval heads appear on the upperside of the eggs on the fourth day. The eggs hatch on the fifth day.


Larva

The larvae live gregariously and are heavily parasitized (up to 90%). The first
instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or ass ...
lasts about five days, second instar for four days, third instar five, fourth instar four, about three days for the fifth instar and another three days for the sixth instar. They begin to pupate after around 22 days of their larval duration.


Pupa

The lower portion of the pupa is yellow or fluorescent green with purple lines. Between the head and the thorax region a projection like a jug-handle is seen. This is distinctive of the species. Below the thoracic region, two yellow spots are seen. The pupal duration is about 15 days.


Food plants

'' Acronychia laurifolia'' and '' Evodia roxburghiana'' of the family
Rutaceae The Rutaceae is a family, commonly known as the rueRUTACEAE
in BoDD – Botanical Derm ...
.Gaden S. Robinson, Phillip R. Ackery, Ian J. Kitching, George W. Beccaloni AND Luis M. Hernández. HOSTS - a Database of the World's Lepidopteran Hostplant

Accessed November 2006


Cited references


Other references

* * * * *


See also

*
Papilionidae Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the larges ...
*
List of butterflies of India The following is a list of the butterflies of India. India has extremely diverse terrain, climate and vegetation, which comprises extremes of heat cold, desert and jungle, of low-lying plains and the highest mountains, of dryness and dampness, i ...
*
List of butterflies of India (Papilionidae) This is a list of the butterflies of family Papilionidae (superfamily Papilionoidea), or the swallowtails, which are found in India. This family of large and beautiful butterflies is well represented with 89 species found within Indian borde ...
{{Taxonbar, from=Q1317880 liomedon Butterflies of Asia Butterflies described in 1874