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''Magericyon'' is an extinct genus of
Amphicyonid Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene (around 45 mya), spread to Europe by the late Eocene (35 mya), and appear in Asia, ...
("bear-dog") that lived during the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
10-9 Ma (
Vallesian The Vallesian age is a period of geologic time (11.6–9.0 Ma) within the Miocene used more specifically with European Land Mammal Ages. It precedes the Turolian age and follows the Astaracian age. The so-called Vallesian Crisis resulted in the e ...
Age) in what is now
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
.


Description

The appearance of this animal was vaguely similar to that of a particularly robust, large felid, but the skull resembles that of a canid or an ursid, like that of many amphicyonids. Unlike most other amphicyonids, ''Magericyon'' had teeth associated with those of a hypercarnivore, with laterally flattened canines, the third premolar having a single root, the absence of second premolars and a metaconid on its lower molars, with a reduction in the second upper molar. The scapula and the front leg showed primitive features such as an acromion in the shoulder with a reduced caudoventral projection and post scapular pit.Peigné, S., Salesa, M. J., Antón, M. & Morales, J., 2008: A new amphicyonine (Carnivora: Amphicyonidae) from the upper Miocene of Batallones-1, Madrid, Spain. Palaeontology: Vol. 51, #4, pp. 943Siliceo, G., Salesa, M. J., Antón, M., Pastor, J. F., Morales, J. 2015. Comparative Anatomy of the Shoulder Region in the Late Miocene Amphicyonid Magericyon anceps (Carnivora): Functional and Paleoecological Inferences. Journal of Mammalian Evolution, 22: 243–258. ''Magericyon'' was roughly equivalent to a large leopard in size, weighing around .


Classification

''Magericyon'' was described for the first time in 2008, based on fossils found in
Cerro de los Batallones Cerro de los Batallones (''Hill of the Battalions'') is a hill at Torrejón de Velasco, Madrid, Spain where a number of fossil sites from the Upper Miocene ( MN10) have been found. Nine sites have been discovered with predominantly vertebrate fo ...
in Spain. The type species is ''Magericyon anceps'', but a second species has also been attributed to the genus as ''M. castellanus'', described in 1981 and initially attributed to the genus ''Amphicyon''. ''Magericyon'' is part of the family of amphicyonidae, a group of very common carnivores ranging from the Eocene to the Miocene, and which occupied many different ecological niches. Magericyon is the last amphicyonid known from Western Europe, but its features are mixed; on one hand the teeth were very specialized, while on the other, front limb and morphology of the scapula was more primitive. Evidence also indicates that Magericyon was closely related to ''
Amphicyon ''Amphicyon'' ("ambiguous dog") is an extinct genus of large carnivorous bone-crushing mammals, popularly known as bear dogs, of the family Amphicyonidae, subfamily Amphicyoninae, from the Burdigalian Epoch until the late Pliocene, with the creat ...
''.


Paleobiology

''Magericyon'' occupied a different ecological niche than other amphicyonids, such as the larger ''Amphicyon'' and ''
Ysengrinia ''Ysengrinia'' is an extinct genus of carnivoran in the family Amphicyonidae (beardogs), which lived in Europe, Asia, and North America during the Early Miocene. It was also reported from Egypt and Namibia, but this material has been reassigned ...
'' (which had lifestyles more akin to bears) or ''
Daphoenodon ''Daphoenodon'' is an extinct genus of terrestrial carnivore, which lived in the early Miocene and belonged to the family Amphicyonidae ("bear dogs") of the suborder Caniformia Caniformia is a suborder within the order Carnivora consistin ...
'' and ''
Temnocyon ''Temnocyon'' is an extinct genus of bear-dogs endemic to North America. It lived from the Oligocene to Early Miocene approximately 30.8—20.4 mya, existing for about . The first fossils are recorded in North America at Logan Butte in the Joh ...
'' of North America, which were more capable runners. ''Magericyon'' probably lived in a similar manner to that of modern felines, being an ambush hunter of large prey. Studies by Gemo Siliceo ''et al'' also revealed that ''Magericyon'' had powerful jaw and neck muscles that helped to stabilize its head and jaws during a bite. The amphicyonid was particularly adept at side-to-side movements and rotations of the head. This feature allowed ''Magericyon'' to swiftly and efficiently process the meat on a carcass, allowing the bear-dog to devour sufficient amounts of flesh before scavengers arrived to steal the predator's hard-earned meal.


Paleoecology

As a carnivore at Cerro de los Battalones, ''Magericyon'' shared the apex predator position with two saber-toothed cat species, the leopard-sized '' Promegantereon ogygia'' and the tiger-sized '' Machairodus aphanistus.'' Evidence indicates that the large carnivores may have co-existed using ''
niche partitioning In ecology, niche differentiation (also known as niche segregation, niche separation and niche partitioning) refers to the process by which competing species use the environment differently in a way that helps them to coexist. The competitive excl ...
.'' A carbon-13 bone analysis matched isotope profiles in prey species with their predators, showing both cats hunted prey that typically live in closed woodland habitats, such as pigs (
Microstonyx ''Microstonyx'' was an extinct genus of suid that existed during the Miocene in Asia and Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its h ...
) and perhaps occasionally young of the gomphotheriid mastodon ''
Tetralophodon ''Tetralophodon'' ("four-ridged tooth") is an extinct proboscidean genus belonging to the superfamily Elephantoidea. Taxonomy The genus ''Tetralophodon'' (meaning "four-ridged tooth") was named in the mid-19th century with the discovery of the ...
''. ''Magericyon'' ate medium-sized prey that live in more open habitats, with the antelope ''Austroportax'' being an important food source and hipparionine horses also present in the diet. Because the site attracted all three species as a
predator trap A predator trap is a natural hazard where prey animals become trapped or incapacitated, and the attracted predators suffer the same fate. More predators, scavengers, insects and birds become attracted to this mounting accumulation of carrion, until ...
, ''Magericyon'' must also have taken carrion or injured animals of various kinds, though its teeth show it was specialized as a
hypercarnivore A hypercarnivore is an animal which has a diet that is more than 70% meat, either via active predation or by scavenging. The remaining non-meat diet may consist of non-animal foods such as fungi, fruits or other plant material. Some extant examp ...
without the bone-cracking adaptations of many other bear-dogs. ''Magericyon'' may have competed at times with large omnivorous bears such as ''
Agriotherium ''Agriotherium'' is an extinct genus of bears whose fossils are found in Miocene through Pleistocene-aged strata of North America, Europe, Africa, and Asia. This long-lived genus persisted from at least ~11.6–2.5 Mya. Materials from the late ...
'' and ''
Indarctos ''Indarctos'' is an extinct genus of bear, endemic to North America, Europe and Asia during the Miocene. It was present from ~11.1 to 5.3 Ma, existing for approximately . The oldest member is from Arizona (~11.1—7.7 Ma) and youngest is (~9.0 ...
'', but these genera focused more on vegetable food''.'' As well as ''Austroportax'' and ''
Hippotherium ''Hippotherium'' is an extinct genus of horse that lived in during the Miocene through Pliocene ~13.65—3.3 Mya, existing for . Species The type species, ''H. primigenius'', is known from Miocene deposits in Europe and the Middle East ...
,'' its prey could have included young of the hornless rhinoceros
Aceratherium ''Aceratherium'' (Greek: "without (a) horn" (keratos), "beast" (therion)) was a genus of rhinoceros of the subfamily Aceratheriinae that lived in Eurasia during the Miocene. Taxonomy ''Aceratherium'' was coined by Kaup (1832) for ''"Rhinocer ...
and possibly the calves of the large silvatherid giraffes and boselaphine antelopes. Since its choice of prey suggests life in open country, but its legs were not designed for speed, ''Magericyon'' probably would have wasted no time in stealing a meal from any of the smaller carnivores in the region or in scavenging when the opportunity presented itself.


References

{{Taxonbar, from=Q25400575 Bear dogs Miocene bear dogs Miocene mammals of Europe Fossil taxa described in 2008 Prehistoric carnivoran genera