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''Maesopsis eminii'', the umbrella tree, is a species of tree in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Rhamnaceae found in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
and
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. It is the only species in the genus ''Maesopsis''. It is often grown as a plantation tree, and as a shade tree in
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. Seeds of ...
plantations and other crops. Birds and monkeys may disperse the seeds. Since this tree grows fast it is often used for regeneration of destroyed forest lands. Its timber is used for construction and firewood and its leaves for animal fodder.


Description

''Maesopsis eminii'' is a large, fast-growing semi-deciduous tree growing to a height of about . The trunk is straight with a clear bole for the lowest third. The bark is greyish-brown and deeply furrowed. The branches are mostly horizontal causing the crown to be flat, although it becomes more rounded as the tree ages. The leaves are simple and elliptic-lanceolate, with toothed margins, , glossy above, with a gland in each tooth. On the underside of the leaves there are
domatia A domatium (plural: domatia, from the Latin "domus", meaning home) is a tiny chamber that houses arthropods, produced by a plant. Ideally domatia differ from galls in that they are produced by the plant rather than being induced by their inhabi ...
in the axils of the secondary nerves. The inflorescence is a many-flowered cyme up to long, each greenish-yellow flower having five calyx lobes that are longer than the corolla lobes and a single anther. The fruit is a drupe up to long, green at first, turning yellow and then purple-black as it ripens. It contains a large stone with one or two black seeds inside.


Distribution and habitat

''Maesopsis eminii'' is native to much of West and Central Africa, its range extending from Liberia, the Congo and Angola to Uganda. It has become established in Sudan, Somalia, Ethiopia, Kenya, Tanzania and Zambia. It was introduced into the
Usambara Mountains The Usambara Mountains of northeastern Tanzania in tropical East Africa, comprise the easternmost ranges of the Eastern Arc Mountains. The ranges of approximately long and about half that wide, are situated in the Lushoto District of the Tan ...
in eastern Tanzania between 1930 and 1970, where it became an invasive species in the submontane rainforest, now being the dominant species of tree there. It is a
pioneer species Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire. Pioneer flora Some lichens grow on rocks without soil, so ...
, found in humid forests, colonising cleared areas and flourishing as secondary growth. It is often present in the border zone between
savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
h and forest.


Ecology

The tree starts flowering when only a few years old. The flowers are
hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have s ...
and are probably pollinated by insects with one or two fruits usually developing per
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
. Monkeys and large birds such as hornbills help to disperse the seed by carrying the fruit away to consume elsewhere. Where the trees become established the previous root mat disappears, the leaf litter becomes thinner, the soil pH rises and the
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...
of ground plants and animals decreases.


Uses

''Maesopsis eminii'' is a useful timber tree. The sapwood is pale and the
heartwood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin t ...
is olive-brown to deep red. It is used for poles, boxes and crates but is susceptible to damage by
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blatto ...
s and to rotting when in continual contact with wet ground. The wood is used for fuel and the foliage for fodder. The tree is planted to provide shade for such crops as
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. Seeds of ...
, cocoa and
cardamom Cardamom (), sometimes cardamon or cardamum, is a spice made from the seeds of several plants in the genera ''Elettaria'' and ''Amomum'' in the family Zingiberaceae. Both genera are native to the Indian subcontinent and Indonesia. They are r ...
, and is often grown domestically in gardens for the same purpose.


References

{{Taxonbar, from1=Q9026752, from2=Q1883624, from3=Q16333755 Rhamnaceae Rhamnaceae genera Monotypic Rosales genera