Macaca Assamensis
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The Assam macaque (''Macaca assamensis'') or Assamese macaque is a
macaque The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and (in one instance) Gibraltar. Macaques are principally ...
of the Old World monkey family native to
South South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
and Southeast Asia. Since 2008, it has been listed as Near Threatened on the IUCN Red List, as it is experiencing significant declines due to poaching, habitat degradation, and
fragmentation Fragmentation or fragmented may refer to: Computers * Fragmentation (computing), a phenomenon of computer storage * File system fragmentation, the tendency of a file system to lay out the contents of files non-continuously * Fragmented distributi ...
.


Characteristics

The Assam macaque has a yellowish-grey to dark brown pelage. The facial skin is dark brownish to purplish. The head has a dark fringe of hair on the cheeks directed backwards to the ears. The hair on the crown is parted in the middle. The shoulders, head and arms tend to be paler than the hindquarters, which are greyish. The tail is well-haired and short. Head-to-body-length measures , and the tail is long. Adult weight is .


Distribution and habitat

The ''Macaca assamensis'' "Nepal population" is endemic to Nepal and likely in some way distinct from the two recognized subspecies, which occupy adjacent areas to the southeast and east of the range of ''M. assamensis''. There is a gap in northeastern India between the two main population pockets, specifically between central Bhutan and the south side of the
Brahmaputra River The Brahmaputra is a trans-boundary river which flows through Tibet, northeast India, and Bangladesh. It is also known as the Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, Luit in Assamese, and Jamuna River in Bangla. It ...
; the east bank of its upper course marks the division between the two recognized subspecies:Groves, C. P. (2001) ''Primate taxonomy.'' Smithsonian Institution Press, Washington, DC, USA. * The eastern Assamese macaque, ''M. a. assamensis'', occurs in Bhutan, Arunachal Pradesh, Assam, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, and Tripura in northeastern India, into northern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, southeast through the Myanmar- Thailand border ranges as far as Chongkrong, to the upper Mekong in Tibet, into the provinces of
Guangxi Guangxi (; ; Chinese postal romanization, alternately romanized as Kwanghsi; ; za, Gvangjsih, italics=yes), officially the Guangxi Zhuang Autonomous Region (GZAR), is an Autonomous regions of China, autonomous region of the People's Republic ...
, Guizhou, Tibet and Yunnan in southwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, in Thateng in northern
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
, and Hoi Xuan in northern Vietnam; * The western Assamese macaque, ''M. a. pelops'', is found from central Nepal through Uttar Pradesh, Sikkim, Assam and northernmost West Bengal in northern India, into central Bhutan and the Sundarbans in Bangladesh. During surveys carried out in 1976, 1978, and 1984 in Nepal, Assam macaques were found to be patchily distributed along rivers in tropical and subtropical forests at altitudes from . They are apparently absent from areas west of the Kaligandaki River. In India, they live in tropical and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
semi
evergreen forests An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zon ...
, dry deciduous and montane forests, from the sea level to altitudes of . They usually inhabit hill areas above , but in the wetter east they may occur even in the lowlands, and frequent areas that only marginally reach this altitude. In Laos and Vietnam, they prefer high altitudes, usually above . In forests on limestone
karst Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...
, they occur in much lower elevations.


Ecology and behaviour

Assam macaques are diurnal, and at times both arboreal and terrestrial. They are omnivorous and feed on fruits, leaves, invertebrates and
cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...
s. In
Namdapha National Park Namdapha National Park is a large protected area in Arunachal Pradesh of Northeast India. The park was established in 1983. With more than 1,000 floral and about 1,400 faunal species, it is a biodiversity hotspot in the Eastern Himalayas. The ...
, Arunachal Pradesh, 15 groups were recorded in 2002 comprising 209 individuals. The population had a group density of 1.11 individuals per , and an average group size of 13.93 individuals.Chetry, D., Medhi, R., Biswas, J., Das, D. and Bhattacharjee, P. C. (2003)
''Nonhuman primates in the Namdapha National Park, Arunachal Pradesh, India''
International Journal of Primatology 24(2): 383–388.
During a survey in Nepal's Langtang National Park in 2007, a total of 213 Assamese macaques were encountered in 9 groups in the study area of . Troop sizes varied between 13 and 35 individuals, with a mean troop size of 23.66 individuals, and comprised 31% adult females, 16% adult males, and their young of various ages. They preferred maize kernals, followed by potato tubers, but also raided fields with wheat, buckwheat, and
millet Millets () are a highly varied group of small-seeded grasses, widely grown around the world as cereal crops or grains for fodder and human food. Most species generally referred to as millets belong to the tribe Paniceae, but some millets al ...
.


Threats

The threats to this species' habitat include selective
logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidding, on-site processing, and loading of trees or logs onto trucks or skeleton cars. Logging is the beginning of a supply chain ...
and various forms of anthropogenic development and activities, alien invasives, hunting and trapping for sport, medicine, food, and the pet trade. Additionally,
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
with adjacent species poses a threat to some populations.Molur, S., Brandon-Jones, D., Dittus, W., Eudey, A., Kumar, A., Singh, M., Feeroz, M. M., Chalise, M., Priya, P. and Walker, S. (2003)
''Status of South Asian Primates: Conservation Assessment and Management Plan Report''
. Workshop Report, 2003. Zoo Outreach Organization/CBSG-South Asia, Coimbatore, India.


Conservation

''Macaca assamensis'' is listed in CITES Appendix II. It is legally protected in all countries of occurrence. For the populations in India, the species is listed under Schedule II, part I of the Indian Wildlife Act.


References


External links


ARKive: Assam macaque (''Macaca assamensis'')
{{Taxonbar, from=Q738925 Assam macaque Primates of Asia Mammals of Bangladesh Mammals of Bhutan Mammals of Myanmar Mammals of China Mammals of India Mammals of Laos Mammals of Nepal Mammals of Thailand Mammals of Vietnam Fauna of Yunnan Fauna of Assam Assam macaque Taxa named by John McClelland (doctor)