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The Maastrichtian () is, in the ICS
geologic timescale The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochr ...
, the latest age (uppermost
stage Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * Sta ...
) of the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
Epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...
or
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', th ...
Series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used in ...
, the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
or System, and of the
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...
Era An era is a span of time defined for the purposes of chronology or historiography, as in the regnal eras in the history of a given monarchy, a calendar era used for a given calendar, or the geological eras defined for the history of Earth. Comp ...
or
Erathem __NOTOC__ In stratigraphy, paleontology, geology, and geobiology an erathem is the total stratigraphic unit deposited during a certain corresponding span of time during an era in the geologic timescale. It can therefore be used as a chronostra ...
. It spanned the interval from . The Maastrichtian was preceded by the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
and succeeded by the
Danian The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretace ...
(part of the Paleogene and
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
). The
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
(formerly known as the Cretaceous–
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
extinction event) occurred at the end of this age. In this
mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. I ...
, many commonly recognized groups such as non-avian
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s, plesiosaurs and
mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on ...
s, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous.


Stratigraphic definitions


Definition

The Maastrichtian was introduced into scientific literature by
Belgian Belgian may refer to: * Something of, or related to, Belgium * Belgians, people from Belgium or of Belgian descent * Languages of Belgium, languages spoken in Belgium, such as Dutch, French, and German *Ancient Belgian language, an extinct languag ...
geologist
André Hubert Dumont André — sometimes transliterated as Andre — is the French and Portuguese form of the name Andrew, and is now also used in the English-speaking world. It used in France, Quebec, Canada and other French-speaking countries. It is a variation ...
in 1849, after studying rock strata of the Chalk Group close to the
Dutch Dutch commonly refers to: * Something of, from, or related to the Netherlands * Dutch people () * Dutch language () Dutch may also refer to: Places * Dutch, West Virginia, a community in the United States * Pennsylvania Dutch Country People E ...
city of
Maastricht Maastricht ( , , ; li, Mestreech ; french: Maestricht ; es, Mastrique ) is a city and a municipality in the southeastern Netherlands. It is the capital and largest city of the province of Limburg. Maastricht is located on both sides of the ...
. These strata are now classified as the
Maastricht Formation The Maastricht Formation (Dutch: ''Formatie van Maastricht''; abbreviation: MMa), named after the city of Maastricht in the Netherlands, is a geological formation in the Netherlands and Belgium whose strata date back to the Late Cretaceous, within ...
- both
formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondar ...
and stage derive their names from the city. The Maastricht Formation is known for its fossils from this age, most notably those of the giant sea reptile ''
Mosasaurus ''Mosasaurus'' (; "lizard of the Meuse River") is the type genus (defining example) of the mosasaurs, an extinct group of aquatic squamate reptiles. It lived from about 82 to 66 million years ago during the Campanian and Maastrichtian sta ...
'', which in turn derives its name from the nearby river Maas (''mosa'' being Latin for the river Maas). The base of the Maastrichtian Stage is at the first appearance of ammonite species '' Pachydiscus neubergicus''. At the original type locality near Maastricht, the stratigraphic record was later found to be incomplete. A reference profile for the base was then appointed in a section along the Ardour river called ''Grande Carrière'', close to the village of
Tercis-les-Bains Tercis-les-Bains (; oc, Tèrcis) is a Communes of France, commune of the Landes (department), Landes Departments of France, department in Nouvelle-Aquitaine in southwestern France. Population See also *Communes of the Landes department Refer ...
in southwestern
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan area ...
. The top of the Maastrichtian Stage is defined to be at the
iridium Iridium is a chemical element with the symbol Ir and atomic number 77. A very hard, brittle, silvery-white transition metal of the platinum group, it is considered the second-densest naturally occurring metal (after osmium) with a density of ...
anomaly at the Cretaceous–Paleogene boundary (K–Pg boundary), which is also characterised by the
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
of many groups of life.


Subdivision

The Maastrichtian is commonly subdivided into two substages (Upper and Lower) and three ammonite
biozone In biostratigraphy, biostratigraphic units or biozones are intervals of geological strata that are defined on the basis of their characteristic fossil taxa, as opposed to a lithostratigraphic unit which is defined by the lithological properties ...
s. The biozones are (from young to old): *zone of '' Anapachydiscus terminus'' *zone of ''Anapachydiscus fresvillensis'' *zone of '' Pachydiscus neubergicus'' till ''Pachydiscus epiplectus'' The Maastrichtian is roughly coeval with the
Lancian The Lancian was a North American faunal stage of the Late Cretaceous. It was the final stage of the Cretaceous period in North America, lasting from approximately 70.6 to 66 million years ago. Geology Terrestrial sedimentary strata from the Judit ...
North American Land Mammal Age The North American land mammal ages (NALMA) establishes a geologic timescale for North American fauna beginning during the Late Cretaceous and continuing through to the present. These periods are referred to as ages or intervals (or stages when re ...
.


Palaeogeography and paleoclimate

The breakup of
Pangaea Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million y ...
was nearly complete in the Maastrichtian, with Australia beginning to break away from Antarctica and Madagascar breaking away from India. However, Arabia had not yet rifted away from Africa. North America was separated from Europe by rift basins, but sea floor spreading had not yet commenced between the two continents. The Pacific Plate was rapidly growing in size as the surrounding oceanic plates were consumed by subduction, and the Pacific-Izanagi Ridge was rapidly approaching Asia. Eruption of the Deccan Traps
large igneous province A large igneous province (LIP) is an extremely large accumulation of igneous rocks, including intrusive (sills, dikes) and extrusive (lava flows, tephra deposits), arising when magma travels through the crust towards the surface. The formation ...
began during the Maastrichtian, at around 67 million years ago. This is thought to be a consequence of India drifting over the
Réunion hotspot The Réunion hotspot is a volcanic hotspot which currently lies under the island of Réunion in the Indian Ocean. The Chagos-Laccadive Ridge and the southern part of the Mascarene Plateau are volcanic traces of the Réunion hotspot. The hotsp ...
. During the Maastrichtian, the global climate began to shift from the warm and humid climate of the Mesozoic to the colder and more arid climate of the Cenozoic. Variation of climate with latitude also became greater. This was likely caused by a major reorganization of oceanic circulation that took place at the boundary between the early and late Maastrichtian. This reorganization was triggered by the breach of tectonic barriers in the South Atlantic, permitting deep ocean water to begin circulating from the nascent North Atlantic to the south. This initiated
thermohaline circulation Thermohaline circulation (THC) is a part of the large-scale ocean circulation that is driven by global density gradients created by surface heat and freshwater fluxes. The adjective ''thermohaline'' derives from '' thermo-'' referring to temp ...
similar to that of the modern oceans. At the same time, the
Laramide orogeny The Laramide orogeny was a time period of mountain building in western North America, which started in the Late Cretaceous, 70 to 80 million years ago, and ended 35 to 55 million years ago. The exact duration and ages of beginning and end of the ...
drained the
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient sea ...
of North America, further contributing to global cooling.


Paleontology

Dinosaurs remained the dominant large terrestrial animals throughout the Maasastrichtian, though mammals with internal organs similar to modern mammals were also present. Both
ammonites Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttl ...
and pterosaurs were in serious decline during the Maastrichtian.


Dinosaurs


Birds

Several archaic clades of birds, such as Enantiornithes,
Ichthyornithes Ichthyornithes is an extinct group of toothed avialans very closely related to the common ancestor of all modern birds. They are known from fossil remains found throughout the late Cretaceous period of North America, though only two genera, ''Ich ...
,
Hesperornithes Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as '' Hesperorn ...
, and
Apsaravis ''Apsaravis'' is a Mesozoic bird genus from the Late Cretaceous. The single known species, ''Apsaravis ukhaana'', lived about 78 million years ago, in the Campanian age of the Cretaceous period. Its fossilized remains were found in the Camel's ...
persisted to the latest Maasatrichtian but became extinct during the Cretaceous-Paleogene extinction event.


Pterosaurs

Traditionally, pterosaur faunas of the Maastrichtian were assumed to be dominated by
azhdarchids Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cre ...
, with other pterosaur groups having become extinct earlier on. However, more recent findings suggest a fairly composite pterosaur diversity: at least six ("Nyctosaurus" ''lamegoi'', a Mexican humerus, a Jordan humerus and several taxa from Morocco) nyctosaurs date to this period, as do a few pteranodontids, and ''
Navajodactylus ''Navajodactylus'' (meaning "Navajo finger") is an extinct genus of pterodactyloid pterosaur from Late Cretaceous (late Campanian stage) deposits of the San Juan Basin in New Mexico, United States. Discovery The holotype specimen of ''Navajoda ...
'', tentatively assigned to Azhdarchidae, lacks any synapomorphies of the group. This seems to underscore a higher diversity of terminal Cretaceous pterosaurs than previously thought.


Flora

The radiation of
angiosperms Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants ...
(flowering plants) was well under way in the Maastrichtian. From 50% to 80% of all genera of land plants were angiosperms, though gymnosperms and
ferns A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except th ...
still covered larger areas of the land surface.


Notes


References


External links


GeoWhen Database - MaastrichtianLate Cretaceous timescale
at the website of the subcommission for stratigraphic information of the ICS
Stratigraphic chart of the Late Cretaceous
at the website of Norges Network of offshore records of geology and stratigraphy
Maastrichtian Microfossils: 60+ images of Foraminifera
{{Authority control *06 Geological ages Cretaceous geochronology Maastricht