Pteranodontidae
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Pteranodontidae
The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head (most famously seen in ''Pteranodon'' itself). The spectacularly-crested ''Nyctosaurus'' is sometimes included in this family, though usually placed in its own family, the Nyctosauridae (Nicholson & Lydekker, 1889). Modern researchers differ in their use of the concept. S. Christopher Bennett and Alexander Kellner have concluded that ''Nyctosaurus'' was not a pteranodontid. In 1994 Bennett defined a clade Pteranodontidae, also including species of the Anhangueridae. However, this definition has not been accepted by other workers. Alexander Kellner, for example, named several additional species for specimens previously classified as ''Pteranodon'', and placed ''P. sternbergi'' in a distinct genus, ''Geosternbergia''. Kellner re-defined Ptera ...
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Tethydraco
''Tethydraco'' is a genus of pterodactyloid pterosaur that lived during the Late Cretaceous period (Maastrichtian stage) of what is now the area of present Morocco, about 66 million years ago. ''Tethydraco'' was originally assigned to the family Pteranodontidae. Subsequently described material suggested that it was an azhdarchid, and possibly synonymous with ''Phosphatodraco''. The type and only species is ''T. regalis''. Discovery and naming Since 2015, a group of paleontologists has been acquiring pterosaur fossils from commercial Moroccan fossil traders, who obtain these from workers in the phosphate mines on the Khouribga plateau, which is located within the Ouled Abdoun Basin. The purpose of this project is to determine pterosaur diversity in the latest Cretaceous. From this stage, no Konservat-Lagerstätten are known, sites combining a large variety of species with exceptional preservation. It is in such sites that the vast majority of pterosaur fossils and taxa have bee ...
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Pteranodontia
Pteranodontia is an extinct group of ornithocheiroid pterodactyloid pterosaurs that lived during the Late Cretaceous period (Coniacian to Maastrichtian stages) of North America and Africa. They were some of the most advanced pterosaurs, and possessed highly specialized cranial crests that may have served as sexual attraction, with males having a much larger crest. Classification Pteranodontia was originally named by Marsh in 1876. In 2003, it was given a phylogenetic definition by David Unwin as the common ancestor of ''Pteranodon'' and ''Nyctosaurus'' plus all its descendants. Though Marsh had originally named this group based on the shared absence of teeth in those species, most analyses show that all of the traditional "ornithocheiroid" pterosaurs are also members of this clade. Below is a cladogram showing the phylogenetic placement of this group from Andres and Myers (2013). In 2018, Longrich, Martill, and Andres revisited the classification, and made a different phylogen ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Volgadraco
''Volgadraco'' ("Volga River dragon") is a genus of pterodactyloid pterosaur from the Upper Cretaceous of European Russia. ''Volgadraco'' was originally classified as an azhdarchid. However, recent studies have concluded that it may belong to either the family Nyctosauridae, or the family Pteranodontidae. ''Volgadraco'' is known from lower beak (holotype SGU, no. 46/104a) and postcranial fragments from the early Campanian-age Rybushka Formation of Saratov, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older Santonian or Turonian azhdarchids like ''Azhdarcho'' and '' Bakonydraco'' and later Maastrichtian azhdarchids like '' Quetzalcoatlus''. ''Volgadraco'' was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The type species is ''V. bogolubovi'', the specific name honouring Russian paleontologist Nikolai Nikolaevich Bogolubov. The authors consider the earlier named genus ''Bogolubovia'' to be a '' no ...
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Dawndraco
''Dawndraco'' is a controversial genus of pteranodontid pterosaur from the Late Cretaceous of North America. It is likely synonymous with the contemporary pteranodontid ''Geosternbergia.'' ''Dawndraco'' was named by Alexander W.A. Kellner in 2010. The type species is ''Dawndraco kanzai''. The generic name combines the Dawn deity of the Iroquois with a Latin ''draco'', "dragon". The specific name refers to the Kanza tribe of Kansas. ''Dawndraco'' is based on the holotype specimen UALVP 24238, a partial skeleton including an almost complete skull and lower jaws. It was recovered in 1974 by Richard C. Fox and Allen Lindoe from rocks of the lower part of the Smoky Hill Chalk Member of the Niobrara Formation in Utica, Kansas. These rocks date to the late Coniacian or early Santonian stage, about 86 million years ago. The specimen was earlier referred to ''Pteranodon sternbergi''. However, when Kellner in 2010 assigned this latter species to the genus ''Geosternbergia'', he ...
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Ornithostoma
''Ornithostoma'' (meaning "bird mouth") is a genus of pterodactyloid pterosaur that lived during the Early Cretaceous period (Albian stage) of Europe, around 110 million years ago. ''Ornithostoma'' was once thought to have been a senior synonym of the pteranodontid ''Pteranodon'' due to its toothless anatomy and prior naming. History In 1869, Harry Govier Seeley, cataloguing the fossils of the Sedgwick Museum at Cambridge, referred three snout fragments of toothless pterosaur specimens from the Lower Cretaceous Albian Cambridge Greensand of England to ''Ornithocheirus simus''. These fragments had in 1859 been described by Richard Owen (who considered the later holotype jaw fragment part of a metacarpal) and referred to ''Pterodactylus sedgwickii'' and ''Pterodactylus fittoni''. By 1871 Seeley had realised ''Ornithocheirus simus'' was a toothed form, different from the fragments. Therefore, he provisionally named them as a separate genus ''Ornithostoma'', the name derived from G ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the Chalk Group c ...
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Nyctosauridae
Nyctosauridae (meaning "night lizards" or "bat lizards") is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly Europe. It was named in 1889 by Henry Alleyne Nicholson and Richard Lydekker.Nicholson, H.A. and Lydekker, R. (1889). ''A manual of palaeontology for the use of students: with a general introduction on the principles of palæontology, Volume II''. Blackwood, 1889. Nyctosaurids are characterized by their lack of all but the wing finger. In most pterosaurs, the hand has four fingers, with the fourth elongated to support the wing, and the remaining three are usually small, clawed, and used in walking or climbing. The lack of functional fingers in nyctosaurids may suggest that they spent almost all of their time in the air, rarely walking on the ground. Nyctosaurids also possessed a distinctively enlarged crest for muscle attachment on their upper arm bone, or humerus, the ''deltopectoral crest'', hatchet shaped li ...
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Nyctosaurus
''Nyctosaurus'' (meaning "night lizard" or "bat lizard") is a genus of nyctosaurid pterosaur from the Late Cretaceous period of what is now the Niobrara Formation of the mid-western United States, which, during the time ''Nyctosaurus'' was alive, was covered in an extensive shallow sea. Some remains belonging to a possible ''Nyctosaurus'' species called ''N.lamegoi'' have been found in Brazil, making ''Nyctosaurus'' more diverse. The genus ''Nyctosaurus'' has had numerous species referred to it, though how many of these may actually be valid requires further study. At least one species possessed an extraordinarily large antler-like cranial crest. ''Nyctosaurus'' was a mid-sized pterosaur that lived along the shores of the Niobrara Formation of the United States, which back then was within a large inland sea called the Western Interior Seaway. It has been suggested that it would have flown similar to modern-day soaring birds such as albatrosses, which consisted of flying very lon ...
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Geosternbergia Sternbergi
''Geosternbergia'' is an extinct genus of pteranodontid pterodactyloid pterosaur from the Late Cretaceous geological period of North America. ''Geosternbergia'' was one of the largest pterosaur genera, and had a wingspan of up to . Discovery and history The first fossils of ''Geosternbergia'' were collected by American paleontologist George F. Sternberg in 1952 from the lower portion of the Niobrara Formation. The fossils of the animal looked similar to those of the species ''Pteranodon longiceps'', but the crests were set upright and in a slightly different position. In 1958, Sternberg and paleontologist Myrl V. Walker published a study about this peculiar find. In 1966, American paleontologist John Christian Harksen assigned the specimens found as a new species of ''Pteranodon'' called ''P. sternbergi'' due to its distinct upright crest that set it apart from ''P. longiceps''. Halsey W. Miller however, concluded a revision of the different species of ''Pteranodon'' in 19 ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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