Lagonimico
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''Lagonimico'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
New World monkey New World monkeys are the five families of primates that are found in the tropical regions of Mexico, Central and South America: Callitrichidae, Cebidae, Aotidae, Pitheciidae, and Atelidae. The five families are ranked together as the Ceboidea ( ...
s from the
Middle Miocene The Middle Miocene is a sub-epoch of the Miocene Epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0.05 Ma to 11.608 ± 0.005 Ma (million y ...
(
Laventan The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes ...
in the
South American land mammal age The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Mya (unit), Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referre ...
s; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of
La Venta La Venta is a pre-Columbian archaeological site of the Olmec civilization located in the present-day Mexican state of Tabasco. Some of the artifacts have been moved to the museum "Parque - Museo de La Venta", which is in nearby Villahermosa, ...
in the Honda Group of
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''L. conclucatus''.''Lagonimico conclucatus''
at
Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals ...
.org


Description

A nearly complete but badly crushed skull and mandible of ''Lagonimico'' were discovered in the
La Victoria Formation The Honda Group ( es, Grupo Honda, Tsh, Ngh) is a geological group of the Upper and Middle Magdalena Basins and the adjacent Central and Eastern Ranges of the Colombian Andes. The group, in older literature also defined as formation, is in its ...
, that has been dated to the
Laventan The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes ...
, about 13.5 to 12.9 Ma.Defler, 2004, p.32Kay, 1994, p.333 ''Lagonimico'', as ''
Micodon ''Micodon'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of Colomb ...
'' and '' Patasola magdalenae'', also from the Honda Group, have been attributed to the
Callitrichinae The Callitrichidae (also called Arctopitheci or Hapalidae) are a family of New World monkeys, including marmosets, tamarins, and lion tamarins. At times, this group of animals has been regarded as a subfamily, called the Callitrichinae, of the fa ...
.Takai et al., 2001, p.290 Features of the dentition suggest ''Lagonimico'' is a sister group to living
Callitrichinae The Callitrichidae (also called Arctopitheci or Hapalidae) are a family of New World monkeys, including marmosets, tamarins, and lion tamarins. At times, this group of animals has been regarded as a subfamily, called the Callitrichinae, of the fa ...
(''
Saguinus The tamarins are squirrel-sized New World monkeys from the family Callitrichidae in the genus ''Saguinus''. They are the first offshoot in the Callitrichidae tree, and therefore are the sister group of a clade formed by the lion tamarins, Goeld ...
'', ''
Leontopithecus The four species of lion tamarins or maned marmosets make up the genus ''Leontopithecus''. They are small New World monkeys named for the mane surrounding their face, similar to the mane of a lion. Description Living in the eastern rainforests ...
'', ''
Callithrix ''Callithrix'' is a genus of New World monkeys of the family Callitrichidae, the family containing marmosets and tamarins. The genus contains the Atlantic Forest marmosets. The name ''Callithrix'' is derived from the Greek words ''kallos'', me ...
'', and ''
Cebuella Pygmy marmosets are two species of small New World monkeys in the genus ''Cebuella''. They are native to rainforests of the western Amazon Basin in South America. These primates are notable for being the smallest monkeys in the world, at just o ...
''). These features include having elongate compressed lower incisors, a reduced P2 lingual moiety, and the absence of upper molar hypocones. The new taxon also has a relatively deep jaw, that rule it out of the direct ancestry of any living callitrichine. The orbits of ''L. conclucatus'' are small, suggesting diurnal habits. Inflated, low-crowned (bunodont) cheek teeth with short, rounded shearing crests, as well as premolar simplification and M3 size reduction, suggest fruit- or gum eating adaptations, as among many living callitrichines. Procumbent and slightly elongate lower incisors suggest this species could use its front teeth as a gouge, perhaps for harvesting tree gum. Estimates from jaw size suggest ''Lagonimico'' weighed about ,Pérez et al., 2013, p.9 about the size of ''
Callicebus ''Callicebus'' is a genus of monkeys known as titi monkeys. Historically, titis were monogeneric, comprising only the genus ''Callicebus'' Thomas, 1903. Owing to the great diversity found across titi monkey species, a new genus-level taxonomy wa ...
'', the living titi monkey of South America.Tejedor, 2013, p.29 Later research reduced the estimated weight to .Silvestro et al., 2017, p.14 Judged from tooth size and jaw length, ''Lagonimico'' would have been slightly smaller than ''Callicebus'', but still larger than ''
Callimico The Goeldi's marmoset or Goeldi's monkey (''Callimico goeldii'') is a small, South American New World monkey that lives in the upper Amazon basin region of Bolivia, Brazil, Colombia, and Peru. It is the only species classified in the genus ''C ...
'' or any living callitrichine. The upper first molar (M1) with a subtriangular outline with a narrow lingual side resembles that of the oldest New World primate discovered to date, the
Late Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "daw ...
'' Perupithecus'' from the Peruvian Amazon.Bond et al., 2015, p.538


Habitat

The Honda Group, and more precisely the "Monkey Beds", are the richest site for fossil primates in South America.Rosenberger & Hartwig, 2001, p.3 Other than most fossil primates found at La Venta, the specimens of ''Lagonimico'' do not come from the "Monkey Beds".Wheeler, 2010, p.137 It has been argued that the monkeys of the Honda Group were living in habitat that was in contact with the
Amazon Amazon most often refers to: * Amazons, a tribe of female warriors in Greek mythology * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon River, in South America * Amazon (company), an American multinational technology c ...
and
Orinoco Basin The Orinoco Basin is the part of South America drained by the Orinoco river and its tributaries. The Orinoco watershed covers an area of about 990000 km2, making it the third largest in South America, covering most of Venezuela and eastern p ...
s, and that La Venta itself was probably seasonally dry forest.Lynch Alfaro et al., 2015, p.520 The evolutionary separation from '' Aotus'' of ''Lagonimico'' has been placed in the Early Miocene at 17.5 Ma.Takai et al., 2001, p.304


See also

*
List of primates of Colombia The primates of Colombia include 41 extant species in 13 genera and five families. Additionally, 12 fossil species in 10 genera and five families have been identified in Colombia, mainly at the La Venta (Colombia), La Venta Lagerstätte of the Hond ...
* ''
Mohanamico ''Mohanamico'' is an extinct genus of New World monkeys from the Middle Miocene (Laventan in the South American land mammal ages; 13.8 to 11.8 Ma). Its remains have been found at the Konzentrat-Lagerstätte of La Venta in the Honda Group of Co ...
'' * ''
Patasola The Patasola or "one leg" is one of many legends in South American folklore about female monsters from the jungle, appearing to male hunters or loggers in the middle of the wilderness when they think about women. The Patasola appears in the form ...
''


References


Bibliography

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Further reading

* * {{Taxonbar, from=Q10315836 Prehistoric primate genera Prehistoric monkeys †Lagonimico Miocene genus first appearances Miocene extinctions Miocene primates of South America Laventan Neogene Colombia Fossils of Colombia Honda Group, Colombia Fossil taxa described in 1994