Los Llanos Formation
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Los Llanos Formation is a geological
formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondar ...
in the in La Rioja Province, northwestern Argentina whose strata date back to the Campanian stage of the Late Cretaceous. Los Llanos Formation over the years has been controversially described as ranging from Late Cretaceous to Miocene, but the Miocene succession was assigned to Las Mulitas Formation in 2019. The formation rests on top of the
Early Permian 01 or '01 may refer to: * The year 2001, or any year ending with 01 * The month of January * 1 (number) Music * '01 (Richard Müller album), 01'' (Richard Müller album), 2001 * 01 (Son of Dave album), ''01'' (Son of Dave album), 2000 * 01 (Urban ...
Los Sauces Los Sauces () is a Chilean town and commune in the Malleco Province, Araucanía Region. Demographics According to the 2002 census of the National Statistics Institute, Los Sauces spans an area of and has 7,581 inhabitants (3,847 men and 3,734 wo ...
and Patquía Formations and in parts on Ordovician crystalline basement. The maximum thickness is estimated at . The sandstones and conglomerates of the formation were deposited ij a
fluvial In geography and geology, fluvial processes are associated with rivers and streams and the deposits and landforms created by them. When the stream or rivers are associated with glaciers, ice sheets, or ice caps, the term glaciofluvial or fluviog ...
environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
. The formation crops out in the , where more than 90 titanosaurid nesting sites were discovered in Los Llanos Formation. The sites were encountered on top of areas characterized by hydrothermal activity as
geyser A geyser (, ) is a spring characterized by an intermittent discharge of water ejected turbulently and accompanied by steam. As a fairly rare phenomenon, the formation of geysers is due to particular hydrogeological conditions that exist only in ...
s and other vents, suggesting a preferred location for the incubation of the dinosaur eggs. Apart from fossil eggs, the formation has provided fossil flora and ostracods. The crocodyliform ''
Llanosuchus ''Llanosuchus'' is an extinct genus of notosuchian mesoeucrocodylian known from the Late Cretaceous Los Llanos Formation in Argentina. It contains a single species In biology, a species is the basic unit of classification and a taxonomi ...
'', described in 2016 from the formation, was named after Los Llanos Formation.


Description

The Los Llanos Formation was first formally described by Bodenbender in 1911.Bodenbender, 2011 The formation crops out in very localized patches in the south of La Rioja Province.Hechenleitner, 2017, p.23 Los Llanos Formation is found on the western and eastern slopes of the eponymous Sierra de Los Llanos, in the Sierra de Chepes, Sierra Ulapes,
Sierra de Velasco 250px, The western part of Sierra de Velasco captures during summer moist air from the Atlantic allowing some sub-tropical forests to grow on it. Sierra de Velasco is a mountain range in the Argentine province of La Rioja. Sierra de Velsco runs fr ...
and Sierra Brava.Hechenleitner, 2017, p.24 Between the original definition of "Los Estratos de Los Llanos" (''Los Llanos Beds'') by Bodenbender and later analysis during the twentieth century, a number of controversies arose. Rusconi (1936), Bordas (1941), Pascual (1954), Guiñazú (1962) and Zuzek (1978) assigned a Miocene age to the formation, based on
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
fossils found in the strata.Hechenleitner, 2017, p.25 This unit is presently known as the Las Mulitas Formation.Chiesa et al., 2019, p.292 The occurrence of typical Cretaceous charophytes confirms the assigned age for Los Llanos Formation. Los Llanos Formation
unconformably An unconformity is a buried erosional or non-depositional surface separating two rock masses or strata of different ages, indicating that sediment deposition was not continuous. In general, the older layer was exposed to erosion for an interval o ...
overlies the Permian Los Sauces Formation in the Sierra de Velasco and the Permian Patquía Formation or Ordovician granitic
basement A basement or cellar is one or more floors of a building that are completely or partly below the ground floor. It generally is used as a utility space for a building, where such items as the furnace, water heater, breaker panel or fuse box, ...
in other places. The maximum recorded thickness in the Sanagasta Geologic Park is ,Hechenleitner, 2017, p.26 while in the region of Tama the formation reaches an estimated thickness of .Hechenleitner, 2017, p.29 It comprises lightgrey and orange sandstones with basal
conglomerate Conglomerate or conglomeration may refer to: * Conglomerate (company) * Conglomerate (geology) * Conglomerate (mathematics) In popular culture: * The Conglomerate (American group), a production crew and musical group founded by Busta Rhymes ** Co ...
beds, deposited in a
fluvial In geography and geology, fluvial processes are associated with rivers and streams and the deposits and landforms created by them. When the stream or rivers are associated with glaciers, ice sheets, or ice caps, the term glaciofluvial or fluviog ...
environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
, which in the Sanagasta park is associated with hydrothermal activity. Hechenleitner, 2017, p.34 The more than 90
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
nests were found in direct association with these hydrothermal vents, suggesting a preferred incubation location of the mother dinosaurs.Hechenleitner, 2017, p.35


Paleontological significance

The formation has provided fossilized nesting sites attributed to a titanosaurid dinosaur. Research conducted in 2015 by Hechenleitner et al. include a comparison with the Cretaceous Sanpetru Formation of Hațeg paleo-island in Romania, the Tremp Formation of the Spanish
Pre-Pyrenees The Pre-Pyrenees are the foothills of the Pyrenees. Description As a mountainous system the Pre-Pyrenees are part of the Pyrenees. They run parallel to the main mountain range in a west to east direction. On the French side the Pyrenees's slop ...
and the Boseong Formation of the
Gyeongsang Basin Gyeongsang ( ko, 경상도, ''Gyeongsang-do''; ) was one of the eight provinces of Korea during the Joseon dynasty. Gyeongsang was located in the southeast of Korea. The provincial capital was Daegu. The region was the birthplace of the Kingdom ...
in South Korea.Hechenleitner et al., 2015, p.6


Fossil content

The following fossils were reported from the formation:Los Llanos Formation
at
Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals ...
.org
*
Dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
**
Aeolosaurini Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found ''Aeolosaurus'', ''Gondwanatitan'', ...
indet. ** Titanosauria indet. ** Abelisauridae indet. *
Reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s ** Crocodyliforms ***
Peirosauridae Peirosauridae is a Gondwanan family of mesoeucrocodylians that lived during the Cretaceous period. It was a clade of terrestrial crocodyliforms that evolved a rather dog-like form, and were terrestrial carnivores. It was phylogenetically defined ...
indet. ***'' Llanosuchus tamaensis''Fiorelli et al., 2016 * Oofossils ** Faveoloolithidae indet.Grellet-Tinner & Fiorelli, 2010 * Invertebrates ** Ostracods *** '' Ilyocypris triebeli''Carignano et al., 2013 *** '' Wolburgiopsis neocretacea'' * Flora ** '' Gobichara (Pseudoharrisichara) groeberi''De Sosa et al., 2017, p.34 ** '' Lychnothamnus (Pseudoharrisichara) tenuis'' ** '' Lychnothamnus (Pseudoharrisichara) walpurgica''


See also

* List of dinosaur-bearing rock formations * Allen Formation, Campanian to Maastrichtian fossiliferous formation of the
Neuquén Basin Neuquén Basin ( es, Cuenca Neuquina) is a sedimentary basin covering most of Neuquén Province in Argentina. The basin originated in the Jurassic and developed through alternating continental and marine conditions well into the Tertiary. The bas ...
* Anacleto Formation, Campanian ooliferous formation of the Neuquén Basin *
Angostura Colorada Formation The Angostura Colorada Formation is a Campanian to Maastrichtian geologic formation of the Neuquén Basin and North Patagonian Massif in the Río Negro Province of Argentina. Dinosaur remains diagnostic to the genus level are among the fossils tha ...
, Campanian to Maastrichtian fossiliferous formation of the
North Patagonian Massif The North Patagonian Massif or Somún Cura Massif (Spanish language, Spanish: ''Macizo Norpatagónico'', ''Macizo Nordpatagónico'' or ''Macizo de Somún Cura'') is a massif in northern Patagonia located in the Argentine provinces of Río Negro Pro ...
* La Colonia Formation, Campanian to Maastrichtian formation of the Cañadón Asfalto Basin * Colorado Formation, Campanian to Maastrichtian fossiliferous formation of the Colorado Basin * Lago Colhué Huapí Formation, Campanian to Maastrichtian fossiliferous formation of the Golfo San Jorge Basin


References


Bibliography

* * * * * * * * * {{cite LSA , last=Bodenbender , first=G , year=1911 , title=Constitución geológica de la parte meridional de La Rioja y regiones limítrofes , journal= Boletín de la Academia Nacional de Ciencias , volume=19 , pages=2-11 Geologic formations of Argentina Upper Cretaceous Series of South America Cretaceous Argentina Campanian Stage Sandstone formations Conglomerate formations Fluvial deposits Ooliferous formations Fossiliferous stratigraphic units of South America Paleontology in Argentina Geology of La Rioja Province, Argentina