Lomatia Fraseri
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''Lomatia fraseri'', commonly known as tree lomatia, forest lomatia or silky lomatia is a plant of the family
Proteaceae The Proteaceae form a family of flowering plants predominantly distributed in the Southern Hemisphere. The family comprises 83 genera with about 1,660 known species. Together with the Platanaceae and Nelumbonaceae, they make up the order Pro ...
native to eastern Australia. It grows as a shrub or small tree, reaching high, with highly variable leaves. The cream to white
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s appear over summer. It is found in
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
margins, gullies and heathland in mountainous regions of
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
and
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
. It regenerates from fire by regrowing from a
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
.


Description

''Lomatia fraseri'' grows as a tall shrub or small tree up to 8–11 metres (25–36 ft) high, though can be much smaller in exposed areas; on heathland in
Werrikimbe National Park The Werrikimbe National Park is a protected national park located in the catchment zone of the Upper Hastings River in New South Wales, Australia. Gazetted in 1975, the park is situated approximately north of Sydney, north-west of , and eas ...
it is reduced to a height of 50 cm (20 in), and can be high in exposed areas in Victoria. It has lanceolate to elliptic leaves which range from entire to deeply pinnatisect and are between 6.5 and 15 (rarely 18) cm long and 1 to 3.5 (rarely 5) cm wide. The leaf margins are generally toothed, though occasionally entire or deeply lobed. The leaves are highly variable, and can even be markedly different on the same plant. The upper surfaces are green and glabrous, while the undersides are yellow-green and silky. The individual flowers are arranged in
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s that are up to 12 cm (5 in) long. They can be more crowded than those of other lomatias. These white to cream
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s appear between December and February in the species' native range, followed by the development of 1.5 to 3 cm long dark grey follicles, which are ripe from April to October.


Taxonomy

Scottish botanist Robert Brown described ''Lomatia fraseri'' in his 1830 work ''
Supplementum primum Prodromi florae Novae Hollandiae ''Supplementum primum Prodromi florae Novae Hollandiae'' ("First supplement to the Prodromus of the flora of New Holland") is an 1830 supplement to Robert Brown's ''Prodromus florae Novae Hollandiae et Insulae Van Diemen''. It may be referred to ...
'', from a specimen that had been collected in the Sydney district (then known as Port Jackson) by Charles Fraser in 1818. George Bentham lumped ''L. fraseri'' and ''L ilicifolia'' together, treating them as one species in his 1870 work ''
Flora Australiensis ''Flora Australiensis: a description of the plants of the Australian Territory'', more commonly referred to as ''Flora Australiensis'', and also known by its standard abbreviation ''Fl. Austral.'', is a seven-volume flora of Australia published be ...
''. Hybrids have been recorded with the river lomatia (''
Lomatia myricoides ''Lomatia myricoides'' , commonly known as the river lomatia, is a shrub native to New South Wales and Victoria in southeastern Australia. Description ''Lomatia myricoides'' grows as a woody shrub or small tree, reaching high, or rarely up to ...
'') on the Southern Tablelands, resulting in plants resembling ''L. fraseri'' but less hairy, and with crinkle bush (''L. silaifolia'') in the
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
area, resulting in plants with highly variable leaf shapes and hairy leaf undersides. Hybrids with native holly ('' L. ilicifolia'') have been recorded, the resulting plants having variable hairiness and prickliness. Analysis of
chloroplast A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in ...
DNA showed that there is extensive hybridization between the five species (''L. arborescens'', ''L. fraseri'', ''L. ilicifolia'', ''L. myricoides'' and ''L. silaifolia'') of mainland southeastern Australia, though each is distinct enough to warrant species status.


Distribution and habitat

''Lomatia fraseri'' has a disjunct distribution, being found the New England district of northern New South Wales north to Tenterfield and from
Budawang National Park Budawang National Park is a national park in New South Wales, Australia, located approximately southwest of Sydney and north of Batemans Bay. It contains part of the Budawang Mountain Range. Budawang National Park is named after Mount Budaw ...
in southern New South Wales into eastern and central
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
to the
Otway Ranges The Great Otway National Park is a national park located in the Barwon South West region of Victoria, Australia. The national park is situated approximately southwest of Melbourne, in the Otway Ranges, a low coastal mountain range. It conta ...
. It is found in
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
margins in mountainous regions. Often found in gullies in Victoria, it grows more in open woodland and heath in New South Wales. ''Lomatia frasieri'' has been documented as being naturalised in New Zealand.


Ecology

''Lomatia fraseri'' regenerates after bushfire by resprouting from a
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
. Fieldwork in the Central Highlands of Victoria predicts populations will be adversely affected by clearfelling and salvage logging after bushfires there.


Cultivation and uses

The marked variation in leaf shape are an interesting horticultural feature. ''Lomatia fraseri'' grows in sun or preferably part-shade, benefiting from extra watering when young and from organic mulch. It is moderately frost-tolerant. Propagation is by seed or cutting.
Joseph Maiden Joseph Henry Maiden (25 April 1859 – 16 November 1925) was a botanist who made a major contribution to knowledge of the Australian flora, especially the genus ''Eucalyptus''. This botanist is denoted by the author abbreviation when citing ...
reported that its wood was light and hard, and easily worked. It had been used for furniture and window-frames in the 19th century.


References

{{Taxonbar, from=Q6669180 Flora of New South Wales Flora of Victoria (state) Plants described in 1830 fraseri Proteales of Australia