Lithospermum Decipiens

''Lithospermum'' is a genus of plants belonging to the family Boraginaceae. The genus is distributed nearly worldwide, but most are native to the Americas and the center of diversity is in the southwestern United States and Mexico. Species are known generally as gromwells or stoneseeds.

Taxonomy

There are about 50, to 60 species in the genus.Cohen, J. I., (2012)
Comparative floral development in ''Lithospermum'' (Boraginaceae) and implications for the evolution and development of heterostyly.
''American Journal of Botany'' 99(5), 797–805.

Some species, such as ''Lithospermum arvense'', are sometimes classified in the genus ''Buglossoides'', but that genus is subsumed into ''Lithospermum'' by works such as the ''Flora of China''. In addition, a 2009 molecular study showed that the genus ''Onosmodium'' should be included within ''Lithospermum''.

Species include:

*''Lithospermum arvense'' (syn. ''Buglossoides arvensis'') – field gromwell, corn gromwell
*''Lithospermum azuayensis''Weigend, M., et al. (2010)
Five new species of ''Lithospermum'' L.(Boraginaceae tribe Lithospermeae) in Andean South America: another radiation in the Amotape-Huancabamba zone.
''Taxon'' 59(4), 1161–1179.
*''Lithospermum bejariense'' – western marbleseed
*'' Lithospermum bolivariensis''
*''Lithospermum californicum'' – California stoneseed
*''Lithospermum calycosum'' – Chinati stoneseed
*''Lithospermum canescens'' – hoary puccoon, Indian-paint
*'' Lithospermum caroliniense'' – Carolina gromwell, hairy puccoon
*'' Lithospermum cobrense'' – smooththroat stoneseed
*'' Lithospermum confine'' – Arizona stoneseed
*'' Lithospermum cuzcoensis''
*'' Lithospermum erythrorhizon'' – purple gromwell, red-root gromwell, 紫草 ''zicao'' ( Pinyin: zǐcǎo), 紫草 ''murasaki･sō'' ( Japanese)
*'' Lithospermum hancockianum''
*'' Lithospermum incisum'' – narrowleaf stoneseed, fringed gromwell
*'' Lithospermum latifolium'' – American stoneseed
*''Lithospermum leymebambensis''
*''Lithospermum macbridei''
*'' Lithospermum matamorense'' – rough stoneseed
*''Lithospermum mirabile'' – San Antonio stoneseed
*''Lithospermum molle'' – softhair marbleseed
*''Lithospermum multiflorum'' – manyflowered stoneseed
*''Lithospermum obtusifolium'' – roundleaf stoneseed
*''Lithospermum officinale'' – common gromwell, European stoneseed
*''Lithospermum parksii'' – Parks' stoneseed
*''Lithospermum parviflorum'' – eastern prairie marbleseed
*'' Lithospermum purpurocaeruleum'' (syn. ''Buglossoides purpurocaerulea'') – purple gromwell
*''Lithospermum rodriguezii''
*''Lithospermum ruderale'' – western stoneseed, Columbia puccoon, wayside gromwell, whiteweed
*'' Lithospermum tuberosum'' – southern stoneseed, tuberous gromwell
*''Lithospermum viride'' – green stoneseed

Ecology

''Lithospermum'' leaves are eaten by the caterpillars of certain Lepidoptera, such as the moth ''Ethmia pusiella'' which has been recorded on ''L. officinale''.

Uses

The dried root of ''Lithospermum erythrorhizon'' is a Chinese herbal medicine with various antiviral and biological activities, including inhibition of human immunodeficiency virus type 1 (HIV-1). '' Lithospermum erythrorhizon'' is native to Japan, where it has been traditionally used to make a purple dye. In southwestern North America, a species of this genus was used as a contraceptive by the Shoshone Native American tribe.

Fossil record

7 petrified nutlets and nutlet fragments of a ''Lithospermum'' species have been described from the Late Miocene Ash Hollow Formation, Ogallala Group, five km south of Martin in Bennett County, South Dakota. †''Lithospermum dakotense'' sp. nov. shows similarities in size, shape, attachment and epidermal cell patterns to extant ''Lithospermum'' species. The fossil nutlets were preserved in various stages of maturity. The fossils closely resemble the nutlets of '' Lithospermum caroliniense'' and '' Lithospermum incisum''.Amer. J. Bot. 74(11): 1690-1693. 1987. A Fossil Lithospermum (Boraginaceae) from the Tertiary of South Dakota by Mark L. Gabel

References

External links

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{{Taxonbar, from=Q932776

Boraginaceae genera