List Of Pyrenula Species
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The genus '' Pyrenula'' consists of
crustose lichen Crustose lichens are lichens that form a crust which strongly adheres to the substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichens consists of a cor ...
s that usually grow on smooth, shaded bark. About 750
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
have been named in the genus, although the majority of these names have been excluded from the genus as they have been transferred to other genera, or are considered
synonyms A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
. In his world
key Key or The Key may refer to: Common meanings * Key (cryptography), a piece of information that controls the operation of a cryptography algorithm * Key (lock), device used to control access to places or facilities restricted by a lock * Key (map ...
to the ''Pyrenula'' species, published in 2012,
André Aptroot André Aptroot (Heemskerk, 1961) is a Dutch mycologist and lichenologist. In 1993 he did his PhD at the University of Utrecht under the supervision of Robbert Gradstein (nl). His dissertation was titled "Systematic studies on pyrenocarpous li ...
accepted 169 species, including 7 not yet formally described. ,
Species Fungorum ''Index Fungorum'' is an international project to index all formal names (scientific names) in the fungus kingdom. the project is based at the Royal Botanic Gardens, Kew, one of three partners along with Landcare Research and the Institute of Mi ...
accepts 168 species of ''Pyrenula''.


A

*'' Pyrenula abditicarpa'' – Brazil *'' Pyrenula acutispora'' *'' Pyrenula adacta'' *'' Pyrenula aggregataspistea'' – South America *'' Pyrenula albonigra'' – Brazil *'' Pyrenula andina'' *'' Pyrenula annulata'' *'' Pyrenula anomala'' *'' Pyrenula arthoniotheca'' – India *'' Pyrenula asahinae'' – Japan *'' Pyrenula aspistea'' *'' Pyrenula astroidea'' *'' Pyrenula aurantiacorubra'' – Brazil *'' Pyrenula aurantioinspersa'' – South America *'' Pyrenula aurantiopileata'' – Thailand *'' Pyrenula aurantiothallina'' – Brazil


B

*'' Pyrenula baileyi'' *'' Pyrenula bataanensis'' *'' Pyrenula bicuspidata'' *'' Pyrenula biseptata'' *'' Pyrenula bispora'' – Brazil *'' Pyrenula borneensis'' – Borneo *'' Pyrenula breutelii''


C

*'' Pyrenula caraibica'' – Panama *'' Pyrenula celaticarpa'' – Brazil *'' Pyrenula cerina'' *'' Pyrenula ceylonensis'' *'' Pyrenula chloroplaca'' – Australia *'' Pyrenula chlorospila'' *'' Pyrenula cinnabarina'' – Brazil *'' Pyrenula clavatispora''
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
*'' Pyrenula coccinea'' – Brazil *'' Pyrenula concatervans'' *'' Pyrenula confinis'' *'' Pyrenula conspurcata'' *'' Pyrenula convexa'' *'' Pyrenula cornutispora'' – South America *'' Pyrenula corticata'' *'' Pyrenula coryli'' *'' Pyrenula crassiuscula'' – Brazil *'' Pyrenula cruenta'' *'' Pyrenula cruentata'' *'' Pyrenula cryptothelia'' – Panama


D

*'' Pyrenula darjeelingensis'' – India *'' Pyrenula decumbens'' *'' Pyrenula defossa'' *'' Pyrenula dermatodes'' *'' Pyrenula diamantinensis'' – Brazil *'' Pyrenula duplicans''


E

*'' Pyrenula endocrocea'' – Philippines


F

*'' Pyrenula fibrata'' *'' Pyrenula filiformis'' – New Caledonia *'' Pyrenula finitima'' *'' Pyrenula flavoinspersa'' – South America *'' Pyrenula fuscoluminata'' *'' Pyrenula fusispora'' *'' Pyrenula fusoluminata'' – Brazil


G

*'' Pyrenula galactina'' *'' Pyrenula gibberulosa'' *'' Pyrenula globifera'' *'' Pyrenula guyanensis'' – South America


H

*'' Pyrenula hawaiiensis'' – Hawaii *'' Pyrenula hibernica'' – Panama *'' Pyrenula howeana'' – Australia


I

*'' Pyrenula immersa'' *'' Pyrenula indusiata'' *'' Pyrenula infracongruens'' *'' Pyrenula infraleucotrypa'' – South America *'' Pyrenula inframamillana'' – South America *'' Pyrenula infrastroidea'' – South America *'' Pyrenula inspersicollaris'' – Brazil *'' Pyrenula inspersoleucotrypa'' – Argentina


L

*'' Pyrenula laevigata'' *'' Pyrenula laii'' – Taiwan *'' Pyrenula leptaleoides'' – Brazil *'' Pyrenula leucostoma'' *'' Pyrenula lilacina'' – Brazil *'' Pyrenula luteopruinosa'' – Panama *'' Pyrenula lyonii''


M

*'' Pyrenula macrospora'' *'' Pyrenula macularis'' *'' Pyrenula mamillana'' *'' Pyrenula maritima'' – South America *'' Pyrenula mastigophora'' South Solomons *'' Pyrenula mastophora'' *'' Pyrenula mastophorizans'' *'' Pyrenula mattickiana'' – South America *'' Pyrenula melaleuca'' *'' Pyrenula microcarpa'' *'' Pyrenula microcarpoides'' *'' Pyrenula micromma'' *'' Pyrenula microtheca'' *'' Pyrenula minae'' *'' Pyrenula minoides'' – South America *'' Pyrenula minutispora'' – Brazil *'' Pyrenula minutissima'' – Iran *'' Pyrenula monospora'' – South America *'' Pyrenula montocensis'' *'' Pyrenula multicolorata'' – Sri Lanka *'' Pyrenula muriciliata''
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It incl ...
*'' Pyrenula musaespora'' – Brazil


N

*'' Pyrenula neojaponica'' – Japan *'' Pyrenula neolaevigata'' – Japan *'' Pyrenula neosandwicensis'' *'' Pyrenula nigrocincta'' *'' Pyrenula nitida'' *'' Pyrenula nitidans'' *'' Pyrenula nitidella'' *'' Pyrenula nitidula''


O

*'' Pyrenula occidentalis'' *'' Pyrenula occulta'' *'' Pyrenula ocellulata'' – Sri Lanka *'' Pyrenula ochraceoflava'' *'' Pyrenula oxysporiza''


P

*'' Pyrenula papillifera'' *'' Pyrenula paraminarum'' – South America *'' Pyrenula parvinuclea'' *'' Pyrenula perfecta'' – South America *'' Pyrenula pinguis'' *'' Pyrenula platystoma'' *'' Pyrenula plicata'' – South America *'' Pyrenula porinoides'' *'' Pyrenula prostrata'' – New Zealand *'' Pyrenula pseudobufonia'' *'' Pyrenula punctoleucotrypa'' – Argentina *'' Pyrenula pyrenastroides'' – New Zealand *'' Pyrenula pyrenuloides''


Q

*'' Pyrenula quartzitica'' – Brazil *'' Pyrenula quassiicola''


R

*'' Pyrenula ravenelii'' *'' Pyrenula reebiae'' – North America *'' Pyrenula reginae'' – Brazil *'' Pyrenula relicta'' *'' Pyrenula rhomboidea'' – Brazil *'' Pyrenula rinodinospora'' – Papua New Guinea *'' Pyrenula rubroanomala'' *'' Pyrenula rubroinspersa'' – South America *'' Pyrenula rubrojavanica'' – Java *'' Pyrenula rubrolateralis'' – Brazil *'' Pyrenula rubromamillana'' – Brazil *'' Pyrenula rubronitidula'' – South America *'' Pyrenula rubrostigma'' – South America *'' Pyrenula rubrostoma''


S

*'' Pyrenula sanguinea'' – Brazil *'' Pyrenula sanguineomeandrata'' *'' Pyrenula sanguineostiolata'' *'' Pyrenula santensis'' *'' Pyrenula schiffneri'' *'' Pyrenula segregata'' *'' Pyrenula seminuda'' – South America *'' Pyrenula sexlocularis'' *'' Pyrenula sexluminata'' *'' Pyrenula shirleyana'' – Australia *'' Pyrenula sipmanii'' – South Korea *'' Pyrenula spissitunicata'' – South Solomons *'' Pyrenula subcongruens'' *'' Pyrenula subcylindrica'' – India *'' Pyrenula subsoluta'' *'' Pyrenula subumbilicata'' – Australia *'' Pyrenula subvariabilis'' *'' Pyrenula subvariolosa'' – Australia *'' Pyrenula supracongruens'' *'' Pyrenula supralaetior'' – Brazil


T

*'' Pyrenula tetraspora'' – South America *'' Pyrenula thailandica'' – Papua New Guinea; India; Thailand *'' Pyrenula tokyoensis'' – Japan *'' Pyrenula triangularis'' – South America


V

*'' Pyrenula velatior'' *'' Pyrenula vermicularis'' – Japan *'' Pyrenula violaceastroidea'' – Brazil *'' Pyrenula viridipyrgilla'' – South America


W

*'' Pyrenula warmingii'' *'' Pyrenula welwitschii''


X

*'' Pyrenula xanthinspersa'' *'' Pyrenula xanthoglobulifera'' – Brazil *'' Pyrenula xanthominuta'' – Australia


Former ''Pyrenula'' species

Many taxa once placed in ''Pyrenula'' have been moved to other genera or have been synonymized; these former ''Pyrenula'' species are listed here. *''Pyrenula achariana'' = '' Melanotheca achariana'' *''Pyrenula achroopora'' = '' Pyrenula dermatodes'' *''Pyrenula addubitans'' = '' Pleospora addubitans'' *''Pyrenula aenea'' = '' Pseudosagedia aenea'' *''Pyrenula aethiobola'' = '' Verrucaria aethiobola'' *''Pyrenula albissima'' = ''
Leptorhaphis epidermidis ''Leptorhaphis'' is a genus of lichens in the family Naetrocymbaceae The Naetrocymbaceae are a family (biology), family of fungi with an uncertain taxonomic placement in the class (biology), class Dothideomycetes. References Pleosporale ...
'' *''Pyrenula americana'' = '' Anisomeridium americanum'' *''Pyrenula analepta'' = '' Arthopyrenia analepta'' *''Pyrenula annularis'' = '' Astrothelium annulare'' *''Pyrenula aractina'' = '' Hydropunctaria aractina'' *''Pyrenula areolata'' = '' Staurothele areolata'' *''Pyrenula arthonioides'' = '' Pyrenula arthoniotheca'' *''Pyrenula biformis'' = '' Microthelia biformis'' *''Pyrenula canellae-albae'' = '' Sulcopyrenula canellae-albae'' *''Pyrenula carpinea'' = '' Segestria carpinea'' *''Pyrenula cartilaginea'' = '' Astrothelium cartilagineum'' *''Pyrenula catalepta'' = '' Verrucaria aethiobola'' *''Pyrenula catervaria'' = ''
Trypethelium variolosum ''Trypethelium'' is a genus of lichenized fungi in the family Trypetheliaceae. The widespread genus contains about 50 species that are predominantly found in tropical areas. ''Trypethelium'' was circumscribed by German botanist Kurt Polycarp Joa ...
'' *''Pyrenula cerasi'' = '' Arthopyrenia cerasi'' *''Pyrenula ceratina'' = '' Astrothelium ceratinum'' *''Pyrenula chilensis'' = '' Parmentaria chilensis'' *''Pyrenula chlorotica'' = '' Pseudosagedia chlorotica'' *''Pyrenula cinchonae'' = '' Constrictolumina cinchonae'' *''Pyrenula circumrubens'' = '' Pyrenula cruenta'' *''Pyrenula clandestina'' = '' Clandestinotrema clandestinum'' *''Pyrenula clopima'' = '' Staurothele clopima'' *''Pyrenula coactella'' = '' Melanothecopsis coactella'' *''Pyrenula collospora'' = '' Bogoriella collospora'' *''Pyrenula diluta'' = '' Pseudopyrenula diluta'' *''Pyrenula discissa'' = '' Phaeotrema discissum'' *''Pyrenula discolor'' = '' Ampliotrema discolor'' *''Pyrenula elaeina '' = '' Verrucaria elaeina'' *''Pyrenula emergens'' = '' Pyrenula pinguis'' *''Pyrenula endococcoidea'' = '' Phaeospora rimosicola'' *''Pyrenula epidermidis'' = ''
Leptorhaphis epidermidis ''Leptorhaphis'' is a genus of lichens in the family Naetrocymbaceae The Naetrocymbaceae are a family (biology), family of fungi with an uncertain taxonomic placement in the class (biology), class Dothideomycetes. References Pleosporale ...
'' *''Pyrenula fallaciosa'' = '' Arthopyrenia fallaciosa'' *''Pyrenula flaventior'' = '' Pyrenula mastophora'' *''Pyrenula fraxini'' = '' Naetrocymbe fraxini'' *''Pyrenula fuliginea'' = '' Thelignya lignyota'' *''Pyrenula funckii'' = '' Verrucaria funckii'' *''Pyrenula fusca'' = '' Pyrenula anomala'' *''Pyrenula gaudichaudii'' = '' Nigrovothelium tropicum'' *''Pyrenula gelatinosa'' = '' Agonimia gelatinosa'' *''Pyrenula gemmata'' = ''
Acrocordia gemmata ''Acrocordia gemmata'' is a species of lichen belonging to the family Monoblastiaceae. Synonym: * ''Arthopyrenia gemmata'' (Ach.) A.Massal., 1852 References {{Taxonbar, from=Q10509861 Dothideomycetes ...
'' *''Pyrenula gemmifera'' = '' Tichothecium gemmiferum'' *''Pyrenula gibbosa'' = ''
Rimularia gibbosa ''Rimularia'' is a genus of lichenized fungi in the family Trapeliaceae. ''Rimularia'' was circumscribed by Finnish botanist William Nylander in 1868. Species *'' Rimularia actinostoma'' *'' Rimularia albotessellata'' *'' Rimularia applanata'' ...
'' *''Pyrenula glabra'' = '' Swinscowia glabra'' *''Pyrenula glabratula'' = '' Pyrenula dermatodes'' *''Pyrenula guayaci'' = '' Parapyrenis guayaci'' *''Pyrenula harrisii'' = '' Pyrenula occidentalis'' *''Pyrenula henatomma'' = '' Ocellularia henatomma'' *''Pyrenula hyalospora'' = '' Lithothelium hyalosporum'' *''Pyrenula hydrela'' = '' Verrucaria hydrela'' *''Pyrenula infernalis'' = '' Megalotremis infernalis'' *''Pyrenula interjungens'' = '' Pseudosagedia interjungens'' *''Pyrenula kakouettae'' = '' Pyrenula acutispora'' *''Pyrenula kunthii'' = '' Pyrenula mamillana'' *''Pyrenula leucocephala'' = '' Lecanactis abietina'' *''Pyrenula leucoplaca'' = '' Eopyrenula leucoplaca'' *''Pyrenula libricola'' = '' Pyrenula leucostoma'' *''Pyrenula lignyota'' = '' Thelignya lignyota'' *''Pyrenula lithina'' = '' Staurothele fissa'' *''Pyrenula marcida'' = '' Astrothelium marcidum'' *''Pyrenula margacea'' = '' Verrucaria margacea'' *''Pyrenula marginata'' = '' Pyrenula mamillana'' *''Pyrenula martinicana'' = '' Pyrenula caraibica'' *''Pyrenula mastoidea'' = '' Clathroporina mastoidea'' *''Pyrenula megalospora'' = '' Acrocordia megalospora'' *''Pyrenula melanospora'' = '' Mycomicrothelia melanospora'' *''Pyrenula micromma'' = '' Pyrenula occulta'' *''Pyrenula microscopica'' = '' Mycoporopsis microscopica'' *''Pyrenula microthelia'' = '' Roselliniella microthelia'' *''Pyrenula neoculata'' = '' Anthracothecium oculatum'' *''Pyrenula nigrescens'' = '' Verrucaria nigrescens'' *''Pyrenula nitens'' = '' Architrypethelium nitens'' *''Pyrenula nitidella'' f. ''chlorospila'' = '' Pyrenula chlorospila'' *''Pyrenula obovata'' = '' Bogoriella obovata'' *''Pyrenula ocellata'' = '' Polymeridium ocellatum'' *''Pyrenula ochraceoflavens'' = '' Pyrenula ochraceoflava'' *''Pyrenula olivacea'' = '' Pseudosagedia borreri'' *''Pyrenula olivacea'' = '' Arthopyrenia analepta'' *''Pyrenula oxyspora'' = ''
Leptorhaphis epidermidis ''Leptorhaphis'' is a genus of lichens in the family Naetrocymbaceae The Naetrocymbaceae are a family (biology), family of fungi with an uncertain taxonomic placement in the class (biology), class Dothideomycetes. References Pleosporale ...
'' *''Pyrenula oxyspora'' = '' Pyrenula oxysporiza'' *''Pyrenula papularis'' = '' Thelidium papulare'' *''Pyrenula perpusilla'' = '' Endococcus rugulosus'' *''Pyrenula pertusarioidea'' = '' Polyblastiopsis pertusarioidea'' *''Pyrenula planorbis'' = '' Constrictolumina planorbis'' *''Pyrenula punctiformis'' = '' Naetrocymbe punctiformis'' *''Pyrenula pupula'' = '' Astrothelium pupula'' *''Pyrenula pygmaea'' = '' Muellerella pygmaea'' *''Pyrenula quercus'' = '' Cyrtidula quercus'' *''Pyrenula quinqueseptata'' = '' Polymeridium quinqueseptatum'' *''Pyrenula quinqueseptata'' = '' Pyrenula sexluminata'' *''Pyrenula rhyponta'' = '' Naetrocymbe rhyponta'' *''Pyrenula salicis'' = '' Arthopyrenia salicis'' *''Pyrenula sphaeroides'' = '' Zignoella sphaeroides'' *''Pyrenula staurospora'' = '' Sulcopyrenula staurospora'' *''Pyrenula subandamanica'' = '' Parmentaria andamanensis'' *''Pyrenula subfarinosa'' = '' Phaeotrema subfarinosum'' *''Pyrenula submersa'' = '' Verrucaria hydrela'' *''Pyrenula subprostans'' = '' Anisomeridium subprostans'' *''Pyrenula subsimplex'' = '' Melanotheca vainioensis'' *''Pyrenula tetracerae'' = '' Porina tetracerae'' *''Pyrenula thelena'' = '' Bogoriella thelena'' *''Pyrenula tremulae'' = ''
Leptorhaphis tremulae ''Leptorhaphis'' is a genus of lichens in the family Naetrocymbaceae. Members of the genus ''Leptorhaphis'' are commonly called birchbark dot lichens. Species * '' Leptorhaphis atomaria'' * ''Leptorhaphis epidermidis ''Leptorhaphis'' is a ...
'' *''Pyrenula tropica'' = '' Nigrovothelium tropicum'' *''Pyrenula uberina'' = '' Architrypethelium uberinum'' *''Pyrenula umbonata'' = '' Pyrenocarpon thelostomum'' *''Pyrenula umbrata'' = '' Thelotrema umbratum'' *''Pyrenula variolosa'' = '' Anthracothecium variolosum'' *''Pyrenula ventosicola'' = '' Muellerella ventosicola'' *''Pyrenula vermicellifera'' = '' Opegrapha vermicellifera'' *''Pyrenula verrucosa'' = '' Polyblastia verrucosa'' *''Pyrenula wallrothii'' = '' Mycomicrothelia wallrothii'' *''Pyrenula zwackhii'' = '' Thelidium zwackhii''


Notes


References

{{Reflist, colwidth=30em, refs= {{cite journal , last1=Acharius , first1=E. , year=1814 , title=Monographie der Lichenen-Gattung ''Pyrenula'' mit Abbildungen aller bisher bekannten Arten , journal=Magazin der Gesellschaft Naturforschenden Freunde Berlin , volume=6 , issue=1 , pages=3–25 , language=de {{cite journal , last1=Aptroot , first1=A. , year=2002 , title=New and interesting lichens and lichenicolous fungi in Brazil , journal=Fungal Diversity , volume=9 , pages=15–45 {{cite journal , last1=Aptroot , first1=A. , year=2003 , title=Pyrenocarpous lichens and related non-lichenized ascomycetes from Taiwan , journal=Journal of the Hattori Botanical Laboratory , volume=93 , pages=155–173 {{cite journal , last1=Aptroot , first1=A. , year=2007 , title=New species, combinations, lectotypifications and synonyms in Australia Pyrenulaceae , journal=Australasian Lichenology , volume=60 , pages=34–41 {{cite journal , last1=Aptroot , first1=A. , last2=Saipunkaew , first2=W. , last3=Sipman , first3=H.J.M. , last4=Sparrius , first4=L.B. , last5=Wolseley , first5=P.A. , year=2007 , title=New lichens from Thailand, mainly microlichens from Chiang Mai , journal=Fungal Diversity , volume=24 , pages=75–134 , url=https://www.researchgate.net/publication/228515656 {{cite journal , last1=Aptroot , first1=André , last2=Lücking , first2=Robert , last3=Sipman , first3=Harrie J.M. , last4=Umana , first4=Loengrin , last5=Chaves , first5=José Luis , year=2008 , title=Pyrenocarpous lichens with bitunicate asci , journal=Bibliotheca Lichenologica , volume=97 , publisher=J. Cramer , location=Berlin/Stuttgart , pages=89–128 , isbn=978-3-443-58076-6 {{cite journal , last=Aptroot , first=André , title=A world key to the species of ''Anthracothecium'' and ''Pyrenula'' , journal=The Lichenologist , volume=44 , issue=1 , date=1 January 2012 , doi=10.1017/s0024282911000624 , pages=5–53, s2cid=86565959 {{cite journal , last1=Aptroot , first1=André , last2=Schumm , first2=Felix , last3=Cáceres , first3=Marcela E.S. , title=Six new species of ''Pyrenula'' from the tropics , journal=The Lichenologist , volume=44 , issue=5 , year=2012 , doi=10.1017/s0024282912000254 , pages=611–618, s2cid=84646514 {{cite journal , last1=Aptroot , first1=André , last2=Valadbeigi , first2=Tahereh , last3=Sipman , first3=Harrie J.M. , title=A new species and new records of the lichen genus ''Pyrenula'' from Iran , journal=The Lichenologist , volume=44 , issue=4 , year=2012 , doi=10.1017/s0024282912000023 , pages=445–448, s2cid=86210236 {{cite journal , last1=Aptroot , first1=André , last2=Cáceres , first2=Marcela Eugenia da Silva , title=Pyrenocarpous lichens (except Trypetheliaceae) in Rondônia , journal=The Lichenologist , volume=45 , issue=6 , year=2013 , doi=10.1017/s0024282913000534 , pages=763–785, s2cid=87480231 {{cite journal , last1=Aptroot , first1=André , last2=Sipman , first2=Harrie J.M. , last3=Cáceres , first3=Marcela Eugenia da Silva , title=Twenty-one new species of ''Pyrenula'' from South America, with a note on over-mature ascospores , journal=The Lichenologist , volume=45 , issue=2 , year=2013 , doi=10.1017/s0024282912000734 , pages=169–198, s2cid=84245404 {{cite journal , last=Aptroot , first=André , title=Two new genera of Arthoniales from New Caledonia and the Solomon Islands, with the description of eight further species , journal=The Bryologist , volume=117 , issue=3 , year=2014 , doi=10.1639/0007-2745-117.3.282 , pages=282–289, s2cid=85916369 {{cite journal , last1=Aptroot , first1=André , last2=Ferraro , first2=Lidia Itatí , last3=Cáceres , first3=Marcela Eugenia da Silva , title=New pyrenocarpous lichens from NE Argentina , journal=The Lichenologist , volume=46 , issue=1 , year=2014 , doi=10.1017/s0024282913000716 , pages=95–102, s2cid=88170280 {{cite journal , last1=Aptroot , first1=Andre , last2=Andrade , first2=Dannyelly Santos , last3=Mendonça , first3=Cléverton , last4=Lima , first4=Edvaneide Leandro De , last5=Cáceres , first5=Marcela Eugenia da Silva , title=Ten new species of corticolous pyrenocarpous lichens from NE Brazil , journal=Phytotaxa , volume=197 , issue=3 , year=2015 , doi=10.11646/phytotaxa.197.3.3 , pages=197–206 {{cite journal , last1=Aptroot , first1=André , last2=Sipman , first2=Harrie J.M. , last3=Mercado Diaz , first3=Joel Alejandro , last4=Mendonça , first4=Cléverton de Oliveira , last5=Feuerstein , first5=Shirley Cunha , last6=Cunha-Dias , first6=Iane Paula Rego , last7=Pereira , first7=Thamires Almeida , last8=Cáceres , first8=Marcela Eugenia da Silva , title=Eight new species of Pyrenulaceae from the Neotropics, with a key to 3-septate ''Pyrgillus'' species , journal=The Lichenologist , volume=50 , issue=1 , year=2018 , doi=10.1017/s0024282917000573 , pages=77–87, s2cid=90118198 {{cite journal , last1=Arnold , first1=F. , year=1887 , title=Lichenologische Fragmente. XXVIII , journal=Flora (Regensburg) , volume=70 , pages=145–164 {{cite journal , last1=Cáceres , first1=Marcela E.S. , last2=Aptroot , first2=André , last3=Nelsen , first3=Matthew P. , last4=Lücking , first4=Robert , title=''Pyrenula sanguinea'' (lichenized Ascomycota: Pyrenulaceae), a new species with unique, trypethelioid ascomata and complex pigment chemistry , journal=The Bryologist , volume=116 , issue=4 , year=2013 , doi=10.1639/0007-2745-116.4.350 , pages=350–357, s2cid=86826602 {{cite web , author=Source dataset.
Species Fungorum ''Index Fungorum'' is an international project to index all formal names (scientific names) in the fungus kingdom. the project is based at the Royal Botanic Gardens, Kew, one of three partners along with Landcare Research and the Institute of Mi ...
Plus: Species Fungorum for CoL+ , title=''Pyrenula'' , url=https://www.catalogueoflife.org/data/taxon/8HQD3 , work=
Catalog of Life The Catalogue of Life is an online database that provides an index of known species of animals, plants, fungi, and microorganisms. It was created in 2001 as a partnership between the global Species 2000 and the American Integrated Taxonomic Info ...
Version 2022-01-14 , access-date=1 March 2022
{{cite journal , last1=Aptroot , first1=André , last2=Common , first2=Ralph S. , title=''Pyrenula clavatispora'', a new species from Florida with narrowly clavate ascospores, with a key to similar species , journal=The Bryologist , volume=120 , issue=3 , year=2017 , doi=10.1639/0007-2745-120.3.270 , pages=270–273, s2cid=90103166 {{cite journal , last1=Diederich , first1=Paul , last2=Ertz , first2=Damien , title=First checklist of lichens and lichenicolous fungi from Mauritius, with phylogenetic analyses and description of new taxa , journal=Plant and Fungal Systematics , volume=65 , issue=1 , year=2020 , doi=10.35535/pfsyst-2020-0003 , pages=13–75 , doi-access=free {{cite book , last1=Eschweiler , first1=F.G. , year=1824 , title=Systema Lichenum, Genera Exhibens rite distincta, Pluribus Novis Adaucta , page=25 , language=la {{cite journal , last1=Etayo , first1=Javiar , last2=Aptroot , first2=André , title=''Pyrenula luteopruinosa'' sp. nov. from Panama and notes on other members of the genus , journal=The Lichenologist , volume=35 , issue=3 , year=2003 , doi=10.1016/s0024-2829(03)00021-5 , pages=233–236, s2cid=85092996 {{cite book , last1=Fée , first1=A.L.A. , year=1825 , title=Essai sur les cryptogames des écorces exotiques officinales , page=74 , location=Paris , publisher=Firmin Didot père et fils , language=fr , url=https://www.biodiversitylibrary.org/page/48999630 {{cite journal , last1=Galloway , first1=D.J. , year=2004 , title=New lichen taxa and names in the New Zealand mycobiota , journal=New Zealand Journal of Botany , volume=42 , pages=105–120, doi=10.1080/0028825X.2004.9512893 , s2cid=84174417 {{cite journal , last1=Gueidan , first1=Cécile , last2=Aptroot , first2=André , last3=Cáceres , first3=Marcela Eugenia da Silva , last4=Binh , first4=Nguyen Quoc , title=Molecular phylogeny of the tropical lichen family Pyrenulaceae: contribution from dried herbarium specimens and FTA card samples , journal=Mycological Progress , volume=15 , issue=1 , year=2015 , doi=10.1007/s11557-015-1154-8 , pages=1–21, s2cid=52807126 {{cite journal , last1=Harada , first1=H. , last2=Okamoto , first2=T. , last3=Yoshimura , first3=I. , year=2004 , title=A checklist of lichens and lichen-allies of Japan , journal=Lichenology , volume=2 , issue=2 , pages=47–166 {{cite journal , last1=Jagadeesh Ram , first1=T. 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M. , last2=Upreti , first2=D.K. , last3=Sinha , first3=G.P. , title=''Pyrenula subcylindrica'', a new pyrenocarpous lichen from India , journal=The Lichenologist , volume=37 , issue=2 , year=2005 , doi=10.1017/s0024282904014604 , pages=109–110, s2cid=85317085 {{cite journal , last1=Harris , first1=D.C. , year=1989 , title=A sketch of the family Pyrenulaceae (Melanommatales) in eastern North America , journal=Memoirs of the New York Botanical Garden , volume=49 , pages=74–107 {{cite journal , last1=Kalb , first1=K. , last2=Hafellner , first2=J. , year=1992 , title=Bemerkenswerte Flechten und lichenicole Pilze von der Insel Madeira , journal=Herzogia , volume=9 , issue=1–2 , pages=45–102 , language=de {{cite journal , last1=Lima , first1=Edvaneide Leandro de , last2=Mendonça , first2=Cléverton de Oliveira , last3=Maia , first3=Leonor Costa , last4=Aptroot , first4=André , last5=Cáceres , first5=Marcela Eugenia da Silva , title=Two new species of ''Pyrenula'' with a red or orange thallus from Vale do Catimbau National Park, Pernambuco, Brazil , journal=The Lichenologist , volume=45 , issue=2 , year=2013 , doi=10.1017/s0024282912000783 , pages=199–202, s2cid=86822681 {{cite journal , last1=Mendonça , first1=Cléverton De Oliveira , last2=Aptroot , first2=André , last3=Cáceres , first3=Marcela Eugenia Da Silva , title=Six new species of the lichen genus ''Pyrenula'' (Pyrenulaceae) from Northeast Brazil , journal=Phytotaxa , volume=286 , issue=3 , year=2016 , doi=10.11646/phytotaxa.286.3.4 , pages=169–176 {{cite journal , last1=Menezes , first1=Aline Anjos , last2=Xavier-Leite , first2=Amanda Barreto , last3=Aptroot , first3=André , last4=da Silva Cáceres , first4=Marcela Eugenia , title=New lichen species from the Caatinga in Chapada do Araripe, northeastern Brazil , journal=The Bryologist , volume=116 , issue=3 , year=2013 , doi=10.1639/0007-2745-116.3.302 , pages=302–305, s2cid=86053009 {{cite journal , last1=Moon , first1=K.H. , last2=Aptroot , first2=A. , year=2009 , title=Pyrenocarpous lichens in Korea , journal=Bibliotheca Lichenologica , volume=99 , pages=297–314 {{cite journal , last1=Müller , first1=J. , year=1882 , title=Lichenologische Beiträge XVI , journal=Flora (Regensburg) , volume=65 , issue=33 , pages=515–519 {{cite journal , last1=Müller , first1=J. , year=1885 , title=Lichenologische Beiträge XXI , journal=Flora (Regensburg) , volume=68 , issue=17 , pages=331–342 , language=la , url= {{cite journal , last1=Müller , first1=J. , year=1885 , title=Lichenologische Beiträge XXII , journal=Flora (Regensburg) , volume=68 , issue=28 , pages=503–518 , language=la {{cite journal , last1=Müller , first1=J. , year=1885 , title=Pyrenocarpeae Cubenses a cl. C. Wright lectae , journal=Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie, volume=6 , pages=375–421 12, language=la {{cite journal , last1=Müller , first1=J. , year=1887 , title=Lichenologische Beiträge XXVI. Flora (Regensburg) , journal=70 , issue=26/27 , pages=423–429 {{cite journal , last1=Müller , first1=J. , year=1891 , title=Lichenes Brisbanenses a cl. F.M. Bailey, Government Botanist, prope Brisbane (Queensland) in Australia orientali lecti , journal=Nuovo Giornale Botanico Italiano , volume=23 , issue=3 , pages=385–404 {{cite journal , last1=Müller , first1=J. , year=1891 , title=Lichenes Bellendenici a cl. 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Victoria lecti , journal=Bulletin de l'Herbier Boissier , volume=1 , pages=33–65 4, language=la {{cite journal , last1=Müller , first1=J. , year=1895 , title=Pyrenocarpeae Queenslandiae , journal=Report of the Australasian Association for the Advancement of Science , volume=6 , pages=449–466 , language=la {{cite journal , last1=Müller , first1=J. , year=1895 , title=Sertum Australiense s. species novae Australienses Thelotremearum, Graphidearum et Pyrenocarpearum , journal=Bulletin de l'Herbier Boissier , volume=3 , pages=313–327 , language=la {{cite journal , last1=Ram , first1=T.A.M. Jagadeesh , last2=Sinha , first2=G.P. , title=A new species and new records of ''Pyrenula'' (Pyrenulaceae) from India , journal=The Lichenologist , volume=42 , issue=1 , year=2009 , doi=10.1017/s0024282909990132 , pages=51–53, s2cid=85867687 {{cite journal , last1=Shirley , first1=J. , year=1894 , title=Notes on Tasmanian lichens , journal=Papers and Proceedings of the Royal Society of Tasmania , volume=1893 , pages=214–219 {{cite journal , last1=Shoemaker , first1=R.A. , last2=LeClair , first2=P.M. , year=1975 , title=Type studies of ''Massaria'' from the Wehmeyer collection , journal=Canadian Journal of Botany , volume=53 , issue=15 , pages=1568–1598, doi=10.1139/b75-188 {{cite journal , last1=Tucker , first1=Shirley C. , last2=Harris , first2=Richard C. , title=New and noteworthy pyrenocarpous lichens from Louisiana and Florida , journal=The Bryologist , volume=83 , issue=1 , year=1980 , doi=10.2307/3242389 , pages=1–20, jstor=3242389 {{cite journal , last1=Upreti , first1=D.K. , year=1998 , title=A key to the lichen genus ''Pyrenula'' from India, with nomenclatural notes , journal=Nova Hedwigia , volume=66 , issue=3–4 , pages=557–576 {{cite journal , last1=Weerakoon , first1=G. , last2=Aptroot , first2=A. , year=2016 , title=Nine new lichen species and 64 new records from Sri Lanka , journal=Phytotaxa , volume=280 , issue=2 , pages=152–162 , doi=10.11646/phytotaxa.280.2.5 {{cite journal , last1=Wijeyaratne , first1=S.C. , last2=Lücking , first2=R. , last3=Lumbsch , first3=H.T. , year=2012 , title=Three new crustose lichen species from Sri Lanka , journal=Nova Hedwigia , volume=94 , issue=3–4 , pages=367–372, doi=10.1127/0029-5035/2012/0008 * Pyrenula