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''Leptoptilos'' is a genus of very large tropical
stork Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family called Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as herons an ...
s, also known as the adjutant bird. The name means thin (''lepto'') feather (''ptilos''). Two species are resident breeders in southern
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
, and the
marabou stork The marabou stork (''Leptoptilos crumenifer'') is a large wading bird in the stork family Ciconiidae native to sub-Saharan Africa. It breeds in both wet and arid habitats, often near human habitation, especially landfill sites. It is someti ...
is found in
Sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
. These are huge
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s, typically 110–150 cm tall with a 210–250 cm wingspan. The three species each have a black upper body and wings, and white belly and undertail. The head and neck are bare like those of a
vulture A vulture is a bird of prey that scavenges on carrion. There are 23 extant species of vulture (including Condors). Old World vultures include 16 living species native to Europe, Africa, and Asia; New World vultures are restricted to North and ...
. The huge
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
is long and thick. Juveniles are a duller, browner version of the adult. ''Leptoptilos'' storks are gregarious colonial breeders in
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
s, building large stick
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
s in trees. They feed on
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s,
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s, young birds,
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s and
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s. They are frequent
scavenger Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...
s, and the naked head and neck are adaptations to this, as are those of the vultures with which they often feed. A feathered head would become rapidly clotted with
blood Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood in the c ...
and other substances when a scavenging bird's head was inside a large corpse, and the bare head is easier to keep clean. Most storks fly with neck outstretched, but the three ''Leptoptilos'' storks retract their necks in flight like a
heron The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychus ...
.


Taxonomy and species

The genus ''Leptoptilos'' was introduced in 1831 by the French naturalist
René Lesson René-Primevère Lesson (20 March 1794 – 28 April 1849) was a French surgeon, naturalist, ornithologist, and herpetologist. Biography Lesson was born at Rochefort, and entered the Naval Medical School in Rochefort at the age of sixteen. He ...
. The genus name combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''leptos'' meaning "delicate" or "slender" with ''ptilon'' meaning "feather". The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
was subsequently designated as the greater adjutant by
George Robert Gray George Robert Gray FRS (8 July 1808 – 6 May 1872) was an English zoologist and author, and head of the ornithological section of the British Museum, now the Natural History Museum, in London for forty-one years. He was the younger brother o ...
. The genus contains three extant species.


Fossils

There is an ample
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
of this genus. Many fossils members of the genus were much larger than living species, standing as tall as a man, with the earliest being '' Leptoptilos falconeri'' from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...
on the Southeast Asian islands of Java (''L''. ''titan'') and Flores ( ''L''. ''robustus''). *†'' Leptoptilos falconeri'' (Early to Late Pliocene of south Asia and east Africa) *†'' Leptoptilos indicus'' (Late Pliocene of
Siwalik The Sivalik Hills, also known as the Shivalik Hills and Churia Hills, are a mountain range of the outer Himalayas that stretches over about from the Indus River eastwards close to the Brahmaputra River, spanning the northern parts of the Indi ...
, India) – formerly ''Cryptociconia indica'', may be the same as ''L. falconeri'' (Louchart ''et al.'' 2005) *†''
Leptoptilos lüi ''Leptoptilos lüi'' is an extinct species of large-bodied Leptoptilini stork that existed during the Middle Pleistocene. Along with '' Aegypius jinniushanensis'', ''L. lüi'' is one of several new species of extinct birds discovered at Jinniush ...
'' (
Middle Pleistocene The Chibanian, widely known by its previous designation of Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. The ...
of
Jinniushan Jinniushan () is a Middle Pleistocene paleoanthropological site, dating to around 260,000 BP, most famous for its archaic hominin fossils. The site is located near Yingkou, Liaoning, China. Several new species of extinct birds were also discovere ...
, Liaoning, China) *†'' Leptoptilos patagonicus'' (Puerto Madryn
Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...
of
Valdés Peninsula The Valdes Peninsula (Spanish: ''Península Valdés'') is a peninsula into the Atlantic Ocean in the Biedma Department of north-east Chubut Province, Argentina. Around in size (not taking into account the isthmus of Carlos Ameghino which connect ...
, Argentina) *†'' Leptoptilos pliocenicus'' (Early Pliocene of Odessa, Ukraine and Urugus, Ethiopia to Late Pliocene of
Koro Toro Koro Toro is a settlement in southern Borkou-Ennedi-Tibesti Region in Chad. It hosts the Koro Toro Airport and a "notorious" maximum security desert prison used by the Chadian government to detain captured fighters of Boko Haram and Chadian rebel ...
, Chad and Olduvai, Tanzania) – includes ''L.'' cf. ''falconeri'', may be the same as ''L. falconeri'' *†'' Leptoptilos richae'' (Beglia Late Miocene of Bled ed Douarah, Tunisia, and Wadi Moghara, Egypt?) *†''
Leptoptilos robustus ''Leptoptilos robustus'' (from reek: thin, slender+ reek: soft featherand atin: strong is an extinct species of large-bodied stork belonging to the genus ''Leptoptilos'' that lived on the island of Flores in Indonesia during the Pleistocene ...
'' (Pleistocene,
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Including the Komodo Islands off its west coast (but excluding the Solor Archipelago to the east of Flores), the land area is 15,530.58 km2, and th ...
, Indonesia) *†'' Leptoptilos titan'' (Notopuro Middle/Late Pleistocene of Watualang, Java, Indonesia) *†''Leptoptilos'' sp. (
Ngorora Formation The Ngorora Formation is a geological formation in Kenya preserving fossils dating to the Miocene. The uppermost member of the formation shows sign of a faunal turnover that occurred around 11 to 10.5 million years ago, coinciding with faunal cha ...
Late Miocene of
Baringo District Baringo County is one of the 47 counties in Kenya. It is located in the former Rift Valley Province. Its headquarters and largest town is Kabarnet. The county is home to Lake Baringo. Geographical location Baringo bounded by Turkana County and ...
, Kenya: Louchart ''et al.'' 2005) †'' Leptoptilos siwalicensis'' from the
Siwalik The Sivalik Hills, also known as the Shivalik Hills and Churia Hills, are a mountain range of the outer Himalayas that stretches over about from the Indus River eastwards close to the Brahmaputra River, spanning the northern parts of the Indi ...
deposits (Late Miocene? to Late Pliocene) may belong to this genus or to a closely related one (Louchart ''et al.'' 2005).


In culture

The adjutant bird features in the arms of
Baron Sinha Baron Sinha, of Raipur in the Presidency of Bengal, is a title in the Peerage of the United Kingdom. It was created in 1919 for Sir Satyendra Prasanna Sinha, a distinguished barrister and ''zamindar'' who was the first (and only) Indian ever t ...
. In the satirical French puppet show '' Le Bébête Show'',
Valéry Giscard d'Estaing Valéry René Marie Georges Giscard d'Estaing (, , ; 2 February 19262 December 2020), also known as Giscard or VGE, was a French politician who served as President of France from 1974 to 1981. After serving as Minister of Finance under prime ...
was, for a time, depicted as a ''Leptoptilos''.


References

{{Taxonbar, from=Q577346 Bird genera Taxa described in 1831 Taxa named by René Lesson