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The lentil (''Lens culinaris'' or ''Lens esculenta'') is an edible
legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock f ...
. It is an
annual plant An annual plant is a plant that completes its life cycle, from germination to the production of seeds, within one growing season, and then dies. The length of growing seasons and period in which they take place vary according to geographical ...
known for its
lens A lens is a transmissive optical device which focuses or disperses a light beam by means of refraction. A simple lens consists of a single piece of transparent material, while a compound lens consists of several simple lenses (''elements''), ...
-shaped
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s. It is about tall, and the seeds grow in pods, usually with two seeds in each. As a food crop, the largest producer is Canada, producing 45% of the world’s total lentils. In
cuisines of the Indian subcontinent Cuisine of the Indian subcontinent includes the cuisines from the Indian subcontinent comprising the traditional cuisines from Bangladesh, Bhutan, India, the Maldives, Nepal, Pakistan and Sri Lanka. Staples and common ingredients Chapati, a ...
, where lentils are a staple, split lentils (often with their hulls removed) known as
dal In Indian cuisine, ''dal'' (also spelled ''daal'' or ''dhal''; pronunciation: , Hindi: दाल, Urdu: ) are dried, split pulses (e.g., lentils, peas, and beans) that do not require soaking before cooking. India is the largest producer of p ...
are often cooked into a thick
curry A curry is a dish with a sauce seasoned with spices, mainly associated with South Asian cuisine. In southern India, leaves from the curry tree may be included. There are many varieties of curry. The choice of spices for each dish in tradit ...
/gravy that is usually eaten with
rice Rice is the seed of the grass species ''Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima ''Oryza glaberrima'', commonly known as African rice, is one of the two domesticated rice species. It was first domesticated and grown i ...
or ''
roti Roti (also known as chapati) is a round flatbread native to the Indian subcontinent. It is popular in India, Sri Lanka, Pakistan, Nepal, Bangladesh, Maldives, Myanmar, Malaysia, Indonesia, Singapore, Thailand, Guyana, Suriname, Jamaica, Trinid ...
s''.


Botanical description


Name

Many different names in different parts of the world are used for the crop lentil. The first use of the word ''lens'' to designate a specific
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
was in the 16th century by the botanist Tournefort. The word "lens" for the lentil is of classical Roman/Latin origin: McGee points out that a prominent Roman family took the name " Lentulus", just as the family name "
Cicero Marcus Tullius Cicero ( ; ; 3 January 106 BC – 7 December 43 BC) was a Roman statesman, lawyer, scholar, philosopher, and academic skeptic, who tried to uphold optimate principles during the political crises that led to the estab ...
" was derived from the chickpea, ''
Cicer arietinum The chickpea or chick pea (''Cicer arietinum'') is an annual legume of the family Fabaceae, subfamily Faboideae. Its different types are variously known as gram" or Bengal gram, garbanzo or garbanzo bean, or Egyptian pea. Chickpea seeds are high ...
'', or " Fabia" (as in Quintus Fabius Maximus) from the fava bean (''
Vicia faba ''Vicia faba'', commonly known as the broad bean, fava bean, or faba bean, is a species of vetch, a flowering plant in the pea and bean family Fabaceae. It is widely cultivated as a crop for human consumption, and also as a cover crop. Variet ...
'').


Systematics

The genus ''Lens'' is part of the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Faboideae The Faboideae are a subfamily of the flowering plant family Fabaceae or Leguminosae. An acceptable alternative name for the subfamily is Papilionoideae, or Papilionaceae when this group of plants is treated as a family. This subfamily is wide ...
which is contained in the flowering
plant family Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of ...
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
or commonly known as
legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock f ...
or
bean A bean is the seed of several plants in the family Fabaceae, which are used as vegetables for human or animal food. They can be cooked in many different ways, including boiling, frying, and baking, and are used in many traditional dishes th ...
family, of the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Fabales The Fabales are an order (biology), order of flowering plants included in the Rosids, rosid group of the eudicots in the APG II system, Angiosperm Phylogeny Group II classification system. In the APG II circumscription, this order includes the f ...
in the
kingdom Kingdom commonly refers to: * A monarchy ruled by a king or queen * Kingdom (biology), a category in biological taxonomy Kingdom may also refer to: Arts and media Television * ''Kingdom'' (British TV series), a 2007 British television drama s ...
Plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
ae. ''Lens'' is a small genus which consists of the cultivated ''L. culinaris'' and six related wild taxa. Among the different taxa of wild lentils, ''L. orientalis'' is considered to be the progenitor of the cultivated lentil and is now generally classified as ''L. culinaris'' subsp. ''orientalis''. Lentil is
hypogeal Hypogeal, hypogean, hypogeic and hypogeous (; ) are biological terms describing an organism's activity below the soil surface. In botany, a seed is described as showing hypogeal germination when the cotyledons of the germinating seed remain n ...
, which means the
cotyledon A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The numb ...
s of the germinating seed stay in the ground and inside the seed coat. Therefore, it is less vulnerable to frost, wind erosion, or insect attack. The plant is a diploid, annual, bushy herb of erect, semierect, or spreading and compact growth and normally varies from in height. It has many hairy branches and its stem is slender and angular. The
rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this c ...
bears 10 to 15 leaflets in five to eight pairs. The leaves are alternate, of oblong-linear and obtuse shape and from yellowish green to dark bluish green in colour. In general, the upper leaves are converted into
tendrils In botany, a tendril is a specialized stem, leaf or petiole with a threadlike shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as ''Cuscuta''. There are many plants that have tendr ...
, whereas the lower leaves are mucronate. If
stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many speci ...
s are present, they are small. The flowers, one to four in number, are small, white, pink, purple, pale purple, or pale blue in colour. They arise from the axils of the leaves, on a slender footstalk almost as long as the leaves. The pods are oblong, slightly inflated, and about long. Normally, each of them contains two seeds, about in diameter, in the characteristic lens shape. The seeds can also be mottled and speckled. The several cultivated varieties of lentil differ in size, hairiness, and colour of the leaves, flowers, and seeds. Lentils are
self-pollinating Self-pollination is a form of pollination in which pollen from the same plant arrives at the stigma of a flower (in flowering plants) or at the ovule (in gymnosperms). There are two types of self-pollination: in autogamy, pollen is transferred to ...
. The flowering begins from the lowermost buds and gradually moves upward, so-called acropetal flowering. About two weeks are needed for all the flowers to open on the single branch. At the end of the second day and on the third day after the opening of the flowers, they close completely and the colour begins to fade. After three to four days, the setting of the pods takes place.


Types

Types can be classified according to their size, whether they are split or whole, or shelled or unshelled. Seed coats can range from light green to deep purple, as well as being tan, grey, brown, black or mottled. Shelled lentils show the colour of the cotyledon which can be yellow, orange, red, or green. Red-cotyledon types: * Nipper (Australia) * Northfield (Australia) * Cobber (Australia) * Digger (Australia) * Nugget (Australia) * Aldinga (Australia) * Masoor dal (unshelled lentils with a brown seed coat and an orange-red cotyledon) * Petite crimson (shelled Masoor lentils) * Red Chief (light tan seed coat and red cotyledon) Small green/brown-seed coat types: * Eston Green * Pardina (Spain) * Verdina (Spain) Medium green/brown-seed coat types * Avondale (United States) * Matilda (Australia) * Richlea Large green/brown-seed coat types: * Boomer (Australia) * Brewer's: a large brown lentil which is often considered the "regular" lentil in the United States * Castellana (Spanish) * Laird: the commercial standard for large green lentils in western Canada * Mason * Merrit * Mosa (Spain) * Naslada (Bulgaria) * Pennell (United States) * Riveland (United States) Other types: * Beluga: black, bead-like, lens-shaped, almost spherical, named for resemblance to
beluga caviar Beluga caviar is caviar consisting of the roe (or eggs) of the beluga sturgeon ''Huso huso''. The fish is found primarily in the Caspian Sea, which is bordered by Iran, Azerbaijan, Kazakhstan, Russia, and Turkmenistan. It can also be found in th ...
. Called Indianhead in Canada. * ''Macachiados'': big yellow Mexican lentils * Puy lentils (var. ''puyensis''): Small dark speckled blue-green lentil from France with a
Protected Designation of Origin The protected designation of origin (PDO) is a type of geographical indication of the European Union and the United Kingdom aimed at preserving the designations of origin of food-related products. The designation was created in 1992 and its main ...
name *
Alb-Leisa The term Alb-Leisa firstly means the ''Öko-Erzeugergemeinschaft Alb-Leisa'' (engl. "Eco-producer association Alb-Leisa"), secondly a trade name and thirdly traditional varieties of lentils from the Swabian Jura, Germany. "Leisa" means lentils in ...
three traditional genotypes of lentils native to the Swabian Jura (Alps) in Germany and protected by the producers' association Öko-Erzeugergemeinschaft Alb-Leisa (engl. "Eco-producer association Alb-Leisa")


Production

In 2020, global production of lentils was 6.5 million
tonne The tonne ( or ; symbol: t) is a unit of mass equal to 1000  kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton ( United State ...
s, led by Canada with 45% and India with 18% of the world total (table).
Saskatchewan Saskatchewan ( ; ) is a Provinces and territories of Canada, province in Western Canada, western Canada, bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and on t ...
is the most productive growing region in Canada, producing 95% of the national total. In India,
Madhya Pradesh Madhya Pradesh (, ; meaning 'central province') is a state in central India. Its capital is Bhopal, and the largest city is Indore, with Jabalpur, Ujjain, Gwalior, Sagar, and Rewa being the other major cities. Madhya Pradesh is the seco ...
and
Uttar Pradesh Uttar Pradesh (; , 'Northern Province') is a state in northern India. With over 200 million inhabitants, it is the most populated state in India as well as the most populous country subdivision in the world. It was established in 1950 ...
are largest producers, together producing more than 70% of the total. Other major producers include
West Bengal West Bengal (, Bengali: ''Poshchim Bongo'', , abbr. WB) is a state in the eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabitants within an area of . West Bengal is the fourt ...
and
Bihar Bihar (; ) is a state in eastern India. It is the 2nd largest state by population in 2019, 12th largest by area of , and 14th largest by GDP in 2021. Bihar borders Uttar Pradesh to its west, Nepal to the north, the northern part of West Be ...
.


Cultivation


History

The cultivated lentil ''Lens culinaris'' subsp. ''culinaris'' was derived from its wild subspecies ''L. culinaris'' subsp. ''orientalis'', although other species may also have contributed some genes, according to Jonathan Sauer (''Historical Geography of Crop Plants'', 2017.) Unlike their wild ancestors, domesticated lentil crops have
indehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...
pods and non-dormant seeds. Lentil was domesticated in the
Fertile Crescent The Fertile Crescent ( ar, الهلال الخصيب) is a crescent-shaped region in the Middle East, spanning modern-day Iraq, Syria, Lebanon, Israel, Palestine and Jordan, together with the northern region of Kuwait, southeastern region of ...
of the
Near East The ''Near East''; he, המזרח הקרוב; arc, ܕܢܚܐ ܩܪܒ; fa, خاور نزدیک, Xāvar-e nazdik; tr, Yakın Doğu is a geographical term which roughly encompasses a transcontinental region in Western Asia, that was once the hist ...
and then spread to
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
,
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
and the Indo-Gangetic plain. The primary center of diversity for the domestic ''Lens culinaris'' as well as its wild progenitor ''L. culinaris ssp. orientalis'' is considered to be the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
. The oldest known carbonized remains of lentil from
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
's
Franchthi Cave Franchthi Cave or Frankhthi Cave ( el, Σπήλαιον Φράγχθι) is an archaeological site overlooking Kiladha Bay, in the Argolic Gulf, opposite the village of Kiladha in southeastern Argolis, Greece. Humans first occupied the cave during ...
are dated to 11,000 BC. In archaeobotanical excavations carbonized remains of lentil seeds have been recovered from widely dispersed places such as
Tell Ramad Tell Ramad ( ar, تل رماد) is a prehistoric, Neolithic tell at the foot of Mount Hermon, about southwest of Damascus in Syria. The tell was the site of a small village of , which was first settled in the late 8th millennium BC. The tell was ...
in
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
(6250-5950 BC), Aceramic Beidha in
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
,
Hacilar Hacilar is an early human settlement in southwestern Turkey, 23 km south of present-day Burdur. It has been dated back 7040 BC at its earliest stage of development. Archaeological remains indicate that the site was abandoned and reoccupied ...
in
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
(5800-5000 BC), Tepe Sabz (Ita. Tepe Sabz) in
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
(5500-5000 BC) and Argissa-Magula Tessaly in Greece (6000-5000 BC), along other places.


Soil requirements

Lentils can grow on various soil types, from sand to clay loam, growing best in deep sandy loam soils with moderate fertility. A soil pH around 7 would be the best. Lentils do not tolerate flooding or water-logged conditions. Lentils improve the physical properties of soils and increase the yield of succeeding cereal crops.
Biological nitrogen fixation Nitrogen fixation is a chemical process by which molecular nitrogen (), with a strong triple covalent bond, in the air is converted into ammonia () or related nitrogenous compounds, typically in soil or aquatic systems but also in industry. Atmos ...
or other rotational effects could be the reason for higher yields after lentils''.''


Climate requirements

The conditions under which lentils are grown differ across different growing regions. In the
temperate climate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
s lentils are planted in the winter and spring under low temperatures and vegetative growth occurs in later spring and the summer. Rainfall during this time is not limited. In the
subtropics The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north and ...
, lentils are planted under relatively high temperatures at the end of the rainy season, and vegetative growth occurs on the residual soil moisture in the summer season. Rainfall during this time is limited. In
West Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...
and
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
, some lentils are planted as a winter crop before snowfall. Plant growth occurs during the time of snow melting. Under such cultivation, seed yields are often much higher.


Seedbed requirements and sowing

The lentil requires a firm, smooth
seedbed A seedbed or seedling bed is the local soil environment in which seeds are planted. Often it comprises not only the soil but also a specially prepared cold frame, hotbed or raised bed used to grow the seedlings in a controlled environment into l ...
with most of the previous
crop residue Crop residues are materials left in an agricultural field after the crop has been harvested. These residues include stalks and stubble (stems), leaves and seed pods. Good management of field residues can increase efficiency of irrigation and c ...
s incorporated. For the seed placement and for later harvesting it is important that the surface is not uneven with large clods, stones, or protruding crop residue. It is also important that the soil be made friable and weed-free, so that seeding can be done at a uniform depth. The plant densities for lentils vary between
genotype The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
s, seed size, planting time and growing conditions, and also from region to region. In South Asia, a seed rate of is recommended. In West Asian countries, a higher seed rate is recommended, and also leads to a higher yield. The seeds should be sown deep. In agriculturally mechanized countries, lentils are planted using grain drills, but many other areas still hand broadcast.


Cultivation management, fertilization

In intercropping systems – a practice commonly used in lentil cultivation –
herbicide Herbicides (, ), also commonly known as weedkillers, are substances used to control undesired plants, also known as weeds.EPA. February 201Pesticides Industry. Sales and Usage 2006 and 2007: Market Estimates. Summary in press releasMain page fo ...
s may be needed to assure crop health. Like many other
legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock f ...
crops, lentils can fix atmospheric nitrogen in the soil with specific
rhizobia Rhizobia are diazotrophic bacteria that fix nitrogen after becoming established inside the root nodules of legumes (Fabaceae). To express genes for nitrogen fixation, rhizobia require a plant host; they cannot independently fix nitrogen. In gene ...
. Lentils grow well under low
fertilizer A fertilizer (American English) or fertiliser (British English; see spelling differences) is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from ...
input conditions, although
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
,
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
,
potassium Potassium is the chemical element with the symbol K (from Neo-Latin ''kalium'') and atomic number19. Potassium is a silvery-white metal that is soft enough to be cut with a knife with little force. Potassium metal reacts rapidly with atmosphe ...
, and
sulfur Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...
may be used for
nutrient A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
-poor soils.


Diseases

Below is a list of the most common lentil diseases.


Fungal diseases


Nematodes, parasitic


Viral diseases


Use by humans


Processing

A combination of gravity, screens and air flow is used to clean and sort lentils by shape and density. After destoning, they may be sorted by a color sorter and then packaged. A major part of the world's red lentil production undergoes a secondary processing step. These lentils are dehulled, split and polished. In the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
, this process is called dal milling. The moisture content of the lentils prior dehulling is crucial to guarantee a good dehulling efficiency. The hull of lentils usually accounts for 6 to 7 percent of the total seed weight, which is lower than most legumes. Lentil flour can be produced by milling the seeds, like cereals.


Culinary use

Lentils can be eaten soaked, germinated, fried, baked or boiled – the most common preparation method. The seeds require a cooking time of 10 to 40 minutes, depending on the variety; small varieties with the husk removed, such as the common red lentil, require shorter cooking times (and unlike most legumes don't require soaking). Most varieties have a distinctive, earthy flavor. Lentils with husks remain whole with moderate cooking, while those without husks tend to disintegrate into a thick purée, which may enable various dishes. The composition of lentils leads to a high
emulsifying An emulsion is a mixture of two or more liquids that are normally immiscible (unmixable or unblendable) owing to liquid-liquid phase separation. Emulsions are part of a more general class of two-phase systems of matter called colloids. Although ...
capacity which can be even increased by dough
fermentation Fermentation is a metabolic process that produces chemical changes in organic substrates through the action of enzymes. In biochemistry, it is narrowly defined as the extraction of energy from carbohydrates in the absence of oxygen. In food ...
in bread making.


Lentil dishes

Lentils are used worldwide in many different dishes. Lentil dishes are most widespread throughout
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...
, the
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...
regions,
West Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...
, and
Latin America Latin America or * french: Amérique Latine, link=no * ht, Amerik Latin, link=no * pt, América Latina, link=no, name=a, sometimes referred to as LatAm is a large cultural region in the Americas where Romance languages — languages derived f ...
. In the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
,
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
,
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It incl ...
,
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...
and the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
, lentil curry is part of the everyday diet, eaten with both rice and
roti Roti (also known as chapati) is a round flatbread native to the Indian subcontinent. It is popular in India, Sri Lanka, Pakistan, Nepal, Bangladesh, Maldives, Myanmar, Malaysia, Indonesia, Singapore, Thailand, Guyana, Suriname, Jamaica, Trinid ...
. Boiled lentils and lentil stock are used to thicken most vegetarian curries. They are also used as stuffing in
dal In Indian cuisine, ''dal'' (also spelled ''daal'' or ''dhal''; pronunciation: , Hindi: दाल, Urdu: ) are dried, split pulses (e.g., lentils, peas, and beans) that do not require soaking before cooking. India is the largest producer of p ...
paratha Paratha () is a flatbread native to South Asia, prevalent throughout the modern-day nations of India, Sri Lanka, Pakistan, Nepal, Bangladesh, Maldives, Afghanistan, Myanmar, Malaysia, Singapore, Mauritius, Fiji, Guyana, Suriname, and Trinidad a ...
s and
puri Puri () is a coastal city and a Nagar Palika, municipality in the state of Odisha in eastern India. It is the district headquarters of Puri district and is situated on the Bay of Bengal, south of the state capital of Bhubaneswar. It is als ...
for breakfast or snacks. Lentils are also used in many regional varieties of sweets. Lentil flour is used to prepare several different bread varieties, such as
papadam A papad is an Indian deep fried dough of black gram bean flour, either fried or cooked with dry heat (flipped over an open flame) until crunchy. Other flours made from lentils, chickpeas, rice, tapioca, millet or potato are also used. ''Papad' ...
. They are frequently combined with
rice Rice is the seed of the grass species ''Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima ''Oryza glaberrima'', commonly known as African rice, is one of the two domesticated rice species. It was first domesticated and grown i ...
, which has a similar cooking time. A lentil and rice dish is referred to in Levantine countries as ''
mujaddara Mujaddara ( ar, مجدرة ''mujadarah'', with alternative spellings in English majadra, mejadra, moujadara, mudardara, and megadarra) consists of cooked lentils together with groats, generally rice, and garnished with sautéed onions. Name and o ...
'' or ''mejadra''. In Iran, rice and lentil is served with fried
raisin A raisin is a dried grape. Raisins are produced in many regions of the world and may be eaten raw or used in cooking, baking, and brewing. In the United Kingdom, Ireland, New Zealand, and Australia, the word ''raisin'' is reserved for the d ...
; this dish is called ''adas polo''. Rice and lentils are also cooked together in ''
khichdi ''Khichdi'' or ''khichri'' (, , , , Odia: ଖେଚୁଡି) is a dish in South Asian cuisine made of rice and lentils (''dal'') with numerous variations. Variations include ''bajra'' and mung ''dal'' ''khichri''. In Indian culture, in ...
'', a popular dish in the Indian subcontinent (India and Pakistan); a similar dish, ''
kushari Koshary, kushari or koshari ( arz, كشري ) is Egypt's national dish and a widely popular street food. A traditional Egyptian cuisine, Egyptian staple, mixing pasta, rice and brown lentils, and topped with a zesty tomato sauce, garlic vinegar ...
'', made in Egypt, is considered one of two
national dish A national dish is a culinary dish that is strongly associated with a particular country. A dish can be considered a national dish for a variety of reasons: * It is a staple food, made from a selection of locally available foodstuffs that can be ...
es. Lentils are used to prepare an inexpensive and nutritious
soup Soup is a primarily liquid food, generally served warm or hot (but may be cool or cold), that is made by combining ingredients of meat or vegetables with stock, milk, or water. Hot soups are additionally characterized by boiling solid ing ...
throughout Europe and
North and South America The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
, sometimes combined with
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adult m ...
or
pork Pork is the culinary name for the meat of the domestic pig (''Sus domesticus''). It is the most commonly consumed meat worldwide, with evidence of pig husbandry dating back to 5000 BCE. Pork is eaten both freshly cooked and preserved; ...
. In Western countries, cooked lentils are often used in salads. In Italy, the traditional dish for New Year's Eve is
Cotechino The ''cotechino'' (, ) is an Italian large pork sausage requiring slow cooking; usually it is simmered at low heat for several hours. Its name comes from ''cotica'' (rind), but it may take different names depending on its various locations of pro ...
served with lentils. Lentils are commonly eaten in
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
in a stew-like dish called ''kik'', or ''kik wot'', one of the dishes people eat with Ethiopia's national food, ''
injera Injera (, ; om, Biddeena; ) is a sour fermented pancake-like flatbread with a slightly spongy texture, traditionally made of teff flour. In Ethiopia, Eritrea, and some parts of Sudan, injera is the staple. Injera is central to the dining proc ...
'' flatbread. Yellow lentils are used to make a non-spicy stew, which is one of the first solid foods Ethiopians feed their babies. Lentils were a chief part of the diet of ancient Iranians, who consumed lentils daily in the form of a stew poured over rice.


Nutritional value


Composition

According to the
USDA National Nutrient Database The USDA National Nutrient Database for Standard Reference is a database produced by the United States Department of Agriculture that provides the nutritional content of many generic and proprietary-branded foods. Released in August 2015 and revised ...
, of raw lentils (variety unspecified) provide of
food energy Food energy is chemical energy that animals (including humans) derive from their food to sustain their metabolism, including their muscle, muscular activity. Most animals derive most of their energy from aerobic respiration, namely combining the ...
; the same weight of cooked lentils provides . Raw lentils are 8% water, 63%
carbohydrates In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or may ...
including 11%
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
, 25%
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
, and 1%
fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers spec ...
(table). Cooked lentils (when boiled) are a rich source (20% or more of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of health ...
, DV) of numerous
essential nutrients A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
, including
folate Folate, also known as vitamin B9 and folacin, is one of the B vitamins. Manufactured folic acid, which is converted into folate by the body, is used as a dietary supplement and in food fortification as it is more stable during processing and ...
(45% DV), iron (25% DV), manganese (24% DV), and phosphorus (26% DV). They are a good source (10% or more of the Daily Value) of several nutrients including
thiamine Thiamine, also known as thiamin and vitamin B1, is a vitamin, an essential micronutrient, that cannot be made in the body. It is found in food and commercially synthesized to be a dietary supplement or medication. Phosphorylated forms of thi ...
(15% DV),
pantothenic acid Pantothenic acid, also called vitamin B5 is a water-soluble B vitamin and therefore an essential nutrient. All animals require pantothenic acid in order to synthesize coenzyme A (CoA) – essential for fatty acid metabolism – as well as to, i ...
(13% DV), vitamin B6 (14% DV),
magnesium Magnesium is a chemical element with the symbol Mg and atomic number 12. It is a shiny gray metal having a low density, low melting point and high chemical reactivity. Like the other alkaline earth metals (group 2 of the periodic ta ...
(10% DV),
copper Copper is a chemical element with the symbol Cu (from la, cuprum) and atomic number 29. It is a soft, malleable, and ductile metal with very high thermal and electrical conductivity. A freshly exposed surface of pure copper has a pinkis ...
(13% DV), and
zinc Zinc is a chemical element with the symbol Zn and atomic number 30. Zinc is a slightly brittle metal at room temperature and has a shiny-greyish appearance when oxidation is removed. It is the first element in group 12 (IIB) of the periodi ...
(13%) (see table). When lentils are cooked by boiling, protein content declines to 9% of total composition, and B vitamins and minerals decrease due to the overall water content increasing (protein itself is not lost). Lentils have the second-highest ratio of protein to food energy of any legume, after
soybean The soybean, soy bean, or soya bean (''Glycine max'') is a species of legume native to East Asia, widely grown for its edible bean, which has numerous uses. Traditional unfermented food uses of soybeans include soy milk, from which tofu an ...
s. Lentils contain the
carotenoid Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic compound, organic pigments that are produced by plants and algae, as well as several bacteria, and Fungus, fungi. Carotenoids give the characteristic color to pumpki ...
s
lutein Lutein (;"Lutein"
zeaxanthin Zeaxanthin is one of the most common carotenoids in nature, and is used in the xanthophyll cycle. Synthesized in plants and some micro-organisms, it is the pigment that gives paprika (made from bell peppers), corn, saffron, goji ( wolfberries), ...
, as well as
polyunsaturated fatty acid Polyunsaturated fatty acids (PUFAs) are fatty acids that contain more than one double bond in their backbone. This class includes many important compounds, such as essential fatty acids and those that give drying oils their characteristic proper ...
s.


Digestive effects

The low levels of readily digestible starch (5 percent) and high levels of slowly digested starch make lentils of potential value to people with
diabetes Diabetes, also known as diabetes mellitus, is a group of metabolic disorders characterized by a high blood sugar level ( hyperglycemia) over a prolonged period of time. Symptoms often include frequent urination, increased thirst and increased ap ...
. The remaining 65% of the starch is a
resistant starch Resistant starch (RS) is starch, including its degradation products, that escapes from digestion in the small intestine of healthy individuals. Resistant starch occurs naturally in foods, but it can also be added as part of dried raw foods, or use ...
classified as RS1. A minimum of 10% in starch from lentils escapes
digestion Digestion is the breakdown of large insoluble food molecules into small water-soluble food molecules so that they can be absorbed into the watery blood plasma. In certain organisms, these smaller substances are absorbed through the small intest ...
and absorption in the
small intestine The small intestine or small bowel is an organ in the gastrointestinal tract where most of the absorption of nutrients from food takes place. It lies between the stomach and large intestine, and receives bile and pancreatic juice through the p ...
(therefore called "resistant starch"). Additional resistant starch is synthesized from gelatinized starch, during cooling, after lentils are cooked. Lentils also have
antinutrient Antinutrients are natural or synthetic compounds that interfere with the absorption of nutrients. Nutrition studies focus on antinutrients commonly found in food sources and beverages. Antinutrients may take the form of drugs, chemicals that natura ...
factors, such as
trypsin inhibitor A trypsin inhibitor (TI) is a protein and a type of serine protease inhibitor (serpin) that reduces the biological activity of trypsin by controlling the activation and catalytic reactions of proteins. Trypsin is an enzyme involved in the breakdown ...
s and a relatively high
phytate Phytic acid is a six-fold dihydrogenphosphate ester of inositol (specifically, of the ''myo'' isomer), also called inositol hexakisphosphate (IP6) or inositol polyphosphate. At physiological pH, the phosphates are partially ionized, resulting in ...
content. Trypsin is an enzyme involved in digestion, and phytates reduce the
bioavailability In pharmacology, bioavailability is a subcategory of absorption and is the fraction (%) of an administered drug that reaches the systemic circulation. By definition, when a medication is administered intravenously, its bioavailability is 100%. H ...
of
dietary minerals In the context of nutrition, a mineral is a chemical element required as an essential nutrient by organisms to perform functions necessary for life. However, the four major structural elements in the human body by weight (oxygen, hydrogen, carbon, ...
. The phytates can be reduced by prolonged soaking and fermentation or sprouting. Cooking nearly completely removes the trypsin inhibitor activity; sprouting is also effective.


Breeding

Although lentils have been an important crop for centuries, lentil breeding and genetic research have a relatively short history compared to that of many other crops. Since the inception of The International Center for Agriculture Research in the Dry Areas (
ICARDA The International Center for Agricultural Research in the Dry Areas (ICARDA), a member of CGIAR, supported by the CGIAR Fund, is a non-profit agricultural research institute that aims to improve the livelihoods of the resource-poor across the ...
) breeding programme in 1977 significant gains have been made. It supplies landraces and breeding lines for countries around the world, supplemented by other programmes in both developing (e.g. India) and developed (e.g. Australia and Canada) countries. In recent years, such collaborations among breeders and agronomists are becoming increasingly important. The focus lies on high yielding and stable cultivars for diverse environments to match the demand of a growing population. In particular, progress in quantity and quality as well as in the resistance to disease and abiotic stresses are the major breeding aims. Several varieties have been developed applyin
conventional breeding methodologies
Serious genetic improvement for yield has been made, however, the full potential of production and productivity could not yet be tapped due to several biotic and abiotic stresses. Wild ''Lens'' species are a significant source of genetic variation for improving the relatively narrow genetic base of this crop. The wild species possess many diverse traits including disease resistances and abiotic stress tolerances. The above-mentioned ''L. nigricans'' and ''L. orientalis'' possess morphological similarities to the cultivated ''L. culinaris''. But only ''L. culinaris'' and ''L. culinaris'' subsp. orientalis are crossable and produce fully fertile seed. Between the different related species hybridisation barriers exist. According to their inter-crossability ''Lens'' species can be divided into three gene pools: # Primary gene pool'':'' ''L. culinaris'' (and ''L. culinaris'' subsp''. orientalis'') and ''L. odemensis'' # Secondary gene pool: ''L. ervoides'' and ''L. nigricans'' # Tertiary gene pool: ''L. lamottei'' and ''L. tomentosus'' Crosses generally fail between members of different gene pools. However,
plant growth regulators Plant hormone (or phytohormones) are signal molecules, produced within plants, that occur in extremely low concentrations. Plant hormones control all aspects of plant growth and development, from embryogenesis, the regulation of organ size, pa ...
and/or
embryo rescue Embryo rescue is one of the earliest and successful forms of in-vitro culture techniques that is used to assist in the development of plant embryos that might not survive to become viable plants. Embryo rescue plays an important role in modern plan ...
allows the growth of viable hybrids between groups. Even if crosses are successful, many undesired genes may be introduced as well in addition to the desired ones. This can be resolved by using a
backcrossing Backcrossing is a crossing of a hybrid with one of its parents or an individual genetically similar to its parent, to achieve offspring with a genetic identity closer to that of the parent. It is used in horticulture, animal breeding, and product ...
programme. Thus,
mutagenesis Mutagenesis () is a process by which the genetic information of an organism is changed by the production of a mutation. It may occur spontaneously in nature, or as a result of exposure to mutagens. It can also be achieved experimentally using la ...
is crucial to create new and desirable varieties. According to Yadav et al. other biotechnology techniques which may impact on lentil breeding are micro-propagation using meristamatic explants, callus culture and regeneration, protoplast culture and doubled haploid production. There is a proposed revision of the gene pools using SNP phylogeny.


See also

*
Dal In Indian cuisine, ''dal'' (also spelled ''daal'' or ''dhal''; pronunciation: , Hindi: दाल, Urdu: ) are dried, split pulses (e.g., lentils, peas, and beans) that do not require soaking before cooking. India is the largest producer of p ...
*
Lentil soup Lentil soup is a soup with lentils as its main ingredient; it may be vegetarian or include meat, and may use brown, red, yellow, green or black lentils, with or without the husk. Dehulled yellow and red lentils disintegrate in cooking, making a ...
* Lentils with Swabian pasta *
Mujaddara Mujaddara ( ar, مجدرة ''mujadarah'', with alternative spellings in English majadra, mejadra, moujadara, mudardara, and megadarra) consists of cooked lentils together with groats, generally rice, and garnished with sautéed onions. Name and o ...
* National Lentil Festival


References


Further reading

* Alan Davidson, ''The Oxford Companion to Food''. * S S Yadav et al. ''Lentil: An Ancient Crop for Modern Times''. (2007). Springer Verlag. .


External links


Lentils – Country Production, Consumption, Exports, and Imports Statistics


{{Taxonbar, from1=Q131226, from2=Q111391147 Pulse crop diseases Fabeae Edible legumes Founder crops