The Lamiaceae ( ) or Labiatae are a

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s commonly known as the mint, deadnettle or sage family. Many of the plants are

aroma An odor (American English) or odour (Commonwealth English; see spelling differences) is caused by one or more volatilized chemical compounds that are generally found in low concentrations that humans and animals can perceive via their sense ...

tic in all parts and include widely used culinary

herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...

s like

basil Basil (, ; ''Ocimum basilicum'' , also called great basil, is a culinary herb of the family Lamiaceae (mints). It is a tender plant, and is used in cuisines worldwide. In Western cuisine, the generic term "basil" refers to the variety also kno ...

mint MiNT is Now TOS (MiNT) is a free software alternative operating system kernel for the Atari ST system and its successors. It is a multi-tasking alternative to TOS and MagiC. Together with the free system components fVDI device drivers, XaAES g ...

rosemary ''Salvia rosmarinus'' (), commonly known as rosemary, is a shrub with fragrant, evergreen, needle-like leaves and white, pink, purple, or blue flowers, native plant, native to the Mediterranean Region, Mediterranean region. Until 2017, it was kn ...

sage Sage or SAGE may refer to: Plants * ''Salvia officinalis'', common sage, a small evergreen subshrub used as a culinary herb ** Lamiaceae, a family of flowering plants commonly known as the mint or deadnettle or sage family ** ''Salvia'', a large ...

savory Savory or Savoury may refer to: Common usage * Herbs of the genus ''Satureja'', particularly: ** Summer savory (''Satureja hortensis''), an annual herb, used to flavor food ** Winter savory (''Satureja montana''), a perennial herb, also used to ...

marjoram Marjoram (; ''Origanum majorana'') is a cold-sensitive perennial herb or undershrub with sweet pine and citrus flavours. In some Middle Eastern countries, marjoram is synonymous with oregano, and there the names sweet marjoram and knotted mar ...

oregano Oregano (, ; ''Origanum vulgare'') is a species of flowering plant in the mint family Lamiaceae. It was native to the Mediterranean region, but widely naturalised elsewhere in the temperate Northern Hemisphere. Oregano is a woody perennial pla ...

hyssop ''Hyssopus officinalis'' or hyssop is a shrub in the Lamiaceae or mint family native to Southern Europe, the Middle East, and the region surrounding the Caspian Sea. Due to its purported properties as an antiseptic, cough reliever, and expecto ...

thyme Thyme () is the herb (dried aerial parts) of some members of the genus ''Thymus'' of aromatic perennial evergreen herbs in the mint family Lamiaceae. Thymes are relatives of the oregano genus ''Origanum'', with both plants being mostly indigenou ...

lavender ''Lavandula'' (common name lavender) is a genus of 47 known species of flowering plants in the mint family, Lamiaceae. It is native to the Old World and is found in Cape Verde and the Canary Islands, and from Europe across to northern and easte ...

, and

perilla ''Perilla'' is a genus consisting of one major Asiatic crop species ''Perilla frutescens'' and a few wild species in nature belonging to the mint family, Lamiaceae. The genus encompasses several distinct varieties of Asian herb, seed, and vegeta ...

, as well as other medicinal herbs such as

catnip ''Nepeta cataria'', commonly known as catnip, catswort, catwort, and catmint, is a species of the genus ''Nepeta'' in the family Lamiaceae, native to southern and eastern Europe, the Middle East, Central Asia, and parts of China. It is widely n ...

salvia ''Salvia'' () is the largest genus of plants in the sage family Lamiaceae, with nearly 1000 species of shrubs, herbaceous plant, herbaceous perennial plant, perennials, and annual plant, annuals. Within the Lamiaceae, ''Salvia'' is part of the ...

bee balm ''Monarda'' is a genus of flowering plants in the mint family, Lamiaceae.Harley, R. M., et al. 2004. "Labiatae". pp 167-275 In: Kubitzki, K. (editor) and J. W. Kadereit (volume editor). ''The Families and Genera of Vascular Plants'' volume VII. ...

, wild dagga, and oriental motherwort. Some species are

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s (such as

teak Teak (''Tectona grandis'') is a tropical hardwood tree species in the family Lamiaceae. It is a large, deciduous tree that occurs in mixed hardwood forests. ''Tectona grandis'' has small, fragrant white flowers arranged in dense clusters (panicl ...

), or, rarely,

vine A vine (Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners themselv ...

s. Many members of the family are widely cultivated, not only for their aromatic qualities, but also their ease of cultivation, since they are readily propagated by stem cuttings. Besides those grown for their edible leaves, some are grown for decorative foliage. Others are grown for seed, such as ''

Salvia hispanica ''Salvia hispanica'', commonly known as chia (), is a species of flowering plant in the mint family, Lamiaceae, native to central and southern Mexico and Guatemala. It is considered a pseudocereal, cultivated for its edible, hydrophilic chia s ...

'' (chia), or for their edible tubers, such as ''

Plectranthus edulis ''Coleus maculosus'' subsp. ''edulis'', synonym ''Plectranthus edulis'', the Ethiopian potato, known as ''welayta dinich'' in Amharic, is a species of perennial plant in the family Lamiaceae. It is indigenous to Ethiopia Ethiopia, , om, ...

'', ''

Plectranthus esculentus ''Coleus esculentus'', synonym ''Plectranthus esculentus'', also known as the kaffir potato or Livingstone potato, is a species of plant in the dicot family Lamiaceae. It is indigenous to Africa, where it is grown for its edible tubers. It is m ...

'', ''

Plectranthus rotundifolius ''Coleus rotundifolius'', synonyms ''Plectranthus rotundifolius'' and ''Solenostemon rotundifolius'', commonly known as native or country potato in Africa and called Chinese potato in India, is a perennial herbaceous plant of the mint family ...

'', and ''

Stachys affinis ''Stachys affinis'', commonly called crosne, Chinese artichoke, Japanese artichoke, knotroot, or artichoke betony, is a perennial herbaceous plant of the family Lamiaceae, originating from China. Its rhizome is a root vegetable that can be e ...

'' (Chinese artichoke). Many are also grown ornamentally, notably

coleus ''Coleus'' is a genus of annual or perennial herbs or shrubs, sometimes succulent, sometimes with a fleshy or tuberous rootstock, found in the Old World tropics and subtropics. The relationship among the genera ''Coleus'', '' Solenostemon'' and ...

, ''

Plectranthus ''Plectranthus'' is a genus of about 85 species of flowering plants from the sage family, Lamiaceae, found mostly in southern and tropical Africa and Madagascar. Common names include spur-flower. ''Plectranthus'' species are herbaceous perennia ...

'', and many ''

Salvia ''Salvia'' () is the largest genus of plants in the sage family Lamiaceae, with nearly 1000 species of shrubs, herbaceous plant, herbaceous perennial plant, perennials, and annual plant, annuals. Within the Lamiaceae, ''Salvia'' is part of the ...

'' species and hybrids. The family has a

cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of the world in appropriate habitats. Such a taxon, usually a species, is said to exhibit cosmopolitanism or cosmopolitism. The ext ...

. The enlarged Lamiaceae contain about 236

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...

and have been stated to contain 6,900 to 7,200

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

, but the World Checklist lists 7,534. The largest genera are ''Salvia'' (900), ''Scutellaria'' (360), ''Stachys'' (300), ''Plectranthus'' (300), ''Hyptis'' (280), ''Teucrium'' (250), ''Vitex'' (250), ''Thymus'' (220), and ''Nepeta'' (200). ''Clerodendrum'' was once a genus of over 400 species, but by 2010, it had been narrowed to about 150. The family has traditionally been considered closely related to the

Verbenaceae The Verbenaceae ( ), the verbena family or vervain family, is a family of mainly tropical flowering plants. It contains trees, shrubs, and herbs notable for heads, spikes, or clusters of small flowers, many of which have an aromatic smell. The ...

;Raymond M. Harley, Sandy Atkins, Andrey L. Budantsev, Philip D. Cantino, Barry J. Conn, Renée J. Grayer, Madeline M. Harley, Rogier P.J. de Kok, Tatyana V. Krestovskaja, Ramón Morales, Alan J. Paton, and P. Olof Ryding. 2004. "Labiatae" pages 167-275. In: Klaus Kubitzki (editor) and Joachim W. Kadereit (volume editor). ''The Families and Genera of Vascular Plants'' volume VII. Springer-Verlag: Berlin; Heidelberg, Germany. in the 1990s,

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

studies Study or studies may refer to: General * Education **Higher education * Clinical trial * Experiment * Observational study * Research * Study skills, abilities and approaches applied to learning Other * Study (art), a drawing or series of drawin ...

suggested that many

classified in the Verbenaceae should be classified in the Lamiaceae Cantino, P.D., Harley, R.M. & Wagstaff, S.J. 1992. Genera of Labiatae: status and classification. Pp. 511-522. In: Raymond M. Harley and Tom Reynolds (editors). ''Advances in Labiate Science.'' Richmond, Royal Botanic Gardens, Kew. or to other families in the

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

Lamiales The order Lamiales (also known as the mint order) are an order in the asterid group of dicotyledonous flowering plants. It includes about 23,810 species, 1,059 genera, and is divided into about 25 families. These families include Acanthaceae, Bi ...

The alternative family name Labiatae refers to the flowers typically having petals fused into an upper lip and a lower lip (' in Latin). The

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

s are bilaterally symmetrical with five united

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

s and five united

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

s. They are usually

bisexual Bisexuality is a romantic or sexual attraction or behavior toward both males and females, or to more than one gender. It may also be defined to include romantic or sexual attraction to people regardless of their sex or gender identity, whi ...

and verticillastrate (a flower cluster that looks like a whorl of flowers, but actually consists of two crowded clusters). Although this is still considered an acceptable alternative name, most botanists now use the name Lamiaceae in referring to this family. The

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

emerge oppositely, each pair at right angles to the previous one (decussate) or whorled. The stems are frequently square in cross section, but this is not found in all members of the family, and is sometimes found in other plant families.

Genera

Leucas aspera at Gandipet, Hyderabad, AP W IMG 9054

Orthosiphon pallidus (Jyoti) in Talakona forest, AP W IMG 8284

The last revision of the entire family was published in 2004. It described and provided

keys Key or The Key may refer to: Common meanings * Key (cryptography), a piece of information that controls the operation of a cryptography algorithm * Key (lock), device used to control access to places or facilities restricted by a lock * Key (map ...

to 236 genera. These are marked with an asterisk (*) in the list below. A few genera have been established or resurrected since 2004. These are marked with a plus sign (+). Other genera have been synonymised. These are marked with a minus sign (-). The remaining genera in the list are mostly of historical interest only and are from a source that includes such genera without explanation."List of genera in Lamiaceae". In: "Lamiaceae". In: "List of families". In: "Families and genera in GRIN. (see ''External links'' below) Few of these are recognized in modern treatments of the family.

Kew Gardens Kew Gardens is a botanical garden, botanic garden in southwest London that houses the "largest and most diverse botany, botanical and mycology, mycological collections in the world". Founded in 1840, from the exotic garden at Kew Park, its li ...

provides a list of genera that includes additional information.List of Genera in Lamiaceae. At: Vascular Plant Families and Genera. At: World Checklist of Selected Plant Families. At: Electronic Plant Information Center. At: Website of ''Royal Botanic Gardens, Kew''. (see ''External Links'' below). A list at the

Angiosperm Phylogeny Website The Angiosperm Phylogeny Website (or APweb) is a website dedicated to research on angiosperm phylogeny and taxonomy. The site is hosted by the Missouri Botanical Garden website and maintained by researchers, Peter F. Stevens and Hilary M. Davis ...

is frequently updated.

Recent changes

The

circumscription Circumscription may refer to: *Circumscribed circle *Circumscription (logic) *Circumscription (taxonomy) * Circumscription theory, a theory about the origins of the political state in the history of human evolution proposed by the American anthrop ...

of several genera has changed since 2004. ''Tsoongia'', ''Paravitex'', and ''Viticipremna'' have been sunk into

synonymy A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

with ''Vitex''. ''Huxleya'' has been sunk into ''Volkameria''. ''Kalaharia'', ''Volkameria'', ''Ovieda'', and ''Tetraclea'' have been segregated from a formerly

polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...

''Clerodendrum''. ''Rydingia'' has been separated from ''Leucas''. The remaining ''Leucas'' is

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

over four other genera.

Subfamilies and tribes

In 2004, the Lamiaceae were divided into seven

subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

, plus 10 genera not placed in any of the subfamilies. The unplaced genera are: ''

Tectona ''Tectona'' is a genus of tropical hardwood trees in the mint family, ''Lamiaceae''.Heywood, V.H., Brummitt, R.K., Culham, A. & Seberg, O. 2007: Flowering Plant Families of the World. Royal Botanic Gardens, Kew. The three species are often collec ...

'', ''

Callicarpa ''Callicarpa'' (beautyberry) is a genus of shrubs and small trees in the family Lamiaceae.Heywood, V.H., Brummitt, R.K., Culham, A. & Seberg, O. 2007: Flowering Plant Families of the World. Royal Botanic Gardens, Kew. They are native to east and ...

'', ''

Hymenopyramis ''Hymenopyramis'' is a genus of plants in the family Lamiaceae, first described in 1843. It is native to Indochina and to the Hainan Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consist ...

'', ''

Petraeovitex ''Petraeovitex'' is a genus of eight climbing shrubs species known to science, of the mint family Lamiaceae (formerly placed within Verbenaceae). Collectively, they grow naturally in Borneo, Peninsular Malaysia, Sumatra, the Philippines, the ...

'', ''

Peronema ''Peronema'' is a genus of flowering plant in the family Lamiaceae, first described in 1822. It contains only one known species, ''Peronema canescens'', native to Thailand, Malaysia, Borneo, Sumatra, and Java Java (; id, Jawa, ; jv, ꦗ� ...

'', ''

Garrettia ''Garrettia'' is a genus of flowering plant in the family Lamiaceae, first described in 1937. It contains only one known species, ''Garrettia siamensis''. It is native to southern China (Yunnan Province), Java, Thailand Thailand ( ), ...

'', ''

Cymaria ''Cymaria'' is a genus of plants in the family Lamiaceae, first described in 1830. It is native to China, New Guinea, and Southeast Asia.Acrymia ''Acrymia'' is a genus of plants in the family Lamiaceae, first described in 1908. Only one species is known, ''Acrymia ajugiflora'', found in the Perak and Selangor regions of Malaysia Malaysia ( ; ) is a country in Southeast Asia. The ...

'', ''

Holocheila ''Holocheila'' is a genus of flowering plant in the family Lamiaceae, first described as a genus in 1962. It contains only one known species, ''Holocheila longipedunculata''. It is endemic to Yunnan Province in China. References

Lamiacea ...

'', and '' Ombrocharis''. The subfamilies are the Symphorematoideae,

Viticoideae Viticoideae is one of seven subfamilies in the sage family, Lamiaceae. Genera This subfamily contains the following 10 genera: * '' Cornutia'' L. * ''Gmelina'' L. * '' Paravitex'' Fletcher * '' Petitia'' Jacq. * '' Pseudocarpidium'' Millsp. * ...

Ajugoideae Ajugoideae is subfamily of the family Lamiaceae. The subfamily name of Teucrioideae is a synonym of Ajugoideae. Genera *Tribe Ajugeae **''Acrymia'' Prain **''Ajuga'' L. **''Cymaria'' Benth. **''Garrettia'' H. R. Fletcher **''Holocheila'' *All o ...

, Prostantheroideae,

Nepetoideae Nepetoideae is a subfamily of plants in the family Lamiaceae. , the Angiosperm Phylogeny Website (APweb) accepted the following genera: The format of the authorities is not the IPNI standard. *'' Acanthomintha'' (A. Gray) Bentham & J. D. Hooker ...

Scutellarioideae Scutellarioideae is a subfamily of plants in the family Lamiaceae. Genera include: * ''Holmskioldia'' * ''Renschia'' * ''Scutellaria'' * ''Tinnea'' * ''Wenchengia'' References

{{Taxonbar, from=Q2889855 Lamiaceae Asterid subfamilies ...

, and

Lamioideae Lamioideae is a subfamily of plants in the family Lamiaceae. Genera include: *'' Acanthoprasium'' *'' Achyrospermum'' *'' Acrotome'' *'' Ajugoides'' *'' Anisomeles'' *''Ballota'' *'' Betonica'' *'' Brazoria'' *'' Chaiturus'' *'' Chamaesphacos'' ...

. The subfamily Viticoideae is probably not

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

. The Prostantheroideae and Nepetoideae are divided into

tribes The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to conflic ...

. These are shown in the

phylogenetic tree A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...

below.

Phylogeny

Most of the genera of Lamiaceae have never been

sampled Sample or samples may refer to: Base meaning * Sample (statistics), a subset of a population – complete data set * Sample (signal), a digital discrete sample of a continuous analog signal * Sample (material), a specimen or small quantity of so ...

for DNA for

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

studies. Most of those that have been are included in the following

. The

phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...

depicted below is based on seven different sources.

References

External links

* *
Lamiaceae
i
L. Watson and M.J. Dallwitz (1992 onwards). The families of flowering plants: descriptions, illustrations, identification, information retrieval.
https://delta-intkey.com {{Authority control Lamiales families