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Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropical lake, and the world's second-largest
fresh water Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
lake by surface area after Lake Superior in North America. In terms of volume, Lake Victoria is the world's ninth-largest continental lake, containing about of water. Lake Victoria occupies a shallow depression in Africa. The lake has an average depth of and a maximum depth of .United Nations, ''Development and Harmonisation of Environmental Laws Volume 1: Report on the Legal and Institutional Issues in the Lake Victoria Basin'', United Nations, 1999, page 17 Its catchment area covers . The lake has a shoreline of when digitized at the 1:25,000 level, with islands constituting 3.7% of this length. The lake's area is divided among three countries: Kenya occupies 6% (), Uganda 45% (), and Tanzania 49% (). Though having multiple local language names ( luo, Nam Lolwe; lg, 'Nnalubaale; rw, Nyanza; also ''Ukerewe''), the lake was renamed after Queen Victoria by the explorer John Hanning Speke, the first Briton to document it in 1858, while on an expedition with Richard Francis Burton. The lake is home to many species of fish which live nowhere else, especially
cichlid Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this ...
s.
Invasive Invasive may refer to: *Invasive (medical) procedure *Invasive species *Invasive observation, especially in reference to surveillance *Invasively progressive spread of disease from one organ in the body to another, especially in reference to cancer ...
fish, such as the Nile perch, have driven many endemic species to extinction.


Geology

Geologically, Lake Victoria is relatively young at about 400,000 years old. It formed when westward-flowing rivers were dammed by an upthrown crustal block. During the Miocene era, what is now the catchment area of the lake was on the western side of an uplifted area that functioned as a continental divide, with streams on the western side flowing into the Congo River basin and streams on the eastern side flowing to the Indian Ocean. As the
East African Rift The East African Rift (EAR) or East African Rift System (EARS) is an active continental rift zone in East Africa. The EAR began developing around the onset of the Miocene, 22–25 million years ago. In the past it was considered to be part of a ...
System formed, the eastern wall of the Albertine Rift (or Western Rift) rose, gradually reversing the drainage towards what is now Lake Victoria. The opening of the main East African Rift and the Albertine Rift downwarped the area between them as the rift walls rose, creating the current Lake Victoria basin. During its geological history, Lake Victoria went through changes ranging from its present shallow depression, through to what may have been a series of much smaller lakes. Geological cores taken from its bottom show Lake Victoria has dried up completely at least three times since it formed. These drying cycles are probably related to past ice ages, which were times when precipitation declined globally. Lake Victoria last dried out about 17,300 years ago, and it refilled 14,700 years ago as the African humid period began.


Hydrology and limnology

Lake Victoria receives 80 percent of its water from direct rainfall. Average evaporation on the lake is between per year, almost double the precipitation of riparian areas. Lake Victoria receives its water additionally from rivers, and thousands of small
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...
s. The Kagera River is the largest river flowing into this lake, with its
mouth In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on ...
on the lake's western shore. Lake Victoria is drained solely by the
Nile River The Nile, , Bohairic , lg, Kiira , Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered the longest rive ...
near Jinja, Uganda, on the lake's northern shore. In the Kenya sector, the main influent rivers are the
Sio Sio may refer to: Places * Sió, an artificial channel in Hungary * Siø, a small Danish island in the South Funen Archipelago * Sio, Burkina Faso, a village in Burkina Faso * Sio, Mali, a commune in Mali * Sio, Papua New Guinea, a town in ...
,
Nzoia The Nzoia River is a Kenyan river, rising from Cherangany hills. It passes through Kapsara, Springer ,Moi's bridge then crosses to Kakamega county.It flows south and then west, eventually flowing into Lake Victoria near the town of Port Victoria. ...
, Yala, Nyando,
Sondu Miriu Sondu is a small town border town in Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates ...
, Mogusi, and Migori. The only outflow from Lake Victoria is the Nile River, which exits the lake near Jinja, Uganda. In terms of contributed water, this makes Lake Victoria the principal source of the longest branch of the Nile. However, the most distal source of the Nile Basin, and therefore the ultimate source of the Nile, is more often considered to be one of the tributary rivers of the Kagera River (the exact tributary remains undetermined), and which originates in either
Rwanda Rwanda (; rw, u Rwanda ), officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of Central Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator ...
or
Burundi Burundi (, ), officially the Republic of Burundi ( rn, Repuburika y’Uburundi ; Swahili language, Swahili: ''Jamuhuri ya Burundi''; French language, French: ''République du Burundi'' ), is a landlocked country in the Great Rift Valley at the ...
. The uppermost section of the Nile is generally known as the Victoria Nile until it reaches Lake Albert. Although it is a part of the same river system known as the
White Nile The White Nile ( ar, النيل الأبيض ') is a river in Africa, one of the two main tributaries of the Nile, the other being the Blue Nile. The name comes from the clay sediment carried in the water that changes the water to a pale color. ...
and is occasionally referred to as such, strictly speaking this name does not apply until after the river crosses the Uganda border into South Sudan to the north. The lake exhibits eutrophic conditions. In 1990–1991, oxygen concentrations in the mixed layer were higher than in 1960–1961, with nearly continuous oxygen supersaturation in surface waters. Oxygen concentrations in hypolimnetic waters (i.e. the layer of water that lies below the thermocline, is noncirculating, and remains perpetually cold) were lower in 1990–1991 for a longer period than in 1960–1961, with values of less than 1 mg per litre (< 0.4  gr/cu ft) occurring in water as shallow as compared with a shallowest occurrence of greater than in 1961. The changes in oxygenation are considered consistent with measurements of higher algal biomass and productivity. These changes have arisen for multiple reasons: successive burning within its basin, soot and ash from which has been deposited over the lake's wide area; from increased nutrient inflows via rivers, and from increased pollution associated with settlement along its shores. Between 2010 and 2022 the surface area of Lake Victoria increased by 15% flooding lakeside communities.


Bathymetry

The lake is considered a shallow lake considering its large geographic area with a maximum depth of approximately and an average depth of . A 2016 project digitized ten-thousand points and created the first true bathymetric map of the lake. The deepest part of the lake is offset to the east of the lake near Kenya and the lake is generally shallower in the west along the Ugandan shoreline and the south along the Tanzanian shoreline.


Native wildlife


Mammals

Many mammal species live in the region of Lake Victoria, and some of these are closely associated with the lake itself and the nearby wetlands. Among these are the hippopotamus,
African clawless otter The African clawless otter (''Aonyx capensis''), also known as the Cape clawless otter or groot otter, is the second-largest freshwater otter species. It inhabits permanent water bodies in savannah and lowland forest areas through most of sub- ...
,
spotted-necked otter The spotted-necked otter (''Hydrictis maculicollis''), or speckle-throated otter, is an otter native to sub-Saharan Africa. Description The spotted-necked otter is a relatively small species, with males measuring from nose to rump, and weigh ...
,
marsh mongoose The marsh mongoose (''Atilax paludinosus''), also known as the water mongoose or the vansire, is a medium-sized mongoose native to sub-Saharan Africa that inhabits foremost freshwater wetlands. It has been listed as Least Concern on the IUCN Red L ...
,
sitatunga The sitatunga or marshbuck (''Tragelaphus spekii'') is a swamp-dwelling antelope found throughout central Africa, centering on the Democratic Republic of the Congo, the Republic of the Congo, Cameroon, parts of Southern Sudan, Equatorial Guinea, ...
,
bohor reedbuck The bohor reedbuck (''Redunca redunca'') is an antelope native to central Africa. The animal is placed under the genus '' Redunca'' and in the family Bovidae. It was first described by German zoologist and botanist Peter Simon Pallas in 1767. T ...
,
defassa waterbuck The waterbuck (''Kobus ellipsiprymnus'') is a large antelope found widely in sub-Saharan Africa. It is placed in the genus '' Kobus'' of the family Bovidae. It was first described by Irish naturalist William Ogilby in 1833. Its 13 subspecies ...
, cane rats, and giant otter shrew.


Reptiles

Lake Victoria and its wetlands has a large population of Nile crocodiles, as well as
African helmeted turtle The African helmeted turtle (''Pelomedusa subrufa''), also known commonly as the marsh terrapin, the crocodile turtle, or in the pet trade as the African side-necked turtle, is a species of omnivorous side-necked terrapin in the family Pelomedus ...
s,
variable mud turtle The variable mud turtle (''Pelusios rhodesianus''), also known as Rhodesian mud turtle, Mashona hinged terrapin or variable hinged terrapin, is a species of turtle in the family Pelomedusidae. It is widely distributed in Central, East, and Southe ...
s, and
Williams' mud turtle Williams' mud turtle (''Pelusios williamsi'') is a species of turtle in the family Pelomedusidae. The species is endemic to Africa. Etymology The specific name, ''williamsi'', is in honor of American herpetologist Ernest E. Williams.Beolens, B ...
.Spawls, Howell, Drewes, and Ashe (2002). ''A Field Guide to the Reptiles of East Africa''. Academic Press, London. . The Williams' mud turtle is restricted to Lake Victoria and other lakes, rivers, and swamps in the upper Nile basin.


Cichlid fish

Lake Victoria formerly was very rich in fish, including many endemics, but a high percentage of these became extinct since the 1940s. The main group in Lake Victoria is the
haplochromine __NOTOC__ The haplochromine cichlids are a tribe of cichlids in subfamily Pseudocrenilabrinae called Haplochromini. This group includes the type genus (''Haplochromis'') plus a number of closely related genera such as '' Aulonocara'', '' Astat ...
cichlids ('' Haplochromis'' '' sensu lato'') with more than 500 species, almost all endemic,DeWeerdt, S. (28 February 2004)
Dark secret of the lake.
New Scientist. Retrieved 26 March 2017.
and including an estimated 300 that still are undescribed.Sayer, C.A., L. Máiz-Tomé, and W.R.T. Darwall (2018). Freshwater biodiversity in the Lake Victoria Basin: Guidance for species conservation, site protection, climate resilience and sustainable livelihoods. Cambridge, UK and Gland, Switzerland: IUCN. This is far more species of fish than any other lake in the world, except Lake Malawi. These are the result of a rapid
adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
in the last circa 15,000 years. Their extraordinary diversity and speed of evolution have been the subjects for many scientists studying the forces that drive the richness of life everywhere. The Victoria haplochromines are part of an older group of more than 700 closely related species, also including those of several smaller lakes in the region, notably Kyoga,
Edward Edward is an English given name. It is derived from the Anglo-Saxon name ''Ēadweard'', composed of the elements '' ēad'' "wealth, fortune; prosperous" and '' weard'' "guardian, protector”. History The name Edward was very popular in Anglo-Sa ...
George, Albert, and Kivu. Most of these lakes are relatively shallow (like Victoria) and part of the present-day upper Nile basin. The exception is Lake Kivu, which is part of the present-day Congo River basin, but is believed to have been connected to Lakes Edward and Victoria by rivers until the uplifting of parts of the
East African Rift The East African Rift (EAR) or East African Rift System (EARS) is an active continental rift zone in East Africa. The EAR began developing around the onset of the Miocene, 22–25 million years ago. In the past it was considered to be part of a ...
. This deep lake may have functioned as an "evolutionary reservoir" for this haplochromine group in periods where other shallower lakes in the region dried out, as happened to Lake Victoria about 15,000 years ago. In recent history only Lake Kyoga was easily accessible to Victoria cichlids, as further downstream movement by the
Victoria Nile The White Nile ( ar, النيل الأبيض ') is a river in Africa, one of the two main tributaries of the Nile, the other being the Blue Nile. The name comes from the clay sediment carried in the water that changes the water to a pale color. ...
(to Lake Albert) is prevented by a series of waterfalls, notably Murchison. In contrast, the
Owen Falls The Owen Falls was a waterfall on the White Nile in Uganda near the city of Jinja. The falls, together with the nearby Ripon Falls, were submerged in 1954 with the completion of the Nalubaale Hydroelectric Power Station. The dam that houses ...
(now flooded by a dam) between Victoria and Kyoga were essentially a series of rapids that did not effectively block fish movements between the two lakes. The Victoria haplochromines are distinctly sexually dimorphic (males relatively brightly colored; females dull), and their ecology is extremely diverse, falling into at least 16 groups, including
detritivore Detritivores (also known as detrivores, detritophages, detritus feeders, or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrates, ...
s, zooplanktivores,
insectivore A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores wer ...
s, prawn-eaters, molluscivores and piscivores. As a result of predation by the introduced Nile perch, eutrophication and other changes to the ecosystem, it is estimated that at least 200 species (about 40 percent) of Lake Victoria haplochromines have become extinct, including more than 100 undescribed species. Initially it was feared that this number was even higher, by some estimates 65 percent of the total species, but several species that were feared extinct have been rediscovered after the Nile perch started to decline in the 1990s.IUCN Red Lists
Geographic Patterns
Eastern Africa. Retrieved 25 March 2017.
Several of the remaining species are seriously threatened and additional extinctions are possible.Fiedler, P.L. and P M. Kareiva, editors (1998). Conservation Biology: For the Coming Decade. 2nd edition. pp. 209–10. Some species have survived in nearby small satellite lakes, have survived in refugias among rocks or papyrus sedges (protecting them from the Nile perch), or have adapted to the human-induced changes in the lake itself. Such adaptions include a larger gill area (adaption for oxygen-poor water), changes in the feeding apparatus, changes to the eyes (giving them a better sight in turbid water) and smaller head/larger caudal peduncle (allowing faster swimming). The piscivorous (affected by both predation and competition from Nile perch), molluscivorous and insectivorous haplochromines were particularly hard hit with many extinctions. Others have become extinct in their pure form, but survive as hybrids between close relatives (especially among the detritivores). The zooplanktivores have been least affected and in the late 1990s had reached densities similar to, or above, the densities before the drastic declines, although consisting of fewer species and often switching their diet towards
macroinvertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate s ...
s. Some of the threatened Lake Victoria cichlid species have captive "insurance" populations in zoos, public aquaria and among private aquarists, and a few species are
extinct in the wild A species that is extinct in the wild (EW) is one that has been categorized by the International Union for Conservation of Nature as known only by living members kept in captivity or as a naturalized population outside its historic range due ...
(only survive in captivity). Before the mass extinction that has occurred among the lake's cichlids in the last 50 years, about 90 percent of the native fish species in the lake were haplochromines. Disregarding the haplochromines, the only native Victoria cichlids are two critically endangered tilapia, the
Singida tilapia ''Oreochromis esculentus'', the Singida tilapia or Graham's tilapia, is a species of cichlid endemic to the Lake Victoria basin, including some of its satellite lakes such as Kyoga, in Tanzania, Uganda, and Kenya. Its common name refers to ...
or ''ngege'' (''Oreochromis esculentus'') and Victoria tilapia (''O. variabilis''). In 1927–1928 Michael Graham conducted the first ever systematic
Fisheries Survey of Lake Victoria Lake Victoria supports Africa's largest inland fishery, with the majority of present catch being the invasive Nile perch, introduced to the Lake in the 1950s. Prior to the introduction of Nile perch as well as Nile tilapia, the fish community w ...
. In his official report of the expedition, Graham wrote that "The ngege or satu ''Tilapia esculenta'', is the most important food fish of the lake, whether for native or non-native consumption. No other fish equals it in the quality of the flesh. It is convenient size for trade, travels well and is found in much greater numbers than other important fish, such as semutundu (Luganda), ''Bagrus sp.''".Graham M. (1929.) The Victoria Nyanza and Its Fisheries: A Report on the Fish Survey of Lake Victoria 1927–1928 and Appendices. London: Crown Agents for the Colonies. 256pp. Furthermore, Graham noted that the introduction of the European flax gill net of 5 inch mesh had undoubtedly caused a diminution in the number of ngege in those parts of the Kavirondo Gulf, the northern shore of the lake, the Sesse Islands and Smith's Sound which are conveniently situated close to markets. Survey catches in 1927–28 included several Haplochromis species that are now thought to be extinct, including: ''
Haplochromis flavipinnis ''Haplochromis flavipinnis'' is a species of cichlid endemic to Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropi ...
'', ''
Haplochromis gowersii ''Haplochromis gowersii'' is a species of cichlid endemic to Lake Victoria. This species can reach a length of SL. The specific name of this taxon honours the former Governor of Uganda William Frederick Gowers Sir William Frederick Gowers, ...
'', ''
Haplochromis longirostris ''Haplochromis longirostris'' is a species of cichlid endemism, endemic to Lake Victoria though it may now be extinct. This species can reach a length of fish measurement, SL. References

Haplochromis, longirostris Fish described in 188 ...
'', ''
Haplochromis macrognathus ''Haplochromis macrognathus'' is a species of cichlid fish endemic to Lake Victoria in East Africa. This piscivorous species can reach a standard length of . Last seen in the early 1980s, it may now be extinct. References macrognathus ''M ...
'', '' Haplochromis michaeli'', ''
Haplochromis nigrescens ''Haplochromis nigrescens'' is a species of cichlid endemic to Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropic ...
'', ''
Haplochromis prognathus ''Haplochromis prognathus'' is a species of cichlid endemic to Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropic ...
''. As well as being due to the introduction of Nile Perch, the extinction of cichlids in the genus '' Haplochromis'' has also been blamed on the lake's eutrophication. The fertility of tropical waters depends on the rate at which nutrients can be brought into solution. The influent rivers of Lake Victoria provide few nutrients to the lake in relation to its size. Because of this, most of Lake Victoria's nutrients are thought to be locked up in lake-bottom deposits. By itself, this vegetative matter decays slowly. Animal flesh decays considerably faster, however, so the fertility of the lake is dependent on the rate at which these nutrients can be taken up by fish and other organisms. There is little doubt that ''Haplochromis'' played an important role in returning detritus and plankton back into solution. With some 80 percent of ''Haplochromis'' species feeding off detritus, and equally capable of feeding off one another, they represented a tight, internal recycling system, moving nutrients and biomass both vertically and horizontally through the water column, and even out of the lake via predation by humans and terrestrial animals. The removal of ''Haplochromis'', however, may have contributed to the increasing frequency of algal blooms, which may in turn be responsible for mass
fish kill The term fish kill, known also as fish die-off, refers to a localized die-off of fish populations which may also be associated with more generalized mortality of aquatic life.University of Florida. Gainesville, FL (2005) ''Plant Management in Fl ...
s.


Other fish

The non-cichlid native fish include African tetras (''
Brycinus ''Brycinus'' is a genus of ray-finned fish in the family Alestiidae. Like other "African characids", they were formerly included in the Characidae but are actually somewhat more distantly related Characiformes. Like some other Alestiidae, they a ...
''), cyprinids (''
Enteromius ''Enteromius'' is a genus of small to medium-sized cyprinid fish native to tropical Africa. Most species were placed in the genus ''Barbus ''Barbus'' is a genus of ray-finned fish in the family Cyprinidae. The type species of ''Barbus'' is ...
'', '' Garra'', '' Labeo'', '' Labeobarbus'', ''
Rastrineobola The silver cyprinid (''Rastrineobola argentea'') also known as the Lake Victoria sardine, mukene,and omena (native language), dagaa (swahili language) is a species of pelagic, freshwater ray-finned fish in the carp family, Cyprinidae from East A ...
'' and '' Xenobarbus''), airbreathing catfish (''
Clariallabes ''Clariallabes'' is a genus of airbreathing catfishes found in Africa. Species There are currently 16 recognized species in this genus: * '' Clariallabes attemsi'' ( Holly, 1927) * '' Clariallabes brevibarbis'' Pellegrin, 1913 * '' Clariallab ...
'', ''
Clarias ''Clarias'' is a genus of catfishes (order Siluriformes) of the family Clariidae, the airbreathing catfishes. The name is derived from the Greek ''chlaros'', which means lively, in reference to the ability of the fish to live for a long time out ...
'' and ''
Xenoclarias The Lake Victoria deepwater catfish (''Xenoclarias eupogon'') is a species of catfish ( order Siluriformes) of the family Clariidae. It is monotypic within the genus ''Xenoclarias''. This species is endemic to Lake Victoria, and is found in dee ...
''), bagrid catfish (''
Bagrus ''Bagrus'' is a genus of bagrid catfishes. These are relatively large catfish found in freshwater habitats in Africa, except for the virtually unknown ''B. tucumanus'' from South America, which likely is a Synonym (biology), synonym of ''Luciop ...
''), loach catfish (''
Amphilius ''Amphilius'' is a genus of catfishes of the family Amphiliidae. ''Amphilius'' catfish have fairly lengthened bodies, with short, depressed, and broad heads. They have three pairs of fringed barbels. The eyes, small and located dorsally, are v ...
'' and ''
Zaireichthys ''Zaireichthys'' is a genus of loach catfishes ( order Siluriformes) of the family Amphiliidae. Distribution ''Zaireichthys'' species are found in Africa. Description ''Zaireichthys'' species are quite small ranging from 2.1 cm (''P. lacu ...
''), silver butter catfish (''Schilbe intermedius''), ''
Synodontis ''Synodontis'' is the largest genus of mochokid catfishes. It is the biggest genus within the 10 genera and 190 different species in the family Mochokidae. ''Synodontis'' has over 131 different species within the genus. ''Synodontis'' are also k ...
'' squeaker catfish, ''
Nothobranchius ''Nothobranchius'' is a genus of small, freshwater killifish, classified in either the family Nothobranchiidaevan der Laan, R., Eschmeyer, W.N. & Fricke, R. (2014)Family-group names of Recent fishes.''Zootaxa, 3882 (2): 1–230.'' or a more inc ...
'' killifish, poeciliids (''
Aplocheilichthys The banded lampeye (''Aplocheilichthys spilauchen'') is a species of poeciliid that is native to Africa, ranging from Senegal to Angola. It is mainly found in coastal brackish habitats (only infrequently in pure fresh or salt water) such as river ...
'' and ''
Micropanchax ''Micropanchax'' is a genus of poeciliids native to Africa. Species The 13 recognized species in this genus are: * ''Micropanchax antinorii'' (Vinciguerra, 1883) (black lampeye) * ''Micropanchax bracheti'' (Heinz Otto Berkenkamp, Berkenkamp, 19 ...
''), the spiny eel '' Mastacembelus frenatus'', elephantfish (''
Gnathonemus ''Gnathonemus'' is a genus of elephantfish in the family Mormyridae. Species There are currently four recognized species in this genus: * '' Gnathonemus barbatus'' Poll 1967 (Angolan mormyrid) * '' Gnathonemus echidnorhynchus'' Pellegrin 192 ...
'', ''
Hippopotamyrus ''Hippopotamyrus'' is a genus of elephantfish in the family Mormyridae. Species There are currently 11 recognized species in this genus: * '' Hippopotamyrus ansorgii'' ( Boulenger 1905) (Slender stonebasher) * '' Hippopotamyrus castor'' Pappe ...
'', ''
Marcusenius ''Marcusenius'' is a genus of elephantfishes native to Africa. Its members are highly diverse in size, with the smallest species reaching less than and the largest more than . The genus is named after Johann Marcusen. Species There are cur ...
'', ''
Mormyrus ''Mormyrus'' is a genus of ray-finned fish in the family Mormyridae. Species There are currently 22 recognized species in this genus: * '' Mormyrus bernhardi'' Pellegrin 1926 (Bernhard's elephant-snout fish) * '' Mormyrus caballus'' Boul ...
'', '' Petrocephalus'', and ''
Pollimyrus ''Pollimyrus'' is a genus of Mormyridae, elephantfishes native to Africa. Species There are currently 19 recognized species in this genus: * ''Pollimyrus adspersus'' (Albert Günther, Günther 1866) * ''Pollimyrus brevis'' (George Albert Boulen ...
''), the climbing gourami '' Ctenopoma muriei'' and marbled lungfish (''Protopterus aethiopicus'').FishBase
Fish Species in Victoria.
Retrieved 25 March 2017.
At a genus level, most of these are widespread in Africa, but the very rare ''Xenobarbus'' and ''Xenoclarias'' are endemic to the lake, and the common ''Rastrineobola'' is near-endemic.


Crustaceans

Four species of
freshwater crab Around 1,300 species of freshwater crabs are distributed throughout the tropics and subtropics, divided among eight families. They show direct development and maternal care of a small number of offspring, in contrast to marine crabs, which relea ...
s are known from Lake Victoria: ''
Potamonautes niloticus ''Potamonautes niloticus'' is a species of freshwater crab in the family Potamonautidae. It is found in the Nile Basin in Egypt, Ethiopia, Rwanda, Sudan, and Uganda. Numerous specimens of ''P. niloticus'' are known from Miocene The Miocene ( ...
'' is widespread in the lake and '' P. emini'' has been recorded from the vicinity of Bukoba in Tanzania, but both are also found elsewhere in Africa. The last were first scientifically described in 2017 and very little is known about them: '' P. entebbe'' is only known from near Entebbe (the only known specimen was collected in 1955 and it is unknown if it was in or near the lake) and '' P. busungwe'' only at Busungwe Island in the northwestern part of the lake. The latter likely is the smallest African freshwater crab with a carapace width up to about , although '' P. kantsyore'' of Kagera River, and ''
Platythelphusa ''Platythelphusa'' is a genus of freshwater crabs endemic to Lake Tanganyika. It has been placed in a number of families, including a monotypic family, Platythelphusidae, as well as Potamidae and its current position in the Potamonautidae, and ha ...
maculata'' and ''P. polita'' of Lake Tanganyika are almost as small. The only shrimp/prawn is ''
Caridina nilotica ''Caridina nilotica'' is a species of freshwater shrimp in the family Atyidae. It is native to Africa from the River Nile in Egypt to Lake Sibaya, South Africa, and is the only species of shrimp in Lake Victoria Lake Victoria is one of th ...
'', which is common and widespread in Lake Victoria.


Molluscs

Lake Victoria is home to 28 species of
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
s (e.g., ''
Bellamya ''Bellamya'' was a supertanker, built in 1976 by Chantiers de l'Atlantique at Saint-Nazaire for the French branch of Shell Oil. She was the second Batillus class supertanker. ''Bellamya'', together with her sister ships ''Batillus, Pierre Guillau ...
'', '' Biomphalaria'', '' Bulinus'', ''
Cleopatra Cleopatra VII Philopator ( grc-gre, Κλεοπάτρα Φιλοπάτωρ}, "Cleopatra the father-beloved"; 69 BC10 August 30 BC) was Queen of the Ptolemaic Kingdom of Egypt from 51 to 30 BC, and its last active ruler.She was also a ...
'', '' Gabbiella'', and '' Melanoides''), including 12 endemic species/subspecies. There are 17 species of
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
s ('' Corbicula'', ''
Coelatura ''Coelatura'' is a genus of bivalve in the Unionidae family. Species Species: *''Coelatura aegyptiaca'' *''Coelatura alluaudi'' *''Coelatura bakeri'' *''Coelatura briarti'' *''Coelatura choziensis'' *''Coelatura cridlandi'' *''Coelatura d ...
'', ''
Sphaerium ''Sphaerium'' is a genus of very small freshwater clams, aquatic bivalve molluscs in the family Sphaeriidae, known as the fingernail clams. The small clams in this genus are unusual in that many of them, such as ''Sphaerium corneum'', can climb ...
'', and '' Byssanodonta''), including 6 endemic species/subspecies. It is likely that undescribed species of snails remain. Conversely, genetic studies indicate that some morphologically distinctive populations, traditionally regarded as separate species, may only be variants of single species. Two of the snail genera, ''Biomphalaria'' and ''Bulinus'', are
intermediate host In biology and medicine, a host is a larger organism that harbours a smaller organism; whether a parasitic, a mutualistic, or a commensalist ''guest'' (symbiont). The guest is typically provided with nourishment and shelter. Examples include a ...
s of the parasite that causes bilharzia (schistosomiasis). Human infections by this parasite are common at Lake Victoria. This may increase as a result of the spread of the invasive water hyacinth (an optimum snail habitat), and the loss of many snail-eating cichlids in the lake.


Spiders

'' Evarcha culicivora'' is a species of jumping spider (family Salticidae) found only around Lake Victoria in Kenya and Uganda. It feeds primarily on female mosquitos.


Fisheries

Lake Victoria supports Africa's largest inland
fishery Fishery can mean either the enterprise of raising or harvesting fish and other aquatic life; or more commonly, the site where such enterprise takes place ( a.k.a. fishing ground). Commercial fisheries include wild fisheries and fish farms, both ...
(as of 1997). Initially the fishery involved native species, especially tilapia and haplochromine cichlids, but also catfish (''Bagrus'', ''Clarias'', ''Synodontis'' and silver butter catfish), elephantfish,
ningu The ningu (''Labeo victorianus'') is a species of ray-finned fish in the family Cyprinidae. It is found in the Lake Victoria basin in Burundi, Kenya, Tanzania, and Uganda. Its natural habitats are rivers, swamps, freshwater lakes, freshwater mars ...
(''Labeo victorianus'') and marbled lungfish (''Protopterus aethiopicus'').Lake Victoria Fisheries Organization (2016). Some of these, including tilapia and ningu (''Labeo victorianus''), had already declined in the first half of the 20th century due to
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in th ...
. To boost fishing, several species of non-native tilapia and Nile perch were introduced to the lake in the 1950s. Nevertheless, the natives continued to dominate fisheries until the 1970s where their decline meant that there was a strong shift towards the non-native Nile tilapia (now 7 percent of catches), non-native Nile perch (60 percent) and the native Lake Victoria sardine (30 percent). Because of its small size, the abundant open-water Lake Victoria sardine only supported minor fisheries until the decline of other natives. At the peak in the early 1990s, of Nile perch were landed annually in Lake Victoria, but this has declined significantly in later years.


Environmental issues

A number of environmental issues are associated with Lake Victoria and the complete disappearance of many endemic cichlid species has been called the "most dramatic example of human-caused extinctions within an ecosystem".


Invasive fish

Starting in the 1950s, many species have been introduced to Lake Victoria where they have become
invasive Invasive may refer to: *Invasive (medical) procedure *Invasive species *Invasive observation, especially in reference to surveillance *Invasively progressive spread of disease from one organ in the body to another, especially in reference to cancer ...
and a prime reason for the extinction of many endemic
haplochromine __NOTOC__ The haplochromine cichlids are a tribe of cichlids in subfamily Pseudocrenilabrinae called Haplochromini. This group includes the type genus (''Haplochromis'') plus a number of closely related genera such as '' Aulonocara'', '' Astat ...
cichlids. Among the introductions are several tilapias: redbreast (''Coptodon rendalli''), redbelly (''C. zillii''), Nile (''Oreochromis niloticus'') and blue-spotted tilapias (''O. leucostictus'').Pringle, R.M. (2005)
The Origins of the Nile Perch in Lake Victoria.
BioScience 55 (9): 780–787.
Although these have contributed to the extinction of native fish by causing significant changes to the ecosystem, outcompeted natives and (in the case of the Nile tilapia) possibly hybridized with the highly threatened native tilapias, the most infamous introduction was the large and highly predatory Nile perch (''Lates niloticus''). As early as the 1920s, it was proposed to introduce a large pelagic predator such as the Nile perch to improve the fisheries in the lake. At the same time it was warned that this could present a serious danger to the native fish species and required extensive research into possible ecological effects before done. These warnings primarily concerned the native tilapia ''O. esculentus'', as the smaller haplochromine cichlids (despite playing an important role in local fisheries) were regarded as "trash fish" by the colonial government. In the following decades, the pressure to introduce the Nile perch continued, as did warnings about the possible effects of doing it. The first introduction of Nile perch to the region, done by the Uganda Game and Fisheries Department (then part of the colonial government) and local African fish guards, happened upstream of Murchison Falls directly after the completion of the Owen Falls Dam in 1954. This allowed it to spread to Lake Kyoga where additional Nile perch were released in 1955, but not Victoria itself. Scientists argued that further introduction should wait until research showed the effect of the introduction in Kyoga, but by the late 1950s, Nile perch began being caught in Lake Victoria. As the species was already present, there were few objections when more Nile perch were transferred to Victoria to further bolster the stock in 1962–63. The origin of the first Victoria introductions in the 1950s is not entirely clear and indisputable evidence is lacking. Uganda Game and Fisheries Department (UGFD) officials denied that they were involved, but circumstantial evidence suggests otherwise and local Africans employed by UGFD have said that they introduced the species in 1954–55 under the directive of senior officials. UGFD officials argued that Nile perch must have spread to Lake Victoria by themselves by passing through the Owen Falls Dam when shut down for maintenance, but this is considered highly unlikely by many scientists. The Nile perch had spread throughout the lake by 1970. Initially the population of the Nile perch was relatively low, but a drastic increase happened, peaking in the 1980s, followed by a decline starting in the 1990s. Due to the presence of the Nile perch, the natural balance of the lake's ecosystem has been disrupted. The food chain is being altered and in some cases, broken by the indiscriminate eating habits of the Nile perch. The subsequent decrease in the number of algae-eating fish allows the algae to grow at an alarming rate, thereby choking the lake. The increasing amounts of algae, in turn, increase the amount of detritus (dead plant material) that falls to the deeper portions of the lake before decomposing. As a by-product of this the oxygen levels in the deeper layer of water are being depleted. Without oxygen, any aerobic life (such as fish) cannot exist in the deeper parts of the lake, forcing all life to exist within a narrow range of depth. In this way, the Nile perch has degraded the diverse and thriving ecosystem that was once Lake Victoria. The abundance of aquatic life is not the only dependent of the lake: more than thirty million people in Tanzania, Kenya and Uganda rely on the lake for its natural resources. Hundreds of endemic species that evolved under the special conditions offered by the protection of Lake Victoria have been lost due to extinction, and several more are still threatened. Their loss is devastating for the lake, the fields of ecology, genetics and evolution biology, and more evidently, for the local fisheries. Local fisheries once depended on catching the lungfish, tilapia, carp and catfish that comprise the local diet. Today, the composition and yields of such fish catches are virtually negligible. Extensive fish kills, Nile perch, loss of habitat and overfishing have caused many fisheries to collapse and many protein sources to be unavailable at the market for local consumption. Few fisheries, though, have been able to make the switch to catching the Nile perch, since that requires a significant amount of capital resources.


Water hyacinth invasion

The water hyacinth has become a major
invasive plant species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
in Lake Victoria. The release of large amounts of untreated wastewater (sewage) and agricultural and industrial runoff directly into Lake Victoria over the past 30 years has greatly increased the nutrient levels of nitrogen and phosphorus in the lake "triggering massive growth of exotic water hyacinth, which colonised the lake in the late 1990s".Luilo, G.B. (August 01, 2008). Lake Victoria water resources management challenges and prospects: a need for equitable and sustainable institutional and regulatory frameworks ''African Journal of Aquatic Science'' 33, 2, 105–13. This invasive weed creates anoxic (total depletion of oxygen levels) conditions in the lake inhibiting decomposing plant material, raising toxicity and disease levels to both fish and people. At the same time, the plant's mat or "web" creates a barrier for boats and ferries to maneuver, impedes access to the shoreline, interferes with hydroelectric power generation, and blocks the intake of water for industries.Kateregga, E., & Sterner, T. (January 01, 2009). "Lake Victoria Fish Stocks and the Effects of Water Hyacinth". ''Journal of Environment & Development'', 18, 1, 62–78.Albright, T.P., Moorhouse, T.G., & McNabb, T.J. (January 1, 2004). "The Rise and Fall of Water Hyacinth in Lake Victoria and the Kagera River Basin, 1989-2001". ''Journal of Aquatic Plant Management'', 42, 73–84. On the other hand, water hyacinth mats can potentially have a positive effect on fish life in that they create a barrier to overfishing and allow for fish growth, there has even been the reappearance of some fish species thought to have been extinct in recent years. The overall effects of the water hyacinth, however, are still unknown. Growth of the water hyacinth in Lake Victoria has been tracked since 1993, reaching its maxima biomass in 1997 and then declining again by the end of 2001. Greater growth was observed in the northern part of the lake, in relatively protected areas, which may be linked to current and weather patterns and could also be due to the climate and water conditions, which are more suitable to the plants growth (as there are large urban areas to the north end of the lake, in Uganda). The invasive weed was first attempted to be controlled by hand, removed manually from the lake; however, re-growth occurred quickly. Public awareness exercises were also conducted. More recently, measures have been used such as the introduction of natural insect predators, including two different water hyacinth weevils and large harvesting and chopping boats, which seem to be much more effective in eliminating the water hyacinth. A green power plant that uses harvested water hyacinth (but also can use other degradable waste) was constructed in Kisumu County in 2013. In addition to the biogas it produces, its by-product can be used as fertilizer. Other factors which may have contributed to the decline of the water hyacinth in Lake Victoria include varying weather patterns, such as El Niño during the last few months of 1997 and first six months of 1998 bringing with it higher levels of water in the lake and thus dislodging the plants. Heavy winds and rains along with their subsequent waves may have also damaged the plants during this same time frame. The plants may not have been destroyed, instead merely moved to another location. Additionally, the water quality, nutrient supply, temperature, and other environmental factors could have played a role. Overall, the timing of the decline could be linked to all of these factors and perhaps together, in combination, they were more effective than any one deterrent would have been by itself. The water hyacinth is in remission and this trend could be permanent if control efforts are continued.


Pollution

Pollution of Lake Victoria is mainly due to discharge of
raw sewage Sewage (or domestic sewage, domestic wastewater, municipal wastewater) is a type of wastewater that is produced by a community of people. It is typically transported through a sewer system. Sewage consists of wastewater discharged from residenc ...
into the lake, dumping of domestic and industrial waste, and fertiliser and chemicals from farms. The Lake Victoria basin, while generally rural, has many major centres of population. Its shores are dotted with key cities and towns, including Kisumu, Kisii, and Homa Bay in Kenya; Kampala, Jinja and Entebbe in Uganda; and Bukoba, Mwanza, and Musoma in Tanzania. These cities and towns are also home to many factories that discharge some chemicals directly into the lake or its influent rivers. The set up of small beaches and local authorities around the lake lack proper sewage treatment facilities allowing pollutants to find their way into the water. Large parts of these urban areas also discharge untreated (raw) sewage into the river, increasing its eutrophication that in turn is helping to increase the invasive water hyacinth. Increased
logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidding, on-site processing, and loading of trees or logs onto trucks or skeleton cars. Logging is the beginning of a supply chain ...
and act of deforestation has led to environmental degradation around the region reducing the absorption of polluting chemicals and deteriorating the water quality.


Environmental data

As of 2016, an environmental data repository exists for Lake Victoria. The repository contains shoreline, bathymetry, pollution, temperature, wind vector, and other important data for both the lake and the wider Basin.


History and exploration

The first recorded information about Lake Victoria comes from Arab traders plying the inland routes in search of gold, ivory, other precious commodities, and
slave Slavery and enslavement are both the state and the condition of being a slave—someone forbidden to quit one's service for an enslaver, and who is treated by the enslaver as property. Slavery typically involves slaves being made to perf ...
s. The lake existed and was known to many Africans in the catchment area long before it was sighted by a
European European, or Europeans, or Europeneans, may refer to: In general * ''European'', an adjective referring to something of, from, or related to Europe ** Ethnic groups in Europe ** Demographics of Europe ** European cuisine, the cuisines of Europe ...
in 1858 when the British explorer John Hanning Speke reached its southern shore while on his journey with Richard Francis Burton to explore central Africa and locate the Great Lakes. Believing he had found the source of the Nile on seeing this "vast expanse of open water" for the first time, Speke named the lake after Queen Victoria. Burton, who had been recovering from illness at the time and resting further south on the shores of
Lake Tanganyika Lake Tanganyika () is an African Great Lake. It is the second-oldest freshwater lake in the world, the second-largest by volume, and the second-deepest, in all cases after Lake Baikal in Siberia. It is the world's longest freshwater lake. ...
, was outraged that Speke claimed to have proved his discovery to have been the true source of the Nile River, which Burton regarded as still unsettled. A very public quarrel ensued, which not only sparked a great deal of intense debate within the scientific community of the day, but also much interest by other explorers keen to either confirm or refute Speke's discovery. In the late 1860s, the famous Scottish explorer and missionary David Livingstone failed in his attempt to verify Speke's discovery, instead pushing too far west and entering the River Congo system instead. Ultimately, the
Welsh-American Welsh Americans ( cy, Americanwyr Cymreig) are an American ethnic group whose ancestry originates wholly or partly in Wales. In the 2008 U.S. Census community survey, an estimated 1.98 million Americans had Welsh ancestry, 0.6% of the total U.S ...
explorer
Henry Morton Stanley Sir Henry Morton Stanley (born John Rowlands; 28 January 1841 – 10 May 1904) was a Welsh-American explorer, journalist, soldier, colonial administrator, author and politician who was famous for his exploration of Central Africa Cen ...
, on an expedition funded by the ''
New York Herald The ''New York Herald'' was a large-distribution newspaper based in New York City that existed between 1835 and 1924. At that point it was acquired by its smaller rival the ''New-York Tribune'' to form the '' New York Herald Tribune''. His ...
'' newspaper, confirmed the truth of Speke's discovery, circumnavigating the lake and reporting the great outflow at Ripon Falls on the lake's northern shore.


Nalubaale Dam

The only outflow for Lake Victoria is at Jinja, Uganda, where it forms the Victoria Nile. The water for at least 12,000 years has drained across a natural rock weir. In 1952, engineers acting for the government of
Colonial Uganda The Protectorate of Uganda was a protectorate of the British Empire from 1894 to 1962. In 1893 the Imperial British East Africa Company transferred its administration rights of territory consisting mainly of the Kingdom of Buganda to the Brit ...
blasted out the weir and reservoir to replace it with an artificial barrage to control the level of the lake and reduce the gradual erosion of the rock weir. A standard for mimicking the old rate of outflow called the "agreed curve" was established, setting the maximum flow rate at 300 to 1,700
cubic metres per second A cubic metre per second (m3s−1, m3/s, cumecs or cubic meter per second in American English) is the unit of volumetric flow rate in the International System of Units (SI) equal to that of a stere or cube with sides of in length exchanged ...
(392–2,224 cu yd/sec) depending on the lake's water level. In 2002, Uganda completed a second
hydroelectric Hydroelectricity, or hydroelectric power, is electricity generated from hydropower (water power). Hydropower supplies one sixth of the world's electricity, almost 4500 TWh in 2020, which is more than all other renewable sources combined and ...
complex in the area, the Kiira Hydroelectric Power Station, with World Bank assistance. By 2006, the water levels in Lake Victoria had reached an 80-year low, and Daniel Kull, an independent hydrologist living in Nairobi, Kenya, calculated that Uganda was releasing about twice as much water as is allowed under the agreement, and was primarily responsible for recent drops in the lake's level.


Water use

Many towns and cities are reliant on Victoria for their water supplies, for farming and other uses.


Lamadi water scheme

The Lamadi water scheme is a water and sanitation project that serves Mwanza and the satellite towns of Lamadi, Misungwi, Magu, Bukoba, and Musoma on the bank of Lake Victoria.  European Investment Bank started the project in 2013 with the aim of protecting the environmental health of the lake, through improved water and sanitation to the towns whose pollution is part of the degradation of the lake. The project aims to provide safe drinking water for an estimated one million people and improved sanitation for 100 000 people. Sediment and suspended solids are filtered out using sand, which acts like a sieve. The water is then ready to be chlorinated or treated in another way. The sand filtration helps reduce water-borne diseases and is based on the use of the local environment.


Transport

Since the 1900s, Lake Victoria ferries have been an important means of transport between Uganda, Tanzania, and Kenya. The main ports on the lake are Kisumu, Mwanza, Bukoba, Entebbe, Port Bell, and Jinja. Until 1963, the fastest and newest ferry, MV ''Victoria'', was designated a Royal Mail Ship. In 1966, train ferry services between Kenya and Tanzania were established with the introduction of and . The ferry MV ''Bukoba'' sank in the lake on 21 May 1996 with a loss of between 800 and 1,000 lives, making it one of Africa's worst maritime disasters. Another tragedy occurred recently on 20 September 2018 that involved the passagers ferry MV Nyerere from Tanzania that caused the deaths of over 200 people. On 6 November 2022, Lake Victoria was the site of a commercial passenger aircraft crash.
Precision Air Flight 494 Precision Air Flight 494 (PW494) was a scheduled domestic passenger flight within Tanzania, from Julius Nyerere International Airport to Bukoba Airport via Mwanza Airport. On 6 November 2022, the ATR 42–500 plane crashed in Lake Victoria wh ...
an ATR 42–500 carrying 39 passengers and four crew, crashed while approaching
Bukoba Airport Bukoba Airport is a domestic airport located in the city of Bukoba, the capital of the Kagera Region in northwest Tanzania. It is adjacent to Lake Victoria. The Bukoba non-directional beacon (Ident: BK) is located on the field. A new asphalt ...
, resulting in 19 fatalities.


See also

* ''
Darwin's Nightmare ''Darwin's Nightmare'' is a 2004 Austrian-French-Belgian documentary film written and directed by Hubert Sauper, dealing with the environmental and social effects of the fishing industry around Lake Victoria in Tanzania. It premiered at the 2004 V ...
'' *
Kishanda Kishanda is a valley and national reserve for elephants in Tanzania. The reserve is part of the Rumanyika-Karagwe National Park created in 2019 and located in Kagera Region Kagera Region (''Mkoa wa Kagera'' in Swahili) is one of Tanzania's ...


References


External links


Decreasing levels of Lake Victoria Worry East African Countries

New Scientist article
on Uganda's violation of the agreed curve for hydroelectric water flow.
Dams Draining Lake Victoria

''A Naturalist on Lake Victoria, with an Account of Sleeping Sickness and the Tse-tse Fly''
(1920). T.F. Unwin Ltd, London; Biodiversity Archive
Video of Lake Victoria
* Institutions of the East African Community
Lake Victoria Fisheries Organisation
{{DEFAULTSORT:Victoria African Great Lakes Lakes of Kenya Lakes of Tanzania Lakes of Uganda Geography of Kampala Nile Kenya–Uganda border Kenya–Tanzania border Tanzania–Uganda border International lakes of Africa Border tripoints Geography of Kagera Region Geography of Mwanza Region Geography of Mara Region Kisumu County Homa Bay County