Kirkwood Formation
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The Kirkwood Formation is a geological
formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondar ...
found in the Eastern and Western Cape provinces in South Africa. It is one of the four formations found within the Uitenhage Group of the Algoa Basin – its type locality – and in the neighbouring Gamtoos Basin. Outcrops of the Kirkwood are also found along the Worcester-Pletmos, Herbertsdale-Riversdale, Heidelberg-Mossel Bay, and Oudtshoorn-Gamtoos basin lines. At these basins the Kirkwood Formation underlies the Buffelskloof Formation and not the
Sundays River Formation The Sundays River Formation is a geological formation found in the Eastern and Western Cape provinces in South Africa. It is the second youngest of the four formations found within the Uitenhage Group of the Algoa Basin, its type locality, and t ...
.


Geology

The Kirwood Formation was formed along the southern section of South Africa during the break up of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
. Dating it has proven tricky, with some regarding it as a Late Jurassic (~145 Ma) or Early Cretaceous deposit (~135 Ma). However, dating the Kirkwood's abundant volcanic ash layers by U-Pb methods is currently underway. The Kirkwood Formation is composed of sedimentary rocks deposited under
fluvial In geography and geology, fluvial processes are associated with rivers and streams and the deposits and landforms created by them. When the stream or rivers are associated with glaciers, ice sheets, or ice caps, the term glaciofluvial or fluviog ...
conditions at or near sea level, such as variegated
mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from '' shale'' by its lack of fissility (parallel layering).Blatt, H., and R.J. Tracy, 1996, ''Petrology. ...
(an iconic feature), medium-grained lithic
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...
(often being charcoal-rich), and sporadic conglomerates. At its base, the Kirkwood Formation includes a significant deposit of fine-to-medium grained, poorly-sorted (often quartzitic) estuarine sandstone with subordinate dark grey to brow shale layers. This deposit was described as the Swartkops Member. The lower portion of the remainder of the Formation is composed predominantly of siltstones, but includes a smaller sandstone layer (described as part of the Bethelsdorp Member). The upper portion of the Formation is composed predominantly of fine, clay-rich mudstones, with multiple smaller layers of estuarine sandstone. The Kirkwood is the second
formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondar ...
of the Uitenhage Group, positioned between the overlying
Sundays River Formation The Sundays River Formation is a geological formation found in the Eastern and Western Cape provinces in South Africa. It is the second youngest of the four formations found within the Uitenhage Group of the Algoa Basin, its type locality, and t ...
and the underlying
Enon Formation The Enon Formation is a geological formation found in the Eastern and Western Cape provinces in South Africa. It is the lowermost of the four formations found within the Uitenhage Group of the Algoa Basin, its type locality, where it has been me ...
. Only three geological
members Member may refer to: * Military jury, referred to as "Members" in military jargon * Element (mathematics), an object that belongs to a mathematical set * In object-oriented programming, a member of a class ** Field (computer science), entries in ...
have been described within it: the Swartkops Member which contains estuarine sandstone deposits and lacks fossils. The Colchester Member is the most fossil-rich containing terrestrial and lacustrine fossils and is composed of dark grey
shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4) and tiny fragments (silt-sized particles) of other minerals, especial ...
,
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility.Blatt ''et al.'' 1980, p ...
s and sandstones. The third, the Bethelsdorp Member, resembles the Colchester in containing dark-grey shales and sandstone, and contains marine microfossils. The upper section of the member is overlain by mudstone and siltstone.


Paleontology

The Kirkwood Formation is the most fossil-rich formation of Late Jurassic - Early Cretaceous age known in South Africa. It has yielded disarticulated remains of
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
and
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world ...
dinosaurs, and several
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
species in which the Kirkwood is particularly rich, and includes indeterminate members of
Diplodocidae Diplodocids, or members of the family Diplodocidae ("double beams"), are a group of sauropod dinosaurs. The family includes some of the longest creatures ever to walk the Earth, including ''Diplodocus'' and '' Supersaurus'', some of which may hav ...
,
Dicraeosauridae Dicraeosauridae is a family of diplodocoid sauropods who are the sister group to Diplodocidae. Dicraeosaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as ''Amargasaurus'', ''Suuwassea'', ''D ...
and
Brachiosauridae The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the ...
, as well as ''
Algoasaurus ''Algoasaurus'' (; "Algoa Bay reptile") is a genus of sauropod dinosaur from the Berriasian- early Valanginian-age Early Cretaceous Upper Kirkwood Formation of Cape Province, South Africa, specifically near a town called Despatch. Discovery an ...
'', a
eusauropod Eusauropoda (meaning "true sauropods") is a derived clade of sauropod dinosaurs. Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of ''Shunosaurus'', and possibly ''Barapasaurus ...
of uncertain affinities. A partial skull and teeth of a stegosaur, ''
Paranthodon ''Paranthodon'' ( ) is a genus of stegosaurian dinosaur that lived in what is now South Africa during the Early Cretaceous, between 139 and 131 million years ago. Discovered in 1845, it was one of the first stegosaurians found. Its only rem ...
'', has also been discovered."Table 16.1," in Weishampel, et al. (2004). Page 345 In addition to ''Paranthodon'', an
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
vertebra has also been described. Fossils belonging to a
dryosaurid Dryosaurids were primitive iguanodonts. They are known from Middle Jurassic to Early Cretaceous rocks of Africa, Europe, and North America. Phylogeny Until recently many dryosaurids have been regarded as dubious ('' Callovosaurus'' and '' Kan ...
ornithopod were unearthed in three expeditions between 1995 and 1999. In 2022, these remains were described as a new genus and species, '' Iyuku raathi''. ''Iyuku'' is primarily known from remains belonging to hatchling and juvenile individuals. Indeterminate members of
Rhynchocephalia Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a diverse g ...
and
Squamata Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians (worm lizards), which are collectively known as squamates or scaled reptiles. With over 10,900 species, ...
have also been discovered. Mesozoic-aged
amber Amber is fossilized tree resin that has been appreciated for its color and natural beauty since Neolithic times. Much valued from antiquity to the present as a gemstone, amber is made into a variety of decorative objects."Amber" (2004). In Ma ...
has also been recovered from the Kirkwood. The most well-known find from the Kirkwood is ''
Nqwebasaurus ''Nqwebasaurus'' (; anglicized as or ) is a basal coelurosaur and is the basal-most member of the coelurosaurian clade Ornithomimosauria from the Early Cretaceous of South Africa. The name ''Nqwebasaurus'' is derived from the Xhosa word "Nqweb ...
'', a basal ornithomimosaur. Fragmentary remains of various reptile, frog, insect, and mammal fossils have also been found, including fish scales and freshwater
bivalves Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
. In addition several plant species such as
bryophyte The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in ...
s,
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s,
conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...
,
cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...
s, and bennettitaleans have been discovered, including
silicified In geology, petrifaction or petrification () is the process by which organic material becomes a fossil through the replacement of the original material and the filling of the original pore spaces with minerals. Petrified wood typifies this proce ...
fossil tree trunks in the sandstone sections which show evidence of being burned.


References

{{Geology of South Africa, formations Geologic formations of South Africa Lower Cretaceous Series of Africa Cretaceous South Africa Berriasian Stage Hauterivian Stage Valanginian Stage Sandstone formations Shale formations Mudstone formations Siltstone formations Fluvial deposits