''Khaan'' (; from Mongol 'lord') was an

oviraptorid Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between on ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

that was found in the

Djadochta Formation The Djadochta Formation (sometimes transcribed and also known as Djadokhta, Djadokata, or Dzhadokhtskaya) is a highly fossiliferous geological formation situated in Central Asia, Gobi Desert, dating from the Late Cretaceous period, about 75 milli ...

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

and lived in the Late

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

(

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

), 75-71

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

Description

''Khaan'' did not differ much from other oviraptorids. At first, its remains were assigned to " Ingenia", but the ''Khaan'' manual structure, lacking the expansion of the upper third

metacarpal In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ...

, was considered to differ sufficiently from that of "Ingenia" for it to be assigned to its own genus. The oviraptorid diet is disputed, with plants and molluscs having been suggested. Like other oviraptorids, ''Khaan'' was probably at least partially a meat eater, feeding on small

vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...

s like

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...

s and possibly other small dinosaurs. It was also probably feathered.

Discovery

The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

''Khaan mckennai'' was in 2001 named by James M. Clark e.a. The genus name is derived from

Mongol The Mongols ( mn, Монголчууд, , , ; ; russian: Монголы) are an East Asian ethnic group native to Mongolia, Inner Mongolia in China and the Buryatia Republic of the Russian Federation. The Mongols are the principal member of ...

''khaan'', "lord" or "ruler". The specific name honours the

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

Malcolm Carnegie McKenna. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

IGM 100/1127, consists of an almost complete skeleton found together with another specimen, IGM 100/1002. Together they were informally referred to as "Romeo and Juliet". These individuals are about four feet long. A third, considerably larger, specimen, IGM 100/973, was referred to the species.

Classification

''Khaan'' was by Clark assigned to the

Oviraptoridae Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one ...

. Among oviraptorids, it was probably more closely related to ''

Conchoraptor ''Conchoraptor'' (meaning "conch plunderer") is a genus of oviraptorid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 70 million years ago. It is known from the Barun Goyot and Nemegt formations of Mongolia. Discovery Wh ...

''. The

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

below follows an analysis by Fanti ''et al.'', 2012.

Paleobiology

According to a 2014 study published in

Nature Nature, in the broadest sense, is the physics, physical world or universe. "Nature" can refer to the phenomenon, phenomena of the physical world, and also to life in general. The study of nature is a large, if not the only, part of science. ...

, ''Khaan'' was possibly

sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...

. Two specimens, the holotype MPC-D 100/1127 and referred specimen MPC-D 100/1002, were analyzed, the dimorphic feature being in the anterior

chevrons Chevron (often relating to V-shaped patterns) may refer to: Science and technology * Chevron (aerospace), sawtooth patterns on some jet engines * Chevron (anatomy), a bone * '' Eulithis testata'', a moth * Chevron (geology), a fold in rock l ...

. Both specimens were of the same size and build, and thus were likely the same age, ruling out

ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...

. In MPC-D 100/1127, the anterior chevrons showed great similarity to those of other theropods, with no great expansion on the distal end. However, in the other individual, the chevrons had a "heel"-like expansion above the distal end, which increased in size along the sequence. The study ruled out a possibly pathologic explanation, finding sexual dimorphism to have more support. It was thought that the reduced spines were a female characteristic, because they would increase space for laying eggs. Also, the larger spines could be for male muscle attachments, which would support a tail fan. A 2022 study of the bite force of ''Khaan'' and comparisons with other oviraptorosaurs such as ''

Incisivosaurus ''Incisivosaurus'' ("incisor lizard") is a genus of small, probably herbivorous theropod dinosaurs from the early Cretaceous Period of what is now the People's Republic of China. The first specimen to be described (by Xu ''et al.'' in 2002), IVPP ...

'', ''

Citipati ''Citipati'' (; meaning "funeral pyre lord") is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality at the Djadokhta F ...

'', and ''

'' suggests that ''Khaan'' had a very strong bite force. The moderate jaw gape seen in oviraptorosaurs is indicative of herbivory in the majority of the group, but it is clear they were likely feeding on much tougher or more types of vegetation than other herbivorous theropods in their environment, such as ornithomimosaurs and therizinosaurs were able to. The examinations suggest oviraptorosaurs may have been powerful-biting generalists or specialists that partook of niche partitioning both in body size and cranial function.

References

{{Taxonbar, from=Q135295 Late Cretaceous dinosaurs of Asia Djadochta fauna Oviraptorids Fossil taxa described in 2001 Taxa named by Mark Norell Taxa named by Rinchen Barsbold

Description

Discovery

Classification

Paleobiology

See also

References