Kansajsuchus
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''Kansajsuchus'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of paralligatorid
mesoeucrocodylia Mesoeucrocodylia is the clade that includes Eusuchia and crocodyliforms formerly placed in the paraphyletic group Mesosuchia. The group appeared during the Early Jurassic, and continues to the present day. Diagnosis It was long known that M ...
n. It is based on
PIN A pin is a device used for fastening objects or material together. Pin or PIN may also refer to: Computers and technology * Personal identification number (PIN), to access a secured system ** PIN pad, a PIN entry device * PIN, a former Dutch de ...
 2399/301, a right
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
, one of the bones of the tip of the snout. This specimen was found in rocks of the lower
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The ...
-age
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Yalovach Svita of
Kansai The or the , lies in the southern-central region of Japan's main island Honshu, Honshū. The region includes the Prefectures of Japan, prefectures of Nara Prefecture, Nara, Wakayama Prefecture, Wakayama, Kyoto Prefecture, Kyoto, Osaka Prefectur ...
, in the
Fergana Basin The Fergana Valley (; ; ) in Central Asia lies mainly in eastern Uzbekistan, but also extends into southern Kyrgyzstan and northern Tajikistan. Divided into three republics of the former Soviet Union, the valley is ethnically diverse and in the ...
of
Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...
. Additional fossils including
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e and bony armor have been assigned to this genus. It would have been a large animal, estimated at up to long. ''Kansajsuchus'' was described in 1975 by Mikhail Efimov. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Kansajsuchus extensus''. Halliday ''et al.'' (2013) confirmed the validity of the species ''K. extensus'', and its
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
position among other basal goniopholidids from Asia. '' "Sunosuchus" shartegensis'' was found to represent its
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
, and both species were placed in a distinct
lineage Lineage may refer to: Science * Lineage (anthropology), a group that can demonstrate its common descent from an apical ancestor or a direct line of descent from an ancestor * Lineage (evolution), a temporal sequence of individuals, populati ...
from the type species of ''Sunosuchus'', '' S. miaoi'', however with a weak support. Therefore, the authors raised the suggestion that later revisions and phylogenetic analyses would result in the abandonment of the name ''Kansajsuchus'', and a referral of its type species to ''Sunosuchus''. However, a subsequent paper by Kuzmin et al. (2019) showed that ''Kansajsuchus'' is a member of the family
Paralligatoridae Paralligatoridae is an extinct family of neosuchian crocodyliforms that existed during the Jurassic and Cretaceous periods. It includes the genera ''Paralligator'', '' Brillanceausuchus'', '' Kansajsuchus'', ''Shamosuchus'', ''Scolomastax'', ''Sa ...
.


References

Late Cretaceous crocodylomorphs of Asia Turonian life Fossils of Uzbekistan Bissekty Formation Fossil taxa described in 1975 Prehistoric pseudosuchian genera {{paleo-archosaur-stub