''Kangnasaurus'' (meaning "Farm Kangnas lizard") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

iguanodont Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', '' Tenontosaurus'', and the hadrosaurids or "duck-bil ...

ian

ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous wo ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

found in supposedly Early Cretaceous rocks of

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...

. It is known from a tooth and possibly some

postcranial Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated s ...

remains found in the early- Aptian Kalahari Deposits Formation. It was probably similar to ''

Dryosaurus ''Dryosaurus'' ( , meaning 'tree lizard', Greek ' () meaning 'tree, oak' and () meaning 'lizard'; the name reflects the forested habitat, not a vague oak-leaf shape of its cheek teeth as is sometimes assumed) is a genus of an ornithopod dinosaur ...

''.

Discovery and naming

''Kangnasaurus'' was named in 1915 by

Sidney H. Haughton Sidney Henry Haughton FRS (7 May 1888 – 24 May 1982) was an English-born South African paleontologist and geologist best known for his description of the sauropodomorph dinosaur '' Melanorosaurus'' in 1924, and his work on the geology of ...

. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...

is ''Kangnasaurus coetzeei''. The generic name refers to the Kangnas farm; the specific name to the farmer, Coetzee. ''Kangnasaurus'' is based on

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...

SAM 2732, a tooth found at a depth of 34 metres in a well at Farm Kangnas, in the

Orange River The Orange River (from Afrikaans/Dutch: ''Oranjerivier'') is a river in Southern Africa. It is the longest river in South Africa. With a total length of , the Orange River Basin extends from Lesotho into South Africa and Namibia to the north ...

valley of northern

Cape Province The Province of the Cape of Good Hope ( af, Provinsie Kaap die Goeie Hoop), commonly referred to as the Cape Province ( af, Kaapprovinsie) and colloquially as The Cape ( af, Die Kaap), was a province in the Union of South Africa and subsequen ...

. The age of these rocks, conglomerates in an ancient

crater lake Crater Lake ( Klamath: ''Giiwas'') is a volcanic crater lake in south-central Oregon in the western United States. It is the main feature of Crater Lake National Park and is famous for its deep blue color and water clarity. The lake partly fill ...

, is unclear; they are thought to be from the Early Cretaceous (probably early- Aptian). Haughton thought SAM 2732 was a tooth from the

upper jaw The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

, but Michael Cooper reidentified it as a

lower jaw In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

tooth in 1985. This had implications for its classification: Haughton thought the tooth was that of an

iguanodontid Iguanodontidae is a family of iguanodontians belonging to Styracosterna, a derived clade within Ankylopollexia. Characterized by their elongated maxillae, they were herbivorous and typically large in size. This family exhibited locomotive dynam ...

, while Cooper identified it as from an animal more like ''

'', a more basal ornithopod. Haughton described several other fossils as possibly belonging to ''Kangnasaurus''. These include five partial thigh bones, a partial thigh bone and

shin bone The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...

, a partial

metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the me ...

, a partial shin and foot,

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...

e, and unidentified bones. Some of the bones apparently came from other deposits, and Haughton was not certain that they all belonged to his new genus. Cooper was also not certain, but described the other specimens as if they did belong to ''Kangnasaurus''.

Classification

''Kangnasaurus'' is usually regarded as dubious, although a 2007 review of dryosaurids by Ruiz-Omeñaca and colleagues retained it as potentially valid, differing from other dryosaurids by details of the thigh bone. Like other basal iguanodontians, it would have been a

bipedal Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...

herbivore. At least two recent studies have found it to be an elasmarian instead of a dryosaurid.

References

{{Taxonbar, from=Q633747 Iguanodonts Aptian life Early Cretaceous dinosaurs of Africa Cretaceous South Africa Fossils of South Africa Fossil taxa described in 1915 Taxa named by Sidney H. Haughton Ornithischian genera