''Jushatyria'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s have been found in the Koltaevo III Locality, district of

Kumertau Kumertau (russian: Кумерта́у; ba, Күмертау) is a town in the Republic of Bashkortostan, Russia, located from Ufa and from Sterlitamak. Population: Administrative and municipal status Within the framework of administrative d ...

near the

Ural Mountains The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western ...

European Russia European Russia (russian: Европейская Россия, russian: европейская часть России, label=none) is the western and most populated part of Russia. It is geographically situated in Europe, as opposed to the cou ...

from the Bukobay Gorizont. The locality dates back to the

Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). ...

stage of the

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and ...

. Additional material has been described from a locality on the banks of the

Berdyanka River Berdyanka or Berdianka may refer to: *Berdyanka River (:ru:Бердянка (приток Урала), Бердянка), Orenburg Oblast, Russia *Berdyanka Tract, part of Berdianskyi State Reserve, Berdyanskyi State Reserve (:uk:Бердянськ ...

that was previously assigned to a rauisuchid-like archosaur. However, this material differed from the original specimens because it lacked slit-like antorbital openings accompanying the

antorbital fossa An antorbital fenestra (plural: fenestrae) is an opening in the skull that is in front of the eye sockets. This skull character is largely associated with archosauriforms, first appearing during the Triassic Period. Among extant archosaurs, birds ...

. Nesbitt (2009) and Gower and Sennikov (2000) suggested that all material currently referred to ''Jushatyria'' most likely does not represent a single taxon.Gower, D. J. and Sennikov, A. G. (2000). Early Archosaurs from Russia ''In:'' Benton, M. J., Kurochkin, E. N., Shishkin, M. A. and Unwin, D. M., eds., ''The Age of Dinosaurs in Russia and Mongolia''. Cambridge, Cambridge University Press; pp. 140–159. Thus, ''Jushatyria'' is known only from its

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

PIN 2867/5, an incomplete left

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

. As the maxilla is damaged, many " rauisuchian" characters could not be verified. ''Jushatyria'' was reassigned as an indeterminate archosaur on the basis of the presence of an

on the lateral surface of the maxilla.

References

External links

''Jushatyria''
in the

Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) originated in the NCEAS-funded Phanerozoic Marine Pale ...

Rauisuchids Extinct animals of Europe Prehistoric pseudosuchian genera {{triassic-reptile-stub