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Julius von Sachs (; 2 October 1832 – 29 May 1897) was a German
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
from Breslau,
Prussian Silesia The Province of Silesia (german: Provinz Schlesien; pl, Prowincja Śląska; szl, Prowincyjŏ Ślōnskŏ) was a province of Prussia from 1815 to 1919. The Silesia region was part of the Prussian realm since 1740 and established as an official p ...
. He is considered the founder of experimental plant physiology and co-founder of modern water culture. Julius von Sachs and
Wilhelm Knop Wilhelm may refer to: People and fictional characters * William Charles John Pitcher, costume designer known professionally as "Wilhelm" * Wilhelm (name), a list of people and fictional characters with the given name or surname Other uses * Moun ...
are monumental figures in the
history of botany The history of botany examines the human effort to understand life on Earth by tracing the historical development of the discipline of botany—that part of natural science dealing with organisms traditionally treated as plants. Rudimentary b ...
by first demonstrating the importance of water culture for the study of
plant nutrition Plant nutrition is the study of the chemical elements and compounds necessary for plant growth and reproduction, plant metabolism and their external supply. In its absence the plant is unable to complete a normal life cycle, or that the element i ...
and plant physiology in the
19th century The 19th (nineteenth) century began on 1 January 1801 ( MDCCCI), and ended on 31 December 1900 ( MCM). The 19th century was the ninth century of the 2nd millennium. The 19th century was characterized by vast social upheaval. Slavery was abolis ...
.


Early life

Sachs was born at Breslau on 2 October 1832. His father, Graveur Sachs, was an engraver by trade, and father taught son delineation and accuracy of line and color. From earliest boyhood Julius was fascinated with plants, making collections of them on many field excursions with his father. He gave much of his time between the ages of thirteen and sixteen to drawing and painting the flowers, fungi, and other specimens which he collected. At the Gymnasium from 1845 to 1850 he was most interested in the natural sciences, collecting skulls, writing a monograph on the crayfish. His natural science teacher, one Krober, showed a singular lack of foresight when he solemnly warned young Sachs against devoting himself to the natural sciences. When he was sixteen years old, his father died, and in the next year both his mother and a brother died of cholera. Suddenly without financial support, he was fortunate to be taken into the family of
Jan Evangelista Purkyně Jan Evangelista Purkyně (; also written Johann Evangelist Purkinje) (17 or 18 December 1787 – 28 July 1869) was a Czech anatomist and physiologist. In 1839, he coined the term '' protoplasm'' for the fluid substance of a cell. He was one of ...
who had accepted a professorship at the University of Prague. Sachs was admitted to the university in 1851. Sachs famously labored long hours in the laboratory for Purkyně, and then long hours for himself each day after his work in the laboratory was finished. After the lab, he could devote himself entirely to establishing how plants grow.


Career

In 1856 Sachs graduated as doctor of philosophy, and then adopted a botanical career, establishing himself as ''
Privatdozent ''Privatdozent'' (for men) or ''Privatdozentin'' (for women), abbreviated PD, P.D. or Priv.-Doz., is an academic title conferred at some European universities, especially in German-speaking countries, to someone who holds certain formal qualific ...
'' for plant physiology. In 1859 he was appointed physiological assistant to the Agricultural Academy of Tharandt (now part of the
Technical University of Dresden TU Dresden (for german: Technische Universität Dresden, abbreviated as TUD and often wrongly translated as "Dresden University of Technology") is a public research university, the largest institute of higher education in the city of Dresden, th ...
) at
Julius Adolph Stöckhardt Julius Adolph Stöckhardt (4 January 1809 – 1 June 1886) was a German agricultural chemist. He is mostly recognized for his work on fertilizers, fume damage of plants and his book ''Die Schule der Chemie'' (''School of Chemistry''), which was ...
; and in 1862 he was called to be director of the Polytechnic at Chemnitz, but was almost immediately transferred to the Agricultural Academy at Poppelsdorf (now part of the
University of Bonn The Rhenish Friedrich Wilhelm University of Bonn (german: Rheinische Friedrich-Wilhelms-Universität Bonn) is a public research university located in Bonn, North Rhine-Westphalia, Germany. It was founded in its present form as the ( en, Rhine U ...
), where he remained until 1867, when he was nominated professor of botany in the
University of Freiburg The University of Freiburg (colloquially german: Uni Freiburg), officially the Albert Ludwig University of Freiburg (german: Albert-Ludwigs-Universität Freiburg), is a public university, public research university located in Freiburg im Breisg ...
. In 1868 he accepted the chair of botany in the
University of Würzburg The Julius Maximilian University of Würzburg (also referred to as the University of Würzburg, in German ''Julius-Maximilians-Universität Würzburg'') is a public research university in Würzburg, Germany. The University of Würzburg is one of ...
, which he continued to occupy (in spite of calls from more prestigious German universities) until his death. Sachs achieved distinction as an investigator, a writer and a teacher; his name will ever be especially associated with the great development of plant physiology which marked the latter half of the 19th century, though there is scarcely a branch of botany to which he did not materially contribute. His earlier papers, scattered through the volumes of botanical journals and of the publications of learned societies (a collected edition was published in 1892–93), are of great and varied interest. Prominent among them is the series of "Keimungsgeschichten," which laid the foundation of our knowledge of microchemical methods, as also of the morphological and physiological details of
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
. Then there is his resuscitation of the method of "water-culture," and the application of it to the investigation of the problems of nutrition. Most important are his experiments, developing the concept of
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
, that the
starch Starch or amylum is a polymeric carbohydrate consisting of numerous glucose units joined by glycosidic bonds. This polysaccharide is produced by most green plants for energy storage. Worldwide, it is the most common carbohydrate in human diets ...
-grains, found in leaf
chloroplasts A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant cell, plant and algae, algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, ...
, depend on sunlight. A leaf that has been in sunlight, then bleached white and stained with
iodine Iodine is a chemical element with the symbol I and atomic number 53. The heaviest of the stable halogens, it exists as a semi-lustrous, non-metallic solid at standard conditions that melts to form a deep violet liquid at , and boils to a vi ...
turns black, proving its starch content, whereas a leaf from the same plant that has been out of the sun will remain white. A demonstration of this experiment is shown in the second episode of
BBC Four BBC Four is a British free-to-air public broadcast television channel owned and operated by the BBC. It was launched on 2 March 2002
's "Botany: A Blooming History" presented by Timothy Walker. Sachs's later papers were almost exclusively published in the three volumes of the ''Arbeiten des botanischen Instituts in Würzburg'' (1871–88). Among these are his investigation of the periodicity of growth in length, in connection with which he devised the self-registering
auxanometer An auxanometer (Gr. '= "to grow" + ''metron''= "measure") is an apparatus for measuring increase of growth in plants. In case of an arc-auxanometer (see picture), there is a thin cord fixed to the plant apex on one end and a dead-weight on the oth ...
, by which he established the retarding influence of the highly refrangible rays of the
spectrum A spectrum (plural ''spectra'' or ''spectrums'') is a condition that is not limited to a specific set of values but can vary, without gaps, across a continuum. The word was first used scientifically in optics to describe the rainbow of colors i ...
on the rate of growth; his research on
heliotropism Heliotropism, a form of tropism, is the diurnal or seasonal motion of plant parts (flowers or leaves) in response to the direction of the Sun. The habit of some plants to move in the direction of the Sun, a form of tropism, was already known by t ...
and
geotropism Gravitropism (also known as geotropism) is a coordinated process of differential growth by a plant in response to gravity pulling on it. It also occurs in fungi. Gravity can be either "artificial gravity" or natural gravity. It is a general featu ...
, in which he introduced the
clinostat A clinostat is a device which uses rotation to negate the effects of gravitational pull on plant growth (gravitropism) and development ( gravimorphism). It has also been used to study the effects of microgravity on cell cultures, animal embryo ...
; his work on the structure and the arrangement of cells in growing-points; the elaborate experimental evidence upon which he based his "imbibition-theory" of the transpiration-current; his exhaustive study of the assimilatory activity of the green
leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
; and other papers of interest. Sachs' first published volume was the ''Handbuch der Experimentalphysiologie des Pflanzen'' (1865; French edition, 1868), which gives an admirable account of the state of knowledge in certain departments of the subject, and includes a great deal of original information. This was followed in 1868 by the first edition of his famous ''Lehrbuch der Botanik''. It is a comprehensive work, giving a summary of the botanical science of the period, enriched with the results of many original investigations. The third edition was translated into French by
Philippe Édouard Léon Van Tieghem Philippe Édouard Léon Van Tieghem (; 19 April 1839 – 28 April 1914) was a French botanist born in Bailleul, Nord, Baillleul in the département of Nord (French department), Nord. He was one of the best known French botanists of the latter nin ...
in 1873, and into English by Alfred Bennett in 1875, and published by Oxford University Press. The fourth and last German edition was published in 1874, and also issued by Oxford in 1882. The ''Lehrbuch'' was eventually superseded by the ''Vorlesungen uber Pflanzenphysiologie'' (1st ed., 1882; 2nd ed., 1887; Eng. ed., Oxford, 1887), a work more limited in scope, but covering more ground than its title would imply; it has not gained the general recognition accorded to the ''Lehrbuch''. Finally, there is the ''Geschichte der Botanik'' (1875); an account of the development of the various branches of botanical science from the middle of the 16th century up to 1860, of which an English edition was published in 1890 by the Oxford Press. A full account of Sachs' life and work was given by Professor Goebel, formerly his assistant, in ''Flora'' (1897), of which an English translation appeared in ''Science Progress'' for 1898. There is also an obituary notice of him in the ''Proc. Roy. Soc.'' vol. lxii. In 1885 he became foreign member of the
Royal Netherlands Academy of Arts and Sciences The Royal Netherlands Academy of Arts and Sciences ( nl, Koninklijke Nederlandse Akademie van Wetenschappen, abbreviated: KNAW) is an organization dedicated to the advancement of science and literature in the Netherlands. The academy is housed ...
.


Evolution

Sachs has been described as a "post-Darwinian botanist" who "integrated the evolutionary theory into his morphological writings." He was originally supportive of
Darwinism Darwinism is a scientific theory, theory of Biology, biological evolution developed by the English naturalist Charles Darwin (1809–1882) and others, stating that all species of organisms arise and develop through the natural selection of smal ...
but in his late career became bitterly opposed to it, instead preferring
non-Darwinian evolution Alternatives to Darwinian evolution have been proposed by scholars investigating biology to explain signs of evolution and the relatedness of different groups of living things. The alternatives in question do not deny that evolutionary changes ove ...
.


Influence

Many pupils of Sachs like
Julius Oscar Brefeld Julius Oscar Brefeld (19 August 1839 – 12 January 1925), usually just Oscar Brefeld, was a German botanist and mycologist. Biography Brefeld was a native of Telgte. He studied pharmacy in Heidelberg and Berlin, and afterwards served as an a ...
, Francis Darwin,
Karl Ritter von Goebel Karl Immanuel Eberhard Ritter von Goebel FRS FRSE (8 March 1855, Billigheim, Baden – 9 October 1932, Munich) was a German botanist. His main fields of study were comparative functional anatomy, morphology, and the developmental physiology of p ...
, Georg Albrecht Klebs, Spiridon Miliarakis, Hermann Müller-Thurgau,
Fritz Noll Fritz Noll (27 August 1858 in Frankfurt am Main – 19 June 1908) was a German botanist who made contributions in the field of plant physiology. He studied natural history and sciences at the Universities of Würzburg, Marburg and Heidelbe ...
,
Wilhelm Pfeffer Wilhelm Friedrich Philipp Pfeffer (9 March 1845 – 31 January 1920) was a German botanist and plant physiologist born in Grebenstein. Academic career He studied chemistry and pharmacy at the University of Göttingen, where his instructors incl ...
, Karl Prantl,
Christian Ernst Stahl Christian Ernst Stahl (21 June 1848 – 3 December 1919) was a German botanist who was a native of Schiltigheim, Alsace. Academic career He studied botany at the University of Strasbourg with Pierre-Marie-Alexis Millardet (1838-1902), and at ...
and
Hugo de Vries Hugo Marie de Vries () (16 February 1848 – 21 May 1935) was a Dutch botanist and one of the first geneticists. He is known chiefly for suggesting the concept of genes, rediscovering the laws of heredity in the 1890s while apparently unaware o ...
became later famous botanists. His scientific statements also influenced
Dennis Robert Hoagland Dennis Robert Hoagland (April 2, 1884 – September 5, 1949) was an American chemist and plant scientist working in the fields of plant nutrition, agricultural chemistry, and physiology. He was Professor of Plant Nutrition at the University o ...
and Daniel Israel Arnon, who followed one of Sachs' principles in developing nutrient solutions, by quoting him as follows: "I mention the quantities (of chemicals) I am accustomed to use generally in water cultures, with the remark, however, that a somewhat wide margin may be permitted with respect to the quantities of the individual salts and the concentration of the whole solution - it does not matter if a little more or less of the one or the other salt is taken - if only the nutritive mixture is kept within certain limits as to quality and quantity, which are established by experience." Sachs' quantity and quality principle and Knop's four-salt mixture contributed significantly to the development of the later
Hoagland solution The Hoagland solution is a hydroponic nutrient solution that was newly developed by Hoagland and Snyder in 1933, modified by Hoagland and Arnon in 1938, and revised by Arnon in 1950. It is one of the most popular artificial solution compositions ...
. The standard botanical author abbreviation Sachs is applied to
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
he described. In 1866, botanist
Griseb. August Heinrich Rudolf Grisebach () was a German botany, botanist and phytogeography, phytogeographer. He was born in Hannover on 17 April 1814 and died in Göttingen on 9 May 1879. Biography Grisebach studied at the Lyceum in Hanover, the clo ...
published '' Sachsia'', which is a genus of
West Indian A West Indian is a native or inhabitant of the West Indies (the Antilles and the Lucayan Archipelago). For more than 100 years the words ''West Indian'' specifically described natives of the West Indies, but by 1661 Europeans had begun to use it ...
and Floridian plants in the elecampane tribe. Then in 1895, botanist
Paul Lindner Paul Lindner (1861 – 1945) was a German chemist and microbiologist, best known for discovering the fission yeast ''Schizosaccharomyces pombe''. Biography and career Lindner was born in 1861 in Giesmannsdorf near the Neisse River, part of Up ...
published '' Sachsia'', which is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of fungi in the
Ascomycota Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The def ...
phylum and
Saccharomycetales Saccharomycetales belongs to the kingdom of Fungi and the division Ascomycota. It is the only order in the class Saccharomycetes. There are currently 13 families recognized as belonging to Saccharomycetales. GBIF also includes; Alloascoideacea ...
order. Both were named in honour of Julius von Sachs.


Publications

* 1859: Physiologische Untersuchungen über die Keimung der Schmikbohne (Phaseolus multiflorus) * 1859: Ueber das abwechselnde Erbleichen und Dunkelwerden der Blätter bei wechselnder Beleuchtung * 1862: Ueber das Vergeilen der Pflanzen * 1863: Ueber den Einfluss des Tageslichtes auf die Neublidung unt Entfaltung verschiedener Pflanzenorgane * 1865: Handbuch der Experimentalphysiologie der Pflanzen * 1868: Lehrbuch der Botanik, 3rd ed. 1873
4th ed. 1874
* 1875: Die Geschichte der Botanik vom 16. Jahrhundert bis 1860
Digital edition from 1875
by the
University and State Library Düsseldorf The University and State Library Düsseldorf (german: Universitäts- und Landesbibliothek Düsseldorf, abbreviated ULB Düsseldorf) is a central service institution of Heinrich Heine University. Along with Bonn and Münster, it is also one of th ...
) * 1878: Ueber die Anordnung der Zellen in jüngsten Pflanzentheilen * 1882: Die Vorlesungen über Pflanzenphysiologie * 1892: Gesammelte Abhandlungen über Pflanzenphysiologie * 1894: Mechanomorphosen und Phylogenie * 1896: Phylogenetische Aphorismen und über innere Gestaltungsursachen oder Automorphosen


References


Other sources

* * *


Bibliography

* ** , see also


External links

* {{DEFAULTSORT:Sachs, Julius von 1832 births 1897 deaths Plant physiologists German scientists People from Wrocław German untitled nobility Foreign associates of the National Academy of Sciences Foreign Members of the Royal Society Members of the Royal Netherlands Academy of Arts and Sciences Charles University alumni Academic staff of Charles University Academic staff of the University of Freiburg Academic staff of the University of Würzburg Academic staff of the University of Bonn Non-Darwinian evolution Musicians from Wrocław People from the Province of Silesia Botanists with author abbreviations 19th-century German botanists Scientists from Würzburg