The Ichthyosauromorpha are an extinct

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

marine reptile Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. The earliest marine reptile mesosaurus (not to be confused with mosasaurus), arose in the Permian period during the ...

s consisting of the

Ichthyosauriformes The Ichthyosauriformes are a group of marine reptiles, belonging to the Ichthyosauromorpha, that lived during the Mesozoic. The stem clade Ichthyosauriformes was in 2014 defined by Ryosuke Motani and colleagues as the group consisting of all ...

and the

Hupehsuchia Hupehsuchia is an order of diapsid reptiles closely related to ichthyosaurs. The group was short-lasting, with a temporal range restricted to the late Olenekian age, spanning only a few million years of the Early Triassic. The order gets its na ...

, living during the Mesozoic. The node clade Ichthyosauromorpha was first defined by Ryosuke Motani ''et al.'' in 2014 as the group consisting of the last common ancestor of ''

Ichthyosaurus communis ''Ichthyosaurus'' (derived from Greek ' () meaning 'fish' and ' () meaning 'lizard') is a genus of ichthyosaurs from the Early Jurassic (Hettangian - Pliensbachian), with possible Late Triassic record, from Europe (Belgium, England, Germany, Swi ...

'' and ''

Hupehsuchus nanchangensis ''Hupehsuchus'' is an extinct genus of small marine reptiles, about 1 m (3 ft) long, found in the area of Hubei in China. This marine reptile lived in the Olenekian stage of the Early Triassic period. Description ''Hupehsuchus'' w ...

'', and all its descendants. Their

synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...

, unique derived traits, include: the presence of an anterior flange on the

humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...

and radius; the lower end of the ulna being as wide as or wider than the upper end, the forelimb being as long as or longer than the hindlimb, the hand having at least three quarters of the length of the upper arm and lower arm combined, the fibula extending behind the level of the thighbone, and the transverse process of the vertebral neural arch being reduced or absent. The Ichthyosauromorpha probably originated in China during the upper

Lower Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a un ...

period, about 248 million years ago. One branch consisted of the Hupehsuchia, and the other of the Ichthyosauriformes, of which ''

Cartorhynchus ''Cartorhynchus'' (meaning "shortened snout") is an extinct genus of early ichthyosauriform marine reptile that lived during the Early Triassic epoch, about 248 million years ago. The genus contains a single species, ''Cartorhynchus lenticarp ...

'' was a basal member. Other ichthyosauriforms were the Ichthyopterygia, containing the Ichthyosauria and allies. The last ichthyosaurs probably became extinct in the middle Cretaceous. Their relationships with other reptiles are unresolved, due to their highly

derived Derive may refer to: * Derive (computer algebra system), a commercial system made by Texas Instruments * ''Dérive'' (magazine), an Austrian science magazine on urbanism *Dérive, a psychogeographical concept See also * *Derivation (disambiguatio ...

morphology, even in their earliest known representatives, though they are usually considered to be

diapsids Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago ...

Taxonomy

* Clade Ichthyosauromorpha ** Order

** Clade

*** Clade

Nasorostra The Ichthyosauriformes are a group of marine reptiles, belonging to the Ichthyosauromorpha, that lived during the Mesozoic. The stem clade Ichthyosauriformes was in 2014 defined by Ryosuke Motani and colleagues as the group consisting of all ...

*** Superorder Ichthyopterygia

Phylogeny

The internal phylogenetic structure of the Ichthyosauromorpha is shown by this cladogram:

References

{{Taxonbar, from=Q18534275 Olenekian first appearances Turonian extinctions