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''Iris stolonifera'' is a plant species in the genus ''
Iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants * Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional ent ...
''; it is also in the subgenus ''
Iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants * Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional ent ...
'', and in the section ''Regelia''. It is a
rhizomatous In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ho ...
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
, from the mountains of
Turkestan Turkestan, also spelled Turkistan ( fa, ترکستان, Torkestân, lit=Land of the Turks), is a historical region in Central Asia corresponding to the regions of Transoxiana and Xinjiang. Overview Known as Turan to the Persians, western Turke ...
, between
Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...
,
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
and
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bordere ...
. It has red-skinned stolon roots and rhizomes, glaucous, long, blue-grey leaves, and bi-coloured flowers, in various shades from milky white, to blue, purple, pale lilac, lavender and brown. It normally has blue to yellow beards on all the petals.


Description

It has a rhizome, that in the spring, sends out thin, and long, (up to long,British Iris Society (1997) ) secondary roots (or
stolons In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...
), which have a red skin. At the end of each stolon, it forms a new rhizome, creating widespread colonies of plants. Other 'Regelia section' irises also have stolons. Also '' Iris japonica'', '' Iris prismatica'' and '' Iris henryi'' produce stolons. It has dark blue-green,James Cullen, Sabina G. Knees, H. Suzanne Cubey (Editors) or glaucous leaves. They are sword-shaped, and long, and 0.8 cm to 2 cm wide. They are prominently veined, and semi-evergreen, disappearing after summer, after the blooming period is over. It has a stem, that can grow up to between tall, or tall. The stem has (scarious) membranous, spathes or
bracts In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
(leaves of the flower bud), which are long. The stems hold 2–3 terminal (top of stem) flowers, blooming in late spring, between April and June, or May, or between May and June.Basak Gardner & Chris Gardner The scented, flowers are in diameter. They are variable in colour, normally bi-coloured, ranging from milky white, to blue, to purple, (or pale lilac, or lavender,) and brown. They normally have a blue (or pale lilac,) centre with a brown, or reddish-bronze, or yellow margin. It has two pairs of petals, three large
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
(outer petals), known as the 'falls' and three inner, smaller petals (or
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
), known as the 'standards'. The falls are long, and wide. The standards are long, and wide. Each petal has a beard. Which is also variable, normally blue, or yellow, or between blue and yellow, Also lilac, and cream beards have been found. It has a brown, 2.5 cm long perianth tube, which is funnel shaped. It has 3 cm long styles, which are variable in colour. It has cream or blue coloured
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
. After the iris has flowered, between May and July, it produces an oblong seed capsule, that is long. It is slightly inflated, and tapers at both ends. Inside the capsule, are light brown seeds that have thick white
aril An aril (pronounced ), also called an arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode or false aril is sometimes distinguished: whereas an aril grows from the attachment point of the see ...
(coatings).


Biochemistry

As most irises are
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
, having two sets of
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
s, this can be used to identify hybrids and classification of groupings. It has a chromosome count: 2n=44.


Taxonomy

It has the common names of 'Pamiriris'. It is known as 'Pobegonosy Iris' in Russia. The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''stolonifera'' derives from its thin stolon-like roots. (See description for more details.) It was first collected by
Olga Fedtschenko Olga Aleksandrovna Fedchenko or Fedtschenko ( Armfeld; 30 October 1845 – 24 April 1921) was a Russian botanist. '' Rosa fedtschenkoana'', which is found in Asia, was named in her honour (with a German transliteration of her name). Early life O ...
in around 1870, It was first published and described by
Karl Maximovich Carl Johann Maximovich (also Karl Ivanovich Maximovich, Russian: Карл Иванович Максимович; 23 November 1827 in Tula, Russia – 16 February 1891 in Saint Petersburg) was a Russian botanist. Maximovich spent most of his life ...
in the 'Bull. Acad. Imp. Sci. Saint-Pétersbourg' Vol.26 on page 535 in 1880. It was also published in 'Mélanges Biol. Bull. Phys.-Math. Acad. Imp. Sci. Saint-Pétersbourg' (Diagn. pl. nov. asiat. or 'Diagnoses Plantarum Asiaticarum') Vol. 10 on page 731 in 1880. It was also published with an illustration in ''
Curtis's Botanical Magazine ''The Botanical Magazine; or Flower-Garden Displayed'', is an illustrated publication which began in 1787. The longest running botanical magazine, it is widely referred to by the subsequent name ''Curtis's Botanical Magazine''. Each of the issue ...
'' 7861 in 1902, and in ''Hort.'' Vol. 7 Issue 8 page 191 on 15 April 1929. ''Iris stolonifera'' is an accepted name by the RHS, and it was verified by
United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
and the
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
on 2 October 2014.


Distribution and habitat

''Iris stolonifera'' is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
areas of central
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
.


Range

It is found in the mountain ranges of the Pamir Alai (or Alay), (including the
Gissar Range Hisar Range ( tg, Қаторкӯҳи Ҳисор; uz, Hisor tizmasi, russian: Гиссарский хребет; uk, Ґаторкугі Гісор; fa, رشته‌کوه حصار; also known as Hissar, Hisor, or Gissar Range) is a mountain range ...
, and Saravschan Range,). which passes through the former Russian (or Soviet), country of Turkestan. (including near
Bokhara Bukhara ( Uzbek: /, ; tg, Бухоро, ) is the seventh-largest city in Uzbekistan, with a population of 280,187 , and the capital of Bukhara Region. People have inhabited the region around Bukhara for at least five millennia, and the city h ...
,) It is also found in Tajikistan, (from the city of
Dushanbe Dushanbe ( tg, Душанбе, ; ; russian: Душанбе) is the capital and largest city of Tajikistan. , Dushanbe had a population of 863,400 and that population was largely Tajik. Until 1929, the city was known in Russian as Dyushambe (r ...
, to Uzbekistan, (including near the city of
Samarkand fa, سمرقند , native_name_lang = , settlement_type = City , image_skyline = , image_caption = Clockwise from the top:Registan square, Shah-i-Zinda necropolis, Bibi-Khanym Mosque, view inside Shah-i-Zinda, ...
near the Baysun ridge, and in the 'Kitab Zapovednik' (Geoological) reserve in the Kashkadarya Province,Gustave Gintzburge ) and in Afghanistan.


Habitat

It grows on dry rock slopes, along dry creeks, and upland meadows. One reference (Flora of S.S.S.R.) mentions wet meadows and stream sides, also mentioned by the British Iris Society Guide, but herbarium specimens do not support this habitat, also the other irises in the Regelia section are also mountainside irises. They can be found at an altitude of above sea level.


Cultivation

It is very cold hardy, to
USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
Zone 5, or between Zone 5 to Zone 8. It is also hardy in Europe to Zone H3. In very cold areas, such as Russia, the plant may die out after 2 or 3 years. It prefers to grow in well-drained soils, which are rich in
humus In classical soil science, humus is the dark organic matter in soil that is formed by the decomposition of plant and animal matter. It is a kind of soil organic matter. It is rich in nutrients and retains moisture in the soil. Humus is the Lati ...
. It can tolerate soils with a
ph level In chemistry, pH (), historically denoting "potential of hydrogen" (or "power of hydrogen"), is a scale used to specify the acidity or basicity of an aqueous solution. Acidic solutions (solutions with higher concentrations of ions) are me ...
of 6.8. It prefers positions in full sun. It needs a dormant dry period (or drought) during the summer. The plants are not tolerant of winds. In non-hardy places, it is best grown in a bulb frame. It can be also grown in rock garden. The plants should be planted at a depth of 2 inches, in October. It is thought to be one of the easiest 'Regelia' section irises to cultivate, as it is more adaptable and vigorous than others.


Hybrids and cultivars

''Iris stolonifera'' has been crossed with ''Iris korolkowii'' to create different coloured hybrids that have dark brown or deep blue beards. It has been crossed with other irises but did not produce very suitable crosses. There are several different cultivars, including 'Chocolate', 'Decorated Blue Beard', 'Decorated Delight', 'Decorated Giant', 'George Barr', 'Here I Am', 'Leichtlini', 'Merlin's Magic', 'Network', 'Real Harmony', 'Red, White, And Blue', 'Conical', 'Turkish Dancer', 'Turkish Delight', 'Vaga', and 'Zwanenburg Beauty'. 'Zwanenburg Beauty' has blue (or blue white ) flowers that are edged in bronze and have bronze veins. It grows to a height of between tall. It has lilac shading on the falls. The beard is bronze-red or cream-white.


References


Sources

* Aldén, B., S. Ryman & M. Hjertson. 2009. Våra kulturväxters namn – ursprung och användning. Formas, Stockholm (Handbook on Swedish cultivated and utility plants, their names and origin). * Czerepanov, S. K. 1995. Vascular plants of Russia and adjacent states (the former USSR). * Dykes, The Genus Iris, page 126, 1913 * Khassanov, F. O. & N. Rakhimova. 2012. Taxonomic revision of the genus Iris L. (Iridaceae Juss.) for the flora of Central Asia. Stapfia 97:177. * Komarov, V. L. et al., eds. 1934–1964. Flora SSSR. * Mathew, B. 1981. The Iris. 64.


External links


Has many galleries of images of the iris
{{Taxonbar, from=Q15572366
stolonifera Stolonifera is a suborder of soft corals in the order Alcyonacea. Members of this taxon are characterised by having separate polyps budding off an encrusting horizontal, branching stolon. The skeletons include spicules or consists of a horny exte ...
Flora of Afghanistan Flora of Tajikistan Flora of Central Asia Flora of Uzbekistan Garden plants of Asia Plants described in 1880