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Intrafusal muscle fibers are skeletal muscle fibers that serve as specialized sensory organs ( proprioceptors). They detect the amount and rate of change in length of a
muscle Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate muscular system and typically are attached by tendons to bones of a skeleton. The muscle cells of skeletal muscles are much longer than in the other types of muscl ...
.Casagrand, Janet (2008) ''Action and Movement: Spinal Control of Motor Units and Spinal Reflexes.'' University of Colorado, Boulder. They constitute the muscle spindle, and are innervated by both sensory (afferent) and motor (efferent) fibers. Intrafusal muscle fibers are not to be confused with extrafusal muscle fibers, which contract, generating skeletal movement and are innervated by
alpha motor neuron Alpha (α) motor neurons (also called alpha motoneurons), are large, multipolar lower motor neurons of the brainstem and spinal cord. They innervate extrafusal muscle fibers of skeletal muscle and are directly responsible for initiating their con ...
s.


Structure


Types

There are two types of intrafusal muscle fibers: nuclear bag fibers and nuclear chain fibers. They bear two types of sensory ending, known as annulospiral and flower-spray endings. Both ends of these fibers contract, but the central region only stretches and does not contract. Intrafusal muscle fibers are walled off from the rest of the muscle by an outer
connective tissue Connective tissue is one of the four primary types of animal tissue, along with epithelial tissue, muscle tissue, and nervous tissue. It develops from the mesenchyme derived from the mesoderm the middle embryonic germ layer. Connective tiss ...
sheath consisting of flattened fibroblasts and
collagen Collagen () is the main structural protein in the extracellular matrix found in the body's various connective tissues. As the main component of connective tissue, it is the most abundant protein in mammals, making up from 25% to 35% of the whole ...
. This sheath has a spindle or "fusiform" shape, hence the name "intrafusal".


Innervation

They are innervated by gamma motor neurons and beta motor neurons. Gamma efferents from small multipolar neurons from anterior gray column innervate it. These form a part of neuromuscular spindles.


Function

Intrafusal muscle fibers detect the amount and rate of change in muscle length. It is by the sensory information from gamma motor neurons and beta motor neurons that an individual is able to judge the position of their muscles.


See also

*
Alpha motor neuron Alpha (α) motor neurons (also called alpha motoneurons), are large, multipolar lower motor neurons of the brainstem and spinal cord. They innervate extrafusal muscle fibers of skeletal muscle and are directly responsible for initiating their con ...
* Beta motor neuron * Extrafusal muscle fiber * Gamma motor neuron * Type Ia sensory fiber * Type II sensory fiber


References


External links


Histology at ucsd.edu


* {{Authority control Muscular system