''Ingerophrynus'' is a genus of

true toads A true toad is any member of the family Bufonidae, in the order Anura (frogs and toads). This is the only family of anurans in which all members are known as toads, although some may be called frogs (such as harlequin frogs). The bufonids now ...

with 12 species. The genus is found in southern

Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...

and

Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...

; from

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...

and

Indochina Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...

to peninsular

Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...

and

Malaya Malaya refers to a number of historical and current political entities related to what is currently Peninsular Malaysia in Southeast Asia: Political entities * British Malaya (1826–1957), a loose collection of the British colony of the Straits ...

Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...

Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...

Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...

, Nias Island,

Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...

, and the

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

. This genus was established after a major taxonomical revision of frogs in 2006.

Taxonomy and systematics

Ten of the species in this genus were formerly considered species of the genus ''

Bufo ''Bufo'' is a genus of true toads in the amphibian family Bufonidae. As traditionally defined, it was a wastebasket genus containing a large number of toads from much of the world, but following taxonomic reviews most of these have been moved to ...

'', most of them in what had been known as the '' Bufo biporcatus'' group; the remaining ones were assigned to this genus based on molecular data. In 2007 a new species, '' Ingerophrynus gollum'', was added to this genus. The

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

of ''Ingerophrynus'' is '' Sabahphrynus''.

Etymology

The generic name ''Ingerophrynus'' honors

Robert F. Inger Robert Frederick Inger (September 10, 1920 – April 12, 2019) was an American herpetologist. During his lifetime, he wrote numerous books and publications about herpetology. He was also the curator for amphibians and reptiles at the Field Mus ...

, an American zoologist from the

Field Museum of Natural History The Field Museum of Natural History (FMNH), also known as The Field Museum, is a natural history museum in Chicago, Illinois, and is one of the largest such museums in the world. The museum is popular for the size and quality of its educational ...

Description

The diagnostic characters of the ''Bufo biporcatus'' group are the presence of , , and supra tympanic crests, lack of a tarsal ridge, presence of

vocal sac The vocal sac is the flexible membrane of skin possessed by most male frogs and toads. The purpose of the vocal sac is usually as an amplification of their mating or advertisement call. The presence or development of the vocal sac is one way o ...

s but absence of melanophores in the surrounding muscle tissue, lack of tibial glands, lack supinator manus humeralis and

adductor longus In the human body, the adductor longus is a skeletal muscle located in the thigh. One of the adductor muscles of the hip, its main function is to adduct the thigh and it is innervated by the obturator nerve. It forms the medial wall of the fem ...

muscles, presence of paired crests on the vertebral column, rugose skull,

squamosal bone The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral co ...

s with broad dorsal plates, and smooth

palatine bone In anatomy, the palatine bones () are two irregular bones of the facial skeleton in many animal species, located above the uvula in the throat. Together with the maxillae, they comprise the hard palate. (''Palate'' is derived from the Latin ''pa ...

Species

There are 12 species:

References

{{Taxonbar, from=Q2303810 Bufonidae Amphibian genera Amphibians of Asia Taxa named by Jonathan A. Campbell Taxa named by Darrel Frost Taxa named by Taran Grant Taxa named by John Douglas Lynch Taxa named by Ronald Archie Nussbaum Taxa named by Christopher John Raxworthy