Ingerophrynus Gollum
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Ingerophrynus Gollum
''Ingerophrynus gollum'' (Gollum's toad) is a toad species in the family Bufonidae, the true toads. It is endemic to the Peninsular Malaysia and has only been recorded from its type locality in the Endau-Rompin National Park, Johor. However, ''I. gollum'' is genetically so close to ''Ingerophrynus divergens'' that its validity as a distinct species can be questioned. Etymology It is called " gollum" with reference of the eponymous character of The Lord of the Rings by J. R. R. Tolkien. Description Three adult males in the type series measure in snout–vent length; females are unknown. The head is relatively large whereas the body and the limbs are slender. The tympanum is distinct. The parotoid gland is elongated, low in profile. Fingers are long with blunt tips. The toes are partially webbed. Skin of the flanks and the dorsum is covered with numerous warts. The ground color of the dorsum and the flanks is orange. The mid-dorsal stripe is white and prominent. The inter-orbi ...
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True Toad
A true toad is any member of the family Bufonidae, in the order Anura (frogs and toads). This is the only family of anurans in which all members are known as toads, although some may be called frogs (such as harlequin frogs). The bufonids now comprise more than 35 genera, ''Bufo'' being the best known. History Bufonidae is thought to have originated in South America. Some studies date the origin of the group to after the breakup of Gondwana, about 78 - 98 million years ago in the Late Cretaceous. In contrast, other studies have dated the origin of the group to the early Paleocene. The bufonids likely radiated out of South America during the Eocene, with the entire radiation occurring during the Eocene to Oligocene, marking an extremely rapid divergence likely facilitated by the Paleogene's changing climatic conditions. Taxonomy The following phylogeny of most genera in the family is based on Portik and Papenfuss, 2015:, Chan ''et al.'', 2016, Chandramouli ''et al.'', 2016 ...
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Iris (anatomy)
In humans and most mammals and birds, the iris (plural: ''irides'' or ''irises'') is a thin, annular structure in the eye, responsible for controlling the diameter and size of the pupil, and thus the amount of light reaching the retina. Eye color is defined by the iris. In optical terms, the pupil is the eye's aperture, while the iris is the diaphragm. Structure The iris consists of two layers: the front pigmented fibrovascular layer known as a stroma and, beneath the stroma, pigmented epithelial cells. The stroma is connected to a sphincter muscle (sphincter pupillae), which contracts the pupil in a circular motion, and a set of dilator muscles ( dilator pupillae), which pull the iris radially to enlarge the pupil, pulling it in folds. The sphincter pupillae is the opposing muscle of the dilator pupillae. The pupil's diameter, and thus the inner border of the iris, changes size when constricting or dilating. The outer border of the iris does not change size. The constricti ...
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Amphibians Described In 2007
Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this. The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in recent decades there has been a dramatic ...
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Amphibians Of Malaysia
The amphibians of Malaysia are diverse. Species * ''Amolops larutensis'' * '' Angular caecilian'' * '' Ansonia albomaculata'' * '' Ansonia endauensis'' * '' Ansonia hanitschi'' * '' Ansonia latidisca'' * '' Ansonia minuta'' * '' Ansonia torrentis'' * '' Borneophrys edwardinae'' * '' Bufo asper'' * '' Calluella brooksii'' * '' Calluella flava'' * '' Calluella guttulata'' * '' Calluella volzi'' * '' Caudacaecilia asplenia'' * ''Caudacaecilia larutensis'' * '' Caudacaecilia nigroflava'' * ''Chaperina fusca'' * '' Chiromantis nongkhorensis'' * ''Duttaphrynus melanostictus'' * ''Fejervarya cancrivora'' * ''Fejervarya limnocharis'' * ''Fejervarya pulla'' * '' Hoplobatrachus rugulosus'' * '' Huia cavitympanum'' * ''Humerana miopus'' * ''Hylarana banjarana'' * ''Hylarana baramica'' * '' Hylarana erythraea'' * ''Hylarana glandulosa'' * ''Hylarana laterimaculata'' * '' Hylarana luctuosa'' * '' Hylarana macrodactyla'' * '' Hylarana nigrovittata'' * '' Hylarana siberu'' * '' Hylarana signata' ...
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Endemic Fauna Of Malaysia
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can be also be referred to as an ''endemism'' or in scientific literature as an ''endemite''. For example '' Cytisus aeolicus'' is an endemite of the Italian flora. '' Adzharia renschi'' was once believed to be an endemite of the Caucasus, but it was later discovered to be a non-indigenous species from South America belonging to a different genus. The extreme opposite of an endemic species is one with a cosmopolitan distribution, having a global or widespread range. A rare alternative term for a species that is endemic is "precinctive", which applies to s ...
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Ingerophrynus
''Ingerophrynus'' is a genus of true toads with 12 species. The genus is found in southern Yunnan and Southeast Asia; from Myanmar and Indochina to peninsular Thailand and Malaya, Sumatra, Borneo, Java, Nias Island, Sulawesi, and the Philippines. This genus was established after a major taxonomical revision of frogs in 2006. Taxonomy and systematics Ten of the species in this genus were formerly considered species of the genus ''Bufo'', most of them in what had been known as the '' Bufo biporcatus'' group; the remaining ones were assigned to this genus based on molecular data. In 2007 a new species, '' Ingerophrynus gollum'', was added to this genus. The sister taxon of ''Ingerophrynus'' is '' Sabahphrynus''. Etymology The generic name ''Ingerophrynus'' honors Robert F. Inger, an American zoologist from the Field Museum of Natural History. Description The diagnostic characters of the ''Bufo biporcatus'' group are the presence of , , and supra tympanic crests, lack of a tarsal r ...
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Oil Palm
''Elaeis'' () is a genus of palms containing two species, called oil palms. They are used in commercial agriculture in the production of palm oil. The African oil palm ''Elaeis guineensis'' (the species name ''guineensis'' referring to its country of origin) is the principal source of palm oil. It is native to west and southwest Africa, occurring between Angola and Gambia. The American oil palm ''Elaeis oleifera'' () is native to tropical Central and South America, and is used locally for oil production. Description Mature palms are single-stemmed, and can grow well over tall. The leaves are pinnate, and reach between long. The flowers are produced in dense clusters; each individual flower is small, with three sepals and three petals. The palm fruit is reddish, about the size of a large plum, and grows in large bunches. Each fruit is made up of an oily, fleshy outer layer (the pericarp), with a single seed (the palm kernel), also rich in oil. Species The two species, '' ...
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Agricultural Expansion
Agricultural expansion describes the growth of agricultural land (arable land, pastures, etc.) especially in the 20th and 21st centuries. The agricultural expansion is often explained as a direct consequence of the global increase in food and energy requirements due to continuing population growth (both which in turn have been attributed to agricultural expansion itself), with an estimated expectation of 10 to 11 billion humans on Earth by end of this century. It is foreseen that most of the world's non-agrarian ecosystems (terrestrial and aquatic) will be affected adversely, from habitat loss, land degradation, overexploitation, and other problems. The intensified food (and biofuel) production will in particular affect the tropical regions. Most modern agriculture relies on intensive methods. Further expansion of the predominant farming types that rest on a small number of highly productive crops has led to a significant loss of biodiversity on a global scale already. More ...
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Above Sea Level
Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The combination of unit of measurement and the physical quantity (height) is called "metres above mean sea level" in the metric system, while in United States customary and imperial units it would be called "feet above mean sea level". Mean sea levels are affected by climate change and other factors and change over time. For this and other reasons, recorded measurements of elevation above sea level at a reference time in history might differ from the actual elevation of a given location over sea level at a given moment. Uses Metres above sea level is the standard measurement of the elevation or altitude of: * Geographic locations such as towns, mountains and other landmarks. * The top of buildings and other structures. * Flying objects such ...
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Sexual Selection In Amphibians
Sex is the biological distinction of an organism between male and female. Sex or SEX may also refer to: Biology and behaviour *Animal sexual behaviour **Copulation (zoology) **Human sexual activity **Non-penetrative sex, or sexual outercourse **Sex drive, a person's overall sexual drive or desire for sexual activity ** Sexual intercourse, also called copulation or coitus *Gender, the distinction between male and female or masculinity and femininity within an individual's gender identity **Sex and gender distinction *Human sexuality *Mating types, a distinction of gametes, whether in anisogamous or isogamous species * Sexing, the act of discerning the sex of an animal *Sexual reproduction, a process of combining and mixing genetic traits, associated with the generation of new individuals, by means of meiosis and fertilization ** Genetic recombination, the process of mixing genetic traits solely, occurring both in organisms with sexual or asexual reproduction Art and entertainment ...
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Vocal Sac
The vocal sac is the flexible membrane of skin possessed by most male frogs and toads. The purpose of the vocal sac is usually as an amplification of their mating or advertisement call. The presence or development of the vocal sac is one way of externally determining the sex of a frog or toad in many species; taking frogs as an example; The vocal sac is open to the mouth cavity of the frog, with two slits on either side of the tongue. To call, the frog inflates its lungs and shuts its nose and mouth. Air is then expelled from the lungs, through the larynx, and into the vocal sac. The vibrations of the larynx emits a sound, which resonates on the elastic membrane of the vocal sac. The resonance causes the sound to be amplified and allows the call to carry further. Muscles within the body wall force the air back and forth between the lungs and vocal sac. Development The development of the vocal sac is different in most species, however they mostly follow the same process. The d ...
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Gular Skin
Gular skin (throat skin), in ornithology, is an area of featherless skin on birds that joins the lower mandible of the beak (or ''bill'') to the bird's neck. Other vertebrate taxa may have a comparable anatomical structure that is referred to as either a gular sac, throat sac, vocal sac or gular fold. In birds Gular skin can be very prominent, for example in members of the order Phalacrocoraciformes as well as in pelicans (which likely share a common ancestor). In many species, the gular skin forms a flap, or gular pouch, which is generally used to store fish and other prey while hunting. In cormorants, the gular skin is often colored, contrasting with the otherwise plain black or black-and-white appearance of the bird. This presumably serves some function in social signalling, since the colors become more pronounced in breeding adults. In frigatebirds, the gular skin (or gular sac or throat sac) is used dramatically. During courtship display, the male forces air into the sac, ...
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