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''Icerya'' is a genus of
scale insect Scale insects are small insects of the order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than the ...
s in the family
Monophlebidae Monophlebidae is a family of scale insects commonly known as the giant scales or monophlebids. They occur in most parts of the world but more genera are found in the tropics than elsewhere.British Mauritius Mauritius was a Crown colony off the Southeast coast of Africa. Formerly part of the French colonial empire, British rule in Mauritius was established de facto with the Invasion of Isle de France in November 1810, and de jure by the subsequent T ...
.


Hermaphroditism

Hermaphroditism is extremely rare in the insect world despite the comparatively common nature of this condition in the crustaceans. Several species of ''Icerya'', including the pestiferous cottony-cushion scale, ''I. purchasi'', are known to be hermaphrodites that reproduce by self-fertilising. Occasionally males are produced from unfertilised eggs, but generally individuals are monoecious with a female-like nature but possessing an ovotestis (a part-testis, part-ovary organ) and sperm is transmitted ovarially from the female to her young. This hermaphroditic sexual self-sufficiency, where a single individual can populate new territory, has contributed to the invasive spread of the cottony-cushion scale insect away from its native Australia.The Insects An outline of Entomology, Gullan & Cranston, Wiley-Blackwell 2001


List of species

* ''
Icerya aegyptiaca ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively co ...
'' (Douglas 1890). * ''
Icerya albolutea ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Cockerell 1898. * '' Icerya bimaculata'' De Lotto 1959. * ''
Icerya brachystegiae ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hall 1940. * '' Icerya brasiliensis'' Hempel 1920. * ''
Icerya callitri ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' (Froggatt 1921). * '' Icerya chilensis'' Hempel 1920. * ''
Icerya colimensis ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Cockerell 1902. * ''
Icerya flava ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1920. * ''
Icerya flocculosa ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1932. * ''
Icerya formicarum ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively co ...
'' Newstead 1897. * ''
Icerya genistae ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively co ...
'' Hempel 1912. * ''
Icerya hanoiensis ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Jashenko & Danzig 1992. * ''
Icerya imperatae ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Rao 1951. * ''
Icerya insulans ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1923. * ''
Icerya koebelei ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Maskell 1892. * ''
Icerya leuderwaldti ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1918. * ''
Icerya littoralis mimosae ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Cockerell 1902. * ''
Icerya littoralis tonilensis ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Cockerell 1902. * ''
Icerya littoralis ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Cockerell 1898. * ''
Icerya longisetosa ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Newstead 1911. * ''
Icerya maxima ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Newstead 1915. * ''
Icerya maynei ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Vayssiere 1926. * ''
Icerya menoni ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Rao 1951. * ''
Icerya minima ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Morrison 1919. * ''
Icerya minor ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Green 1908. * ''
Icerya montserratensis ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Riley & Howard 1890. * ''
Icerya morrisoni ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Rao 1951. * '' Icerya nigroareolata'' Newstead 1917. * '' Icerya palmeri'' Riley & Howard 1890. * ''
Icerya paulista ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1920. * ''
Icerya pilosa ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Green 1896. * '' Icerya pulchra'' (Leonardi 1907). * ''
Icerya purchasi citriperda ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1920. * ''
Icerya purchasi crawii ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively c ...
'' Cockerell 1897. * '' Icerya purchasi maskelli'' Cockerell 1897. * ''
Icerya purchasi ''Icerya purchasi'' ( common name: cottony cushion scale) is a scale insect that feeds on more than 80 families of woody plants, most notably on ''Citrus'' and ''Pittosporum''. Originally described in 1878 from specimens collected in New Zealand ...
'' Maskell 1878 - cottony cushion scale * ''
Icerya rileyi ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Cockerell 1896. * ''
Icerya schoutedeni ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Vayssiere 1926. * ''
Icerya schrottkyi ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1900. * ''
Icerya seychellarum (type) ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' (Westwood 1855). * ''
Icerya seychellarum cristata ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Newstead 1909. * ''
Icerya similis ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Morrison 1923. * '' Icerya splendida'' Lindinger 1913. * ''
Icerya subandina ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Leonardi 1911. * '' Icerya sulfurea pattersoni'' Newstead 1917. * ''
Icerya sulfurea ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Lindinger 1913. * ''
Icerya sumatrana ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Rao 1951. * ''
Icerya taunayi ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Hempel 1920. * ''
Icerya travancorensis ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Rao 1951. * ''
Icerya tremae ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively co ...
'' Vayssiere 1926. * ''
Icerya zeteki ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively co ...
'' Cockerell 1914. * ''
Icerya zimmermani ''Icerya'' is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius. Hermaphroditism Hermaphroditism is extremely rare in the insect world despite the comparatively com ...
'' Green 1932.


References

Monophlebidae Sternorrhyncha genera {{Coccoidea-stub