''Huanhepterus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

ctenochasmatid Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early C ...

pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...

from the

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

period of what is now

Qingyang Qingyang () is a prefecture-level city in eastern Gansu province, China. Geography and climate Qingyang is the easternmost prefecture-level division of Gansu and is thus sometimes referred to as "Longdong" (). It forms an administrative penins ...

Gansu Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibet ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

History

The genus was named by

Dong Zhiming Dong Zhiming (Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at the ...

in 1982. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Huanhepterus quingyangensis''. The genus name refers to the Huang Jian (not the

Yellow River The Yellow River or Huang He (Chinese: , Standard Beijing Mandarin, Mandarin: ''Huáng hé'' ) is the second-longest river in China, after the Yangtze River, and the List of rivers by length, sixth-longest river system in the world at th ...

or "Huang He", but a smaller tributary of the Jinghe River in Gansu), and combines it with a Latinized Greek ''pteron'', "wing". The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

refers to Qinyang County. ''Huanhepterus'' is based on the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

specimen, IVPP V9070, a partial articulated skeleton consisting mostly of impressions of the left half of the body and the

beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...

-end of the skull. The

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

was in May 1978 found in a quarry operated by the Sanshilipu-

commune A commune is an alternative term for an intentional community. Commune or comună or comune or other derivations may also refer to: Administrative-territorial entities * Commune (administrative division), a municipality or township ** Communes of ...

, when an explosion exposed a vertebra. Its force obliterated the right half of the specimen. IVPP V9070 hails from the Early Cretaceous-age Huachihuanhe Formation of the

Zhidan Group The Zhidan Group is a geological formation in China whose strata date back to the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Earl ...

Description

''Huanhepterus'' had a long, low skull, with a low crest running along the midline that was higher toward the tip of the snout and became smaller toward the eyes. The

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

, about 26 pairs in the upper and 25 in the lower jaws, were slender and numerous, and became shorter farther from the 11th pair, both to the front as to the back, where they become absent completely in posterior part of the snout. The

cervical In anatomy, cervical is an adjective that has two meanings: # of or pertaining to any neck. # of or pertaining to the female cervix: i.e., the ''neck'' of the uterus. *Commonly used medical phrases involving the neck are **cervical collar **cervic ...

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

e were long, as were the toes, and there was no fused complex of the front dorsal vertebrae (

notarium Notarium or os dorsale is a bone consisting of the fused vertebra of the shoulder in birds and some pterosaurs. The structure helps brace the chest against the forces generated by the wings. In birds, the vertebrae are only in contact with adjace ...

), as seen in other pterosaurs. The

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...

of the type individual was estimated at 2.5 m (8.2 ft). This genus was described as most like ''

Gnathosaurus ''Gnathosaurus'' (meaning "jawed lizard") is a genus of ctenochasmatid pterosaur containing two species: ''G. subulatus'', named in 1833 from the Solnhofen Limestone of Germany, and ''G. macrurus'', known from the Purbeck Limestone of the UK. I ...

''.Dong, Z.-M. (1982). On a new Pterosauria (''Huanhepterus quingyangensis'' gen.et. sp.nov.) from Ordos, China. ''Vertebrata PalAsiatica'' 20(2):115-121. David Unwin later referred it the

Gnathosaurinae Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early ...

, a subgroup of the

Ctenochasmatidae Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early C ...

Classification

Below is

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

following a topology by Andres, Clark and Xu (2014). In the analysis, they recovered ''Huanhepterus'' within the family Ctenochasmatidae, more precisely within the subfamily

, though placed in the basalmost position.

Biology

Like ''

'', it may have used its tightly-packed, slender teeth to filter food from water.

References

External links