Holoaden
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''Holoaden'', the highland frogs, is a small genus of
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s in the family Strabomantidae. The species are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to south-eastern Brazil.


Taxonomy

''Holoaden'' is the type genus of
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Holoadeninae that was erected in 2008 and placed in the newly erected family Strabomantidae. Prior to this, it had been placed in the genus
Brachycephalidae The Brachycephalidae () are a family of frogs confined to eastern and southern Brazil and northern Argentina. The family is composed of two externally quite different genera: the tiny, often (but not always) colourful and plump saddleback toads ( ...
.


Description

''Holoaden'' grow to snout–vent length at most. Head is narrower than the body. Differentiated tympanic membrane and tympanic annulus are absent. Dorsum is highly glandular. Venter is areolate.


Species

There are four recognized species in this genus: * '' Holoaden bradei'' Lutz, 1958 * ''
Holoaden luederwaldti ''Holoaden luederwaldti'' is a species of frog in the family Strabomantidae. It is endemic to Brazil. Its natural habitats are subtropical or tropical high-altitude shrubland, subtropical or tropical high-altitude grassland, and rocky areas. It ...
'' Miranda-Ribeiro, 1920 * '' Holoaden pholeter'' Pombal, Siqueira, Dorigo, Vrcibradic, and Rocha, 2008 * '' Holoaden suarezi'' Martins and Zaher, 2013


References

{{Taxonbar, from=Q2129401 Strabomantidae Endemic fauna of Brazil Amphibians of South America Amphibian genera Taxa named by Alípio de Miranda-Ribeiro