Heteropodidae

Huntsman spiders, members of the family Sparassidae (formerly Heteropodidae), are known by this name because of their speed and mode of hunting. They are also called giant crab spiders because of their size and appearance. Larger species sometimes are referred to as wood spiders, because of their preference for woody places (forests, mine shafts, woodpiles, wooden shacks). In southern Africa the genus ''Palystes'' are known as rain spiders or lizard-eating spiders. Commonly, they are confused with baboon spiders from the Mygalomorphae infraorder, which are not closely related.

More than a thousand Sparassidae species occur in most warm temperate to tropical regions of the world, including much of Australasia, Africa, Asia, the Mediterranean Basin, and the Americas.

Several species of huntsman spider can use an unusual form of locomotion. The wheel spider (''Carparachne aureoflava'') from the Namib uses a cartwheeling motion which gives it its name, while ''Cebrennus rechenbergi'' uses a handspring motion.

Description

Sparassids are eight-eyed spiders. The eyes appear in two largely forward-facing rows of four on the anterior aspect of the prosoma. Many species grow very large – in Laos, male giant huntsman spiders (''Heteropoda maxima'') attain a legspan of . Persons unfamiliar with spider taxonomy commonly confuse large species with tarantulas, but huntsman spiders can generally be identified by their legs, which, rather than being jointed vertically relative to the body, are twisted in such a way that in some attitudes the legs extend forward in a crab-like fashion.

On their upper surfaces the main colours of huntsman spiders are inconspicuous shades of brown or grey, but many species have undersides more or less aposematically marked in black-and-white, with reddish patches over the mouthparts. Their legs bear fairly prominent spines, but the rest of their bodies are smoothly furry. They tend to live under rocks, bark and similar shelters, but human encounters are common in sheds, garages and other infrequently-disturbed places. The banded huntsman (''Holconia'') is large, grey to brown with striped bands on its legs. The badge huntsman (''Neosparassus'') is larger still, brown and hairy. The tropical or brown huntsman (''Heteropoda'') is also large and hairy, with mottled brown, white and black markings. The eyesight of these spiders is not nearly as good as that of the ''Salticidae'' ( jumping spiders). Nevertheless, their vision is quite sufficient to detect approaching humans or other large animals from some distance.

File:Sparassidae Palystes superciliosus Male Ventral Aspect 0110s.jpg, ''Palystes superciliosus'', ventral aspect, showing aposematic coloration, plus typically masculine gracile build and clavate pedipalps armed with mating spurs
File:Huntsman spider with meal.jpg, A huntsman spider consuming a small beetle
File:Huntsman Spider feeding Grasshopper.jpg, A female ''Heteropoda venatoria'' consuming a katydid
File:Huntsman spider on log.jpg, Adult social huntsman ''Delena cancerides'' on the underside of a log in Victoria, Australia
File:Olios argelasius MHNT.jpg, ''Olios argelasius''

Identification

They can be distinguished from other spider families by their appearance, as other spiders similar to them are smaller in size. They are often confused for tarantulas due to their hairy nature, but can easily be distinguished by their laterigrade legs, similar to those of crabs. Members of this family are also typically less bulky than tarantulas. They possess two claws, as is the case for most spiders that actively hunt their prey. If this is not enough to fully identify them, they also possess eight eyes divided into two regular rows.

Size, venom, and aggression

On average, a huntsman spider's leg-span can reach up to , while their bodies measure about long.

Like most spiders, Sparassidae use venom to immobilize prey. They have been known to inflict serious defensive bites on humans.

There have been reports of members of various genera such as ''Palystes'', ''Neosparassus'' and several others, inflicting severe bites. The effects vary, including local swelling and pain, nausea, headache, vomiting, irregular pulse rate, and heart palpitations, indicating some systemic neurological toxin effects, especially when the bites were severe or repeated. However, the formal study of spider bites is fraught with complications, including unpredictable infections, dry bites, shock, nocebo effects, and even bite misdiagnosis by medical professionals and specimen misidentification by the general public.

It is not always clear what provokes Sparassidae to attack and bite humans and animals, but it is known that female members of this family will aggressively defend their egg-sacs and young against perceived threats. Bites from sparassids usually do not require hospital treatment.

Sound-production in mating-rituals

Males of ''Heteropoda venatoria'', one of the huntsman spiders that seems to easily find its way around the world, have recently been found to deliberately make a substrate-borne sound when they detect a chemical ( pheromone) left by a nearby female of their species. The males anchor themselves firmly to the surface onto which they have crawled and then use their legs to transmit vibrations from their bodies to the surface. Most of the sound emitted is produced by strong vibrations of the abdomen. The characteristic frequency of vibration and the pattern of bursts of sound identify them to females of their species, who will approach if they are interested in mating. This sound can often be heard as a rhythmic ticking, somewhat like a quartz clock, which fades in and out and can be heard by human ears in a relatively quiet environment.

Genera

, the World Spider Catalog accepted the following genera:

*''Adcatomus'' Karsch, 1880 — Venezuela, Peru
*''Anaptomecus'' Simon, 1903 — Central America, South America
*''Anchonastus'' Simon, 1898 — Cameroon, Congo
*''Arandisa'' Lawrence, 1938 — Namibia
*''Barylestis'' Simon, 1910 — Africa, Asia, Europe
*''Beregama'' Hirst, 1990 — Australia, Papua New Guinea
*''Berlandia'' Lessert, 1921 — East Africa
*''Bhutaniella'' Jäger, 2000 — Asia
*''Borniella'' Grall & Jäger, 2022 — Borneo
*''Caayguara'' Rheims, 2010 — Brazil
*''Carparachne'' Lawrence, 1962 — Namibia
*''Cebrennus'' Simon, 1880 — Africa, Asia, Malta
*'' Cerbalus'' Simon, 1897 — Israel, Jordan, Egypt
*''Chrosioderma'' Simon, 1897 — Madagascar
*''Clastes'' Walckenaer, 1837 — Indonesia, Papua New Guinea
*''Curicaberis'' Rheims, 2015 — North America, Central America, Brazil
*'' Damastes'' Simon, 1880 — Madagascar, Mozambique, Seychelles
*''Decaphora'' Franganillo, 1931 — North America, Caribbean, Central America, Colombia
*''Deelemanikara'' Jäger, 2021 — Madagascar
*''Defectrix'' Petrunkevitch, 1925 — Panama
*''Delena'' Walckenaer, 1837 — Australia, New Zealand
*''Dermochrosia'' Mello-Leitão, 1940 — Brazil
*''Diminutella'' Rheims & Alayón, 2018 — Cuba
*''Eusparassus'' Simon, 1903 — Asia, Africa, Europe, Peru
*''Exopalystes'' Hogg, 1914 — Papua New Guinea
*''Extraordinarius'' Rheims, 2019 — Brazil
*''Geminia'' Thorell, 1897 — Myanmar
*''Gnathopalystes'' Rainbow, 1899 — Asia, Oceania
*''Guadana'' Rheims, 2010 — Brazil, Peru, Ecuador
*''Heteropoda'' Latreille, 1804 — Oceania, Asia, South America, Greece
*''Holconia'' Thorell, 1877 — Australia
*''Irileka'' Hirst, 1998 — Australia
*''Isopeda'' L. Koch, 1875 — Australia, Philippines, Papua New Guinea
*''Isopedella'' Hirst, 1990 — Australia, Papua New Guinea, Indonesia
*''Keilira'' Hirst, 1989 — Australia
*''Leucorchestris'' Lawrence, 1962 — Angola, Namibia
*''Macrinus'' Simon, 1887 — South America, Tobago, United States
*''Martensopoda'' Jäger, 2006 — India
*''May'' Jäger & Krehenwinkel, 2015 — Namibia, South Africa
*''Megaloremmius'' Simon, 1903 — Madagascar
*''Menarik'' Grall & Jäger, 2022 — Borneo
*''Meri'' Rheims & Jäger, 2022 — South America
*'' Micrommata'' Latreille, 1804 — Spain, Africa, Asia
*''Micropoda'' Grall & Jäger, 2022 — Papua New Guinea
*''Microrchestris'' Lawrence, 1962 — Namibia
*''Neosparassus'' Hogg, 1903 — Australia
*''Neostasina'' Rheims & Alayón, 2016 — Caribbean
*''Nolavia'' Kammerer, 2006 — Brazil
*''Nungara'' Pinto & Rheims, 2016 — Brazil, Ecuador
*''Olios'' Walckenaer, 1837 — Asia, South America, Oceania, Africa, Central America, North America, Caribbean
*'' Orchestrella'' Lawrence, 1965 — Namibia
*''Origes'' Simon, 1897 — Argentina, Peru, Ecuador
*''Paenula'' Simon, 1897 — Ecuador
*''Palystella'' Lawrence, 1928 — Namibia
*''Palystes'' L. Koch, 1875 — Africa, India, Australia
*''Panaretella'' Lawrence, 1937 — South Africa
*''Pandercetes'' L. Koch, 1875 — Asia, Oceania
*''Parapalystes'' Croeser, 1996 — South Africa
*''Pediana'' Simon, 1880 — Indonesia, Australia
*''Platnickopoda'' Jäger, 2020 — East Africa
*''Pleorotus'' Simon, 1898 — Seychelles
*''Polybetes'' Simon, 1897 — South America
*'' Prusias'' O. Pickard-Cambridge, 1892 — Brazil, Mexico, Panama
*''Prychia'' L. Koch, 1875 — Papua New Guinea, Fiji, Philippines
*''Pseudomicrommata'' Järvi, 1914 — Africa
*'' Pseudopoda'' Jäger, 2000 — Asia
*''Quemedice'' Mello-Leitão, 1942 — Brazil, Argentina
*''Remmius'' Simon, 1897 — Africa
*''Rhacocnemis'' Simon, 1897 — Seychelles
*''Rhitymna'' Simon, 1897 — Asia, Africa
*''Sadala'' Simon, 1880 — South America
*''Sagellula'' Strand, 1942 — Japan, China
*''Sarotesius'' Pocock, 1898 — East Africa
*''Sinopoda'' Jäger, 1999 — Asia
*''Sivalicus'' Dyal, 1957 — India
*''Sparianthina'' Banks, 1929 — South America, Tobago, Central America
*''Sparianthis'' Simon, 1880 — Colombia
*'' Spariolenus'' Simon, 1880 — Asia
*''Staianus'' Simon, 1889 — Madagascar
*''Stasina'' Simon, 1877 — South America, Gabon, Asia, Cuba
*'' Stasinoides'' Berland, 1922 — Ethiopia
*'' Stipax'' Simon, 1898 — Seychelles
*''Strandiellum'' Kolosváry, 1934 — Papua New Guinea
*'' Thelcticopis'' Karsch, 1884 — Asia, Oceania, Africa
*''Thomasettia'' Hirst, 1911 — Seychelles
*''Thunberga'' Jäger, 2020 — Madagascar
*''Tibellomma'' Simon, 1903 — Venezuela
*''Tiomaniella'' Grall & Jäger, 2022 — Malaysia
*'' Tychicus'' Simon, 1880 — Philippines, Papua New Guinea, Indonesia
*''Typostola'' Simon, 1897 — Australia, Papua New Guinea
*''Uaiuara'' Rheims, 2013 — Panama, South America
*''Vindullus'' Simon, 1880 — South America, Guatemala
*''Yiinthi'' Davies, 1994 — Australia, Papua New Guinea
*''Zachria'' L. Koch, 1875 — Australia

Distribution and habitat

Members of the Sparassidae are native to tropical and warm temperate regions worldwide. A few species are native to colder climates, like the green huntsman spider (''Micrommata virescens'') which is native to Northern and Central Europe. Some tropical species like ''Heteropoda venatoria'' (Cane huntsman) and ''Delena cancerides'' (Social huntsman) have been accidentally introduced to many subtropical parts of the world, including New Zealand (which has no native sparassid species). The huntsman spiders found in southern parts of Florida are of the ''Heteropoda venatoria'' species and considered an invasive species transplanted from Asia. Because of their speed, they commonly hunt and eat cockroaches and are found in many homes.

As adults, huntsman spiders do not build webs, but hunt and forage for food: their diet consists primarily of insects and other invertebrates, and occasionally small skinks and geckos. They live in the crevices of tree bark, but will frequently wander into homes and vehicles. They are able to travel extremely quickly, often using a springing jump while running, and walk on walls and even on ceilings. They also tend to exhibit a "cling" reflex if picked up, making them difficult to shake off and much more likely to bite. The females are fierce defenders of their egg sacs and young. They will generally make a threat display if provoked, and if the warning is ignored they may attack and bite. The egg sacs differ fairly widely among the various genera. For example, in ''Heteropoda'' spp. egg sacs are carried underneath the female's body, while in other species like ''Palystes'' and ''Pseudomicrommata'' spp., females generally attach egg sacs to vegetation.

File:Huntsman Spider egg sac hatching 23 Jan 11.JPG, Australian sparassid egg sac hatching
File:Palystes castaneus (Sparassidae egg purse IMG 2662.jpg, ''Palystes castaneus'' egg purse

See also

* Cultural depictions of spiders (section: Modern myths and urban legends)
* List of Sparassidae species
* Spider wasp
* Table of spider families

References

Inline citations

General references

*

External links

Images and information on Australian Huntsman Spiders
Minibeast Wildlife

Information page on Huntsman Spiders
from the Australian Museum

Badge Huntsman Spider
at Victorian Museum


University of Southern Queensland – pictures & descriptions


a
American Arachnological Society


at University of Florida Institute of Food and Agricultural Sciences

Video of Olios sp. from Costa Rica

{{Taxonbar, from=Q11721

*
Taxa named by Philipp Bertkau