Herring are

forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the food ...

, mostly belonging to the family of

Clupeidae Clupeidae is a family of ray-finned fishes, comprising, for instance, the herrings, shads, sardines, hilsa, and menhadens. The clupeoids include many of the most important food fishes in the world, and are also commonly caught for production of ...

. Herring often move in large

schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...

around

fishing bank An ocean bank, sometimes referred to as a fishing bank or simply bank, is a part of the seabed that is shallow compared to its surrounding area, such as a shoal or the top of an underwater hill. Somewhat like continental slopes, ocean bank sl ...

s and near the coast, found particularly in shallow, temperate waters of the

North Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

and

North Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Africa, Europe an ...

s, including the

Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and from ...

, as well as off the west coast of South America. Three species of ''Clupea'' (the

type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal f ...

of the herring family Clupeidae) are recognised, and comprise about 90% of all herrings captured in fisheries. The most abundant of these species is the

Atlantic herring Atlantic herring (''Clupea harengus'') is a herring in the family (biology), family Clupeidae. It is one of the most abundant fish species in the world. Atlantic herrings can be found on both sides of the Atlantic Ocean, congregating in large ...

, which comprises over half of all herring capture. Fish called herring are also found in the

Arabian Sea The Arabian Sea ( ar, اَلْبَحرْ ٱلْعَرَبِيُّ, Al-Bahr al-ˁArabī) is a region of the northern Indian Ocean bounded on the north by Pakistan, Iran and the Gulf of Oman, on the west by the Gulf of Aden, Guardafui Channel ...

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by th ...

, and

Bay of Bengal The Bay of Bengal is the northeastern part of the Indian Ocean, bounded on the west and northwest by India, on the north by Bangladesh, and on the east by Myanmar and the Andaman and Nicobar Islands of India. Its southern limit is a line between ...

. Herring played an important role in the history of marine fisheries in Europe, and early in the 20th century, their study was fundamental to the development of

fisheries science Fisheries science is the academic discipline of managing and understanding fisheries. It is a multidisciplinary science, which draws on the disciplines of limnology, oceanography, freshwater biology, marine biology, meteorology, conservation, eco ...

. These

oily fish Oily fish are fish species with oil (fats) in soft tissues and in the coelomic cavity around the gut. Their fillets may contain up to 30% oil, although this figure varies both within and between species. Examples of oily fish include small ...

also have a long history as an important

food fish Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is ingest ...

, and are often salted,

smoked Smoking is the process of flavoring, browning, cooking, or preserving food by exposing it to smoke from burning or smoldering material, most often wood. Meat, fish, and ''lapsang souchong'' tea are often smoked. In Europe, alder is the tradi ...

, or

pickled Pickling is the process of preserving or extending the shelf life of food by either anaerobic fermentation in brine or immersion in vinegar. The pickling procedure typically affects the food's texture and flavor. The resulting food is called ...

. Herring are also known as "silver darlings".

Species

A number of different species, most belonging to the family

, are commonly referred to as herrings. The origins of the term "herring" is somewhat unclear, though it may derive from the

Old High German Old High German (OHG; german: Althochdeutsch (Ahd.)) is the earliest stage of the German language, conventionally covering the period from around 750 to 1050. There is no standardised or supra-regional form of German at this period, and Old High ...

''heri'' meaning a "host, multitude", in reference to the large schools they form. The

of the herring family Clupeidae is ''

Clupea ''Clupea'' is genus of planktivorous bony fish belonging to the family Clupeidae, commonly known as herrings. They are found in the shallow, temperate waters of the North Pacific and the North Atlantic oceans, including the Baltic Sea. Two main ...

''.Pauly, Daniel (2004
''Darwin's Fishes: An Encyclopedia of Ichthyology, Ecology, and Evolution''
Page 109, Cambridge University Press. . ''Clupea'' contains only two species: the

(the type species) found in the North Atlantic, and the

Pacific herring The Pacific herring (''Clupea pallasii'') is a species of the herring family associated with the Pacific Ocean environment of North America and northeast Asia. It is a silvery fish with unspined fins and a deeply forked caudal fin. The distribut ...

mainly found in the North Pacific. Subspecific divisions have been suggested for both the Atlantic and Pacific herrings, but their biological basis remains unclear. In addition, a number of related species, all in the Clupeidae, are commonly referred to as herrings. The table immediately below includes those members of the family Clupeidae referred to by

FishBase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.

as herrings which have been assessed by the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

. Also, a number of other species are called herrings, which may be related to clupeids or just share some characteristics of herrings (such as the

lake herring ''Coregonus artedi'', commonly known as the cisco, is a North American species of freshwater whitefish in the family Salmonidae. The number of species and definition of species limits in North American ciscoes is a matter of debate. Accordingly ...

, which is a

salmonid Salmonidae is a family of ray-finned fish that constitutes the only currently extant family in the order Salmoniformes . It includes salmon (both Atlantic and Pacific species), trout (both ocean-going and landlocked), chars, freshwater whitefis ...

). Just which of these species are called herrings can vary with locality, so what might be called a herring in one locality might be called something else in another locality. Some examples:

Characteristics

The species of ''Clupea'' belong to the larger family Clupeidae (herrings,

shad The Alosinae, or the shads,Alosinae

sardines "Sardine" and "pilchard" are common names for various species of small, oily forage fish in the herring family Clupeidae. The term "sardine" was first used in English during the early 15th century, a folk etymology says it comes from the I ...

menhaden Menhaden, also known as mossbunker and bunker and "the most important fish in the sea", are forage fish of the genera ''Brevoortia'' and ''Ethmidium'', two genera of marine fish in the family Clupeidae. ''Menhaden'' is a blend of ''poghaden'' ...

s), which comprises some 200 species that share similar features. These silvery-coloured fish have a single

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

, which is soft, without spines. They have no

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...

and have a protruding lower jaw. Their size varies between subspecies: the

Baltic herring Atlantic herring (''Clupea harengus'') is a herring in the family Clupeidae. It is one of the most abundant fish species in the world. Atlantic herrings can be found on both sides of the Atlantic Ocean, congregating in large schools. They can ...

(''Clupea harengus membras'') is small, 14 to 18 cm; the proper Atlantic herring (''Clupea harengus harengus'') can grow to about 46 cm (18 in) and weigh up 700 g (1.5 lb); and

grow to about 38 cm (15 in).

Lifecycle

At least one stock of Atlantic herring

spawn Spawn or spawning may refer to: * Spawn (biology), the eggs and sperm of aquatic animals Arts, entertainment, and media * Spawn (character), a fictional character in the comic series of the same name and in the associated franchise ** '' Spawn: Ar ...

s in every month of the year. Each spawns at a different time and place (spring, summer, autumn, and winter herrings).

Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...

populations spawn in of water, while North Sea (bank) herrings spawn at down to in autumn. Eggs are laid on the sea bed, on rock, stones, gravel, sand or beds of

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

. Females may deposit from 20,000 to 40,000 eggs, according to age and size, averaging about 30,000. In sexually mature herring, the genital organs grow before spawning, reaching about one-fifth of its total weight. The eggs sink to the bottom, where they stick in layers or clumps to gravel, seaweed, or stones, by means of their mucous coating, or to any other objects on which they chance to settle. If the egg layers are too thick they suffer from oxygen depletion and often die, entangled in a maze of

mucus Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It is ...

. They need substantial water microturbulence, generally provided by

wave action In fluid dynamics, a wind wave, water wave, or wind-generated water wave, is a surface wave that occurs on the free surface of bodies of water as a result from the wind blowing over the water surface. The contact distance in the direction of t ...

or coastal

current Currents, Current or The Current may refer to: Science and technology * Current (fluid), the flow of a liquid or a gas ** Air current, a flow of air ** Ocean current, a current in the ocean *** Rip current, a kind of water current ** Current (stre ...

s. Survival is highest in crevices and behind solid structures, because predators feast on openly exposed eggs. The individual eggs are in diameter, depending on the size of the parent fish and also on the local race. Incubation time is about 40 days at , 15 days at , or 11 days at . Eggs die at temperatures above . The larvae are long at hatching, with a small yolk sac that is absorbed by the time the larvae reach . Only the eyes are well pigmented. The rest of the body is nearly transparent, virtually invisible under water and in natural lighting conditions. The dorsal fin forms at , the anal fin at about —the ventral fins are visible and the tail becomes well forked at 30 to — at about , the larva begins to look like a herring. Herring larvae are very slender and can easily be distinguished from all other young fish of their range by the location of the vent, which lies close to the base of the tail; however, distinguishing clupeoids one from another in their early stages requires critical examination, especially telling herring from

sprat Sprat is the common name applied to a group of forage fish belonging to the genus ''Sprattus'' in the family Clupeidae. The term also is applied to a number of other small sprat-like forage fish (''Clupeoides'', ''Clupeonella'', '' Corica'', ' ...

s. At one year, they are about long, and they first spawn at three years.

Ecology

Prey

Herrings consume

copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...

arrow worm The Chaetognatha or chaetognaths (meaning ''bristle-jaws'') are a phylum of predatory marine worms that are a major component of plankton worldwide. Commonly known as arrow worms, about 20% of the known Chaetognatha species are benthic zone, be ...

s, pelagic amphipods,

mysid Mysida is an order of small, shrimp-like crustaceans in the malacostracan superorder Peracarida. Their common name opossum shrimps stems from the presence of a brood pouch or "marsupium" in females. The fact that the larvae are reared in thi ...

s, and

krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...

in the

pelagic zone The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...

. Conversely, they are a central prey item or

for higher

trophic level The trophic level of an organism is the position it occupies in a food web. A food chain is a succession of organisms that eat other organisms and may, in turn, be eaten themselves. The trophic level of an organism is the number of steps it i ...

s. The reasons for this success are still enigmatic; one speculation attributes their dominance to the huge, extremely fast cruising

they inhabit. Herring feed on

phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...

, and as they mature, they start to consume larger organisms. They also feed on zooplankton, tiny animals found in

ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...

ic surface waters, and small fish and fish larvae. Copepods and other tiny crustaceans are the most common zooplankton eaten by herring. During daylight, herring stay in the safety of deep water, feeding at the surface only at night when the chance of being seen by predators is less. They swim along with their mouths open, filtering the plankton from the water as it passes through their gills. Young herring mostly hunt copepods individually, by means of "particulate feeding" or "raptorial feeding",Kils U (1992)
The ATOLL Laboratory and other Instruments Developed at Kiel
' U.S. GLOBEC News, Technology Forum Number 8: 6–9. a feeding method also used by adult herring on larger prey items like krill. If prey concentrations reach very high levels, as in microlayers, at fronts, or directly below the surface, herring become

filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...

s, driving several meters forward with wide open mouth and far expanded opercula, then closing and cleaning the

gill raker Gill rakers in fish are bony or cartilaginous processes that project from the branchial arch (gill arch) and are involved with suspension feeding tiny prey. They are not to be confused with the gill filaments that compose the fleshy part of the ...

s for a few milliseconds. Copepods, the primary zooplankton, are a major item on the forage fish menu. Copepods are typically long, with a teardrop-shaped body. Some scientists say they form the largest animal

biomass Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms bi ...

on the planet. Copepods are very alert and evasive. They have large antennae (see photo below left). When they spread their antennae, they can sense the pressure wave from an approaching fish and jump with great speed over a few centimetres. If copepod concentrations reach high levels, schooling herrings adopt a method called

ram feeding Aquatic feeding mechanisms face a special difficulty as compared to feeding on land, because the density of water is about the same as that of the prey, so the prey tends to be pushed away when the mouth is closed. This problem was first identifi ...

. In the photo below, herring ram feed on a school of copepods. They swim with their mouths wide open and their operculae fully expanded. The fish swim in a grid where the distance between them is the same as the jump length of their prey, as indicated in the animation above right. In the animation, juvenile herring hunt the copepods in this synchronised way. The copepods sense with their antennae the pressure wave of an approaching herring and react with a fast escape jump. The length of the jump is fairly constant. The fish align themselves in a grid with this characteristic jump length. A copepod can dart about 80 times before it tires. After a jump, it takes it 60 milliseconds to spread its antennae again, and this time delay becomes its undoing, as the almost endless stream of herring allows a herring to eventually snap up the copepod. A single juvenile herring could never catch a large copepod. Other pelagic prey eaten by herring includes fish eggs, larval

snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...

diatom A diatom (Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising sev ...

s by herring larvae below ,

tintinnid Tintinnids are ciliates of the choreotrich order Tintinnida, distinguished by vase-shaped shells, the name deriving from a Latin source meaning a small tinkling bell, that are called'' loricae'', which are mostly protein but may incorporate min ...

s by larvae below ,

mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is esti ...

an larvae,

larvae,

s, smaller fishes, pteropods,

annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...

s, ''

Calanus ''Calanus'' is a genus of marine copepod in the family Calanidae (Order Calanoida). The genus was split in 1974, with some species being placed in a new genus, '' Neocalanus''. The following species are recognised: *'' Calanus aculeatus'' Bra ...

'' spp.,

Centropagidae Centropagidae is a family of copepods in the order Calanoida. Its members are particularly known as plankton in coastal waters and in fresh water in Australia and southern South America. They are also found on subantarctic islands and in lakes i ...

, and ''

Meganyctiphanes norvegica Northern krill (''Meganyctiphanes norvegica'') is a species of krill that lives in the North Atlantic Ocean. It is an important component of the zooplankton, providing food for whales, seals, fish and birds. (In the Southern Ocean, Antarctic kril ...

''. Herrings, along with

Atlantic cod The Atlantic cod (''Gadus morhua'') is a benthopelagic fish of the family Gadidae, widely consumed by humans. It is also commercially known as cod or codling.sprat Sprat is the common name applied to a group of forage fish belonging to the genus ''Sprattus'' in the family Clupeidae. The term also is applied to a number of other small sprat-like forage fish (''Clupeoides'', ''Clupeonella'', '' Corica'', ' ...

, are the most important commercial species to humans in the Baltic Sea.Friedrich W. Köster, et al. "Developing Baltic Cod Recruitment Models. I. Resolving Spatial And Temporal Dynamics Of Spawning Stock And Recruitment For Cod, Herring, And Sprat." ''Canadian Journal of Fisheries & Aquatic Sciences'' 58.8 (2001): 1516. Academic Search Premier. Web. 21 Nov. 2011. p. 1516

/ref> The analysis of the stomach contents of these fish indicate Atlantic cod is the top predator, preying on the herring and sprat. Sprat are competitive with herring for the same food resources. This is evident in the two species'

vertical migration Diel vertical migration (DVM), also known as diurnal vertical migration, is a pattern of movement used by some organisms, such as copepods, living in the ocean and in lakes. The word ''diel'' comes from the Latin ''dies'' day, and means a 24- ...

in the Baltic Sea, where they compete for the limited zooplankton available and necessary for their survival.Casini, Michele, Cardinale, Massimiliano, and Arrheni, Fredrik. "Feeding preferences of herring (Clupea harengus) and sprat (Sprattus sprattus) in the southern Baltic Sea." ''ICES Journal of Marine Science'', 61 (2004): 1267–1277. Science Direct. Web. 22 November 2011. p. 1268

/ref> Sprat are highly selective in their diet and eat only zooplankton, while herring are more eclectic, adjusting their diet as they grow in size. In the Baltic, copepods of the genus ''

Acartia ''Acartia'' is a genus of marine calanoid copepods. They are epipelagic, estuarine, zooplanktonic found throughout the oceans of the world, primarily in temperate regions. Classification This genus contains the majority of species in the fam ...

'' can be present in large numbers. However, they are small in size with a high escape response, so herring and sprat avoid trying to catch them. These copepods also tend to dwell more in surface waters, whereas herring and sprat, especially during the day, tend to dwell in deeper waters.

Predators

Humpback whale lunging through a herring school

Predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...

s of herring include

seabirds Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine (ocean), marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ev ...

, marine mammals such as

dolphins A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (t ...

porpoises Porpoises are a group of fully aquatic marine mammals, all of which are classified under the family Phocoenidae, parvorder Odontoceti (toothed whales). Although similar in appearance to dolphins, they are more closely related to narwhals an ...

whales Whales are a widely distributed and diverse group of fully aquatic placental marine mammals. As an informal and colloquial grouping, they correspond to large members of the infraorder Cetacea, i.e. all cetaceans apart from dolphins and ...

seal Seal may refer to any of the following: Common uses * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impr ...

s, and

sea lion Sea lions are pinnipeds characterized by external ear flaps, long foreflippers, the ability to walk on all fours, short and thick hair, and a big chest and belly. Together with the fur seals, they make up the family Otariidae, eared seals. ...

s, predatory fish such as

sharks Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorp ...

billfish The term billfish refers to a group of saltwater predatory fish characterised by prominent pointed bills (rostra), and by their large size; some are longer than . Extant billfish include sailfish and marlin, which make up the family Istiophor ...

tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...

salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...

striped bass The striped bass (''Morone saxatilis''), also called the Atlantic striped bass, striper, linesider, rock, or rockfish, is an anadromous perciform fish of the family Moronidae found primarily along the Atlantic coast of North America. It has al ...

cod Cod is the common name for the demersal fish genus '' Gadus'', belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gadus'' is commonly not call ...

, and

halibut Halibut is the common name for three flatfish in the genera '' Hippoglossus'' and ''Reinhardtius'' from the family of right-eye flounders and, in some regions, and less commonly, other species of large flatfish. The word is derived from ''h ...

Fishermen A fisher or fisherman is someone who captures fish and other animals from a body of water, or gathers shellfish. Worldwide, there are about 38 million commercial and subsistence fishers and fish farmers. Fishers may be professional or recreat ...

also catch and eat herring. The predators often cooperate in groups, using different techniques to panic or herd a school of herring into a tight

bait ball A bait ball, or baitball, occurs when small fish swarm in a tightly packed spherical formation about a common centre. It is a last-ditch defensive measure adopted by small schooling fish when they are threatened by predators. Small schooling f ...

. Different predatory species then use different techniques to pick the fish off in the bait ball. The

sailfish The sailfish is one or two species of marine fish in the genus ''Istiophorus'', which belong to the family Istiophoridae (marlins). They are predominantly blue to gray in colour and have a characteristically large dorsal fin known as the sa ...

raises its sail to make it appear much larger.

Swordfish Swordfish (''Xiphias gladius''), also known as broadbills in some countries, are large, highly migratory predatory fish characterized by a long, flat, pointed bill. They are a popular sport fish of the billfish category, though elusive. Swordfis ...

charge at high speed through the bait balls, slashing with their swords to kill or stun prey. They then turn and return to consume their "catch".

Thresher shark Thresher sharks are large lamniform sharks of the family Alopiidae found in all temperate and tropical oceans of the world; the family contains three extant species, all within the genus ''Alopias''. All three thresher shark species have been ...

s use their long tails to stun the shoaling fish. These sharks compact their prey school by swimming around them and splashing the water with their tails, often in pairs or small groups. They then strike them sharply with the upper lobe of their tails to stun them.Seitz, J.C
Pelagic Thresher
. ''Florida Museum of Natural History.'' Retrieved on December 22, 2008.

Spinner shark The spinner shark (''Carcharhinus brevipinna'') is a species of requiem shark, in the family Carcharhinidae, named for the spinning leaps it makes as a part of its feeding strategy. This species occurs in tropical and warm temperate waters worldw ...

s charge vertically through the school, spinning on their axes with their mouths open and snapping all around. The sharks' momentum at the end of these spiraling runs often carries them into the air. Some whales

lunge feed A bait ball, or baitball, occurs when small fish swarm in a tightly packed spherical formation about a common centre. It is a last-ditch defensive measure adopted by small schooling fish when they are threatened by predators. Small schooling f ...

on bait balls. Lunge feeding is an extreme feeding method, where the whale accelerates from below the bait ball to a high velocity and then opens its mouth to a large gape angle. This generates the water pressure required to expand its mouth and engulf and filter a huge amount of water and fish. Lunge feeding by

rorqual Rorquals () are the largest group of baleen whales, which comprise the family Balaenopteridae, containing ten extant species in three genera. They include the largest animal that has ever lived, the blue whale, which can reach , and the fin wha ...

s, a family of huge baleen whales that includes the

blue whale The blue whale (''Balaenoptera musculus'') is a marine mammal and a baleen whale. Reaching a maximum confirmed length of and weighing up to , it is the largest animal known to have ever existed. The blue whale's long and slender body can ...

, is said to be the largest biomechanical event on Earth.

Fisheries

Adult herring are harvested for their flesh and eggs, and they are often used as

baitfish 300px, Feeder Goldfish are common baitfish. Bait fish (or baitfish) are small-sized fish caught and used by anglers as bait to attract larger predatory fish, particularly game fish. Baitfish species are typically those that are common and bree ...

. The trade in herring is an important sector of many national economies. In Europe, the fish has been called the "silver of the sea", and its trade has been so significant to many countries that it has been regarded as the most commercially important fishery in history. Purse seining for herring, southeast Alaska, nd (COBB 212)

Purse seining for herring, southeast Alaska, nd (COBB 212)

Environmental Defense Environmental Defense Fund or EDF (formerly known as Environmental Defense) is a United States-based nonprofit environmental advocacy group. The group is known for its work on issues including global warming, ecosystem restoration, oceans, and hu ...

have suggested that the

(''Clupea harengus'') fishery is an environmentally responsible fishery.

As food

Herring has been a

staple food A staple food, food staple, or simply a staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard diet for a given person or group of people, supplying a large fraction of energy needs and ...

source since at least 3000 BC. The fish is served numerous ways, and many regional recipes are used: eaten raw, fermented,

, or cured by other techniques, such as being smoked as

kipper A kipper is a whole herring, a small, oily fish, that has been split in a butterfly fashion from tail to head along the dorsal ridge, gutted, salted or pickled, and cold-smoked over smouldering wood chips (typically oak). In the United Ki ...

s. Herring are very high in the long-chain

omega-3 fatty acid Omega−3 fatty acids, also called Omega-3 oils, ω−3 fatty acids or ''n''−3 fatty acids, are polyunsaturated fatty acids (PUFAs) characterized by the presence of a double bond, three atoms away from the terminal methyl group in their chem ...

EPA The Environmental Protection Agency (EPA) is an independent executive agency of the United States federal government tasked with environmental protection matters. President Richard Nixon proposed the establishment of EPA on July 9, 1970; it be ...

and

DHA DHA, Dha and dha may refer to: Chemicals * Docosahexaenoic acid, a 22:6 omega-3 fatty acid * Dehydroandrosterone, an endogenous androgenic steroid * Dehydroascorbic acid, an oxidized form of ascorbic acid * Dehydroacetic acid, a pyrone derivati ...

. They are a source of

vitamin D Vitamin D is a group of fat-soluble secosteroids responsible for increasing intestinal absorption of calcium, magnesium, and phosphate, and many other biological effects. In humans, the most important compounds in this group are vitamin D3 (c ...

Water pollution Water pollution (or aquatic pollution) is the contamination of water bodies, usually as a result of human activities, so that it negatively affects its uses. Water bodies include lakes, rivers, oceans, aquifers, reservoirs and groundwater. Water ...

influences the amount of herring that may be safely consumed. For example, large Baltic herring slightly exceeds recommended limits with respect to

PCB PCB may refer to: Science and technology * Polychlorinated biphenyl, an organic chlorine compound, now recognized as an environmental toxin and classified as a persistent organic pollutant * Printed circuit board, a board used in electronics * ...

and

dioxin Dioxin may refer to: * 1,2-Dioxin or 1,4-Dioxin, two unsaturated heterocyclic 6-membered rings where two carbon atoms have been replaced by oxygen atoms, giving the molecular formula C4H4O2 *Dibenzo-1,4-dioxin, the parent compound also known as ...

, although some sources point out that the cancer-reducing effect of omega-3 fatty acids is statistically stronger than the

carcinogen A carcinogen is any substance, radionuclide, or radiation that promotes carcinogenesis (the formation of cancer). This may be due to the ability to damage the genome or to the disruption of cellular metabolic processes. Several radioactive substan ...

ic effect of PCBs and dioxins. The contaminant levels depend on the age of the fish which can be inferred from their size. Baltic herrings larger than may be eaten twice a month, while herrings smaller than 17 cm can be eaten freely.

Mercury in fish The presence of mercury in fish is a health concern for people who eat them, especially for women who are or may become pregnant, nursing mothers, and young children. Fish and shellfish concentrate mercury in their bodies, often in the form of m ...

also influences the amount of fish that women who are pregnant or planning to be pregnant within the next one or two years may safely eat.

History

The herring has played a highly significant role in history both socially and economically. During the Middle Ages, herring prompted the founding of

Great Yarmouth Great Yarmouth (), often called Yarmouth, is a seaside town and unparished area in, and the main administrative centre of, the Borough of Great Yarmouth in Norfolk, England; it straddles the River Yare and is located east of Norwich. A pop ...

Amsterdam Amsterdam ( , , , lit. ''The Dam on the River Amstel'') is the Capital of the Netherlands, capital and Municipalities of the Netherlands, most populous city of the Netherlands, with The Hague being the seat of government. It has a population ...

, and

Copenhagen Copenhagen ( or .; da, København ) is the capital and most populous city of Denmark, with a proper population of around 815.000 in the last quarter of 2022; and some 1.370,000 in the urban area; and the wider Copenhagen metropolitan ar ...

. In 1274, while on his deathbed at the monastery of Fossanova (south of Rome, Italy), when encouraged to eat something to regain his strength,

Thomas Aquinas Thomas Aquinas, OP (; it, Tommaso d'Aquino, lit=Thomas of Aquino; 1225 – 7 March 1274) was an Italian Dominican friar and priest who was an influential philosopher, theologian and jurist in the tradition of scholasticism; he is known wi ...

asked for fresh herring.Process of Canonization of St. Thomas Aquinas, Testimony of Br. Peter of Montesangiovanni
/ref>

References

Citations

Sources

* * Dewhurst HW (1834
''Clupea harengis'' or the common herring
In: ''The Natural History of the Order Cetacea'', Oxford University. Pages 232–246. * Geffen, Audrey J (2009
Advances in herring biology: from simple to complex, coping with plasticity and adaptability:
''ICES Journal of Marine Science'', 66 (8): 1688–1695. * Gilpen JB (1867
"On the common herring (''Clupea elongata'')"
''Proceedings and Transactions of the Nova-Scotian Institute of Natural Science'', 1 (1): 4–11. * O'Clair, Rita M. and O'Clair, Charles E., "Pacific herring," ''Southeast Alaska's Rocky Shores: Animals''. pg. 343–346. Plant Press: Auke Bay, Alaska (1998). * Stephenson RL (2001
The role of herring investigations in shaping fisheries science
In F. Funk, J. Blackburn, D. Hay, A.J. Paul, R. Stephen- son, R. Toresen, and D. Witherell (eds.) ''Herrings: Expectations for a New Millennium'', Alaska Sea Grant College Program. AK-SG-01-04. pp. 1–20. . * Stephenson, R. L., Melvin, G. D., and Power, M. J. (2009
"Population integrity and connectivity in Northwest Atlantic herring: a review of assumptions and evidence"
''ICES Journal of Marine Science'', 66: 1733–1739. * Whitehead PJP, Nelson GJ and Wongratana T (1988) FAO species catalogue, volume 2: ''Clupeoid Fishes of the World''

FAO Fisheries Synopsis 125, Rome. . tp://ftp.fao.org/docrep/fao/009/t0835e/t0835e.zip Download ZIP (16 MB)

External links

from National Geographic (2003)
Atlantic Herring
from the Gulf of Maine Research Institute
Nutrition Facts for Herring

Prospecting herring waste – from ScienceNordic

PNAS Population-scale sequencing reveals genetic differentiation due to local adaptation in Atlantic herring.
{{Authority control Clupeidae Seafood Commercial fish Oily fish Fish common names

Species

Characteristics

Lifecycle

Ecology

Prey

Predators

Fisheries

As food

History

See also

References

Citations

Sources

Further reading

External links