Hemiphractus Panamensis
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''Hemiphractus'' is a genus of frogs, the horned treefrogs, in the family Hemiphractidae. These overall brownish frogs have a pointed snout and a casque on the head. They are
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
, relatively rare and native to humid lowland and highland forests in northern South America ( Amazon east as far as Tapajós, Chocó-Magdalena and northern Andes) and Panama, where typically found on the ground or at relatively low levels in vegetation.


Description

''Hemiphractus'' are robust-bodied frogs. The genus is characterized by a fleshy proboscis on the tip of the snout and fleshy tubercles on the eyelids, skull that is highly casqued with prominent lateral occipital processes projecting backwards, and fang-like maxillary and premaxillary teeth. Females grow larger than males, and depending on exact species the snout–to–vent length generally is between .


Behaviour

''Hemiphractus'' are believed to be specialized predators of other frogs, and hence confined to areas with high density of frogs. Nevertheless, they have also been recorded feeding on lizards, large insects and snails in the wild, and in addition to other frogs captive ''Hemiphractus'' will eat geckos, newborn mice and earthworms. They can threaten any would-be predator by opening their mouth, exposing the bright yellow inside and tongue. If this fails to scare an adversary, they willingly bite, which is relatively painful for a frog of their size. Female frogs carry eggs openly on their back; the eggs adhere to the mother's back with gelatinous material. Eggs hatch as fully developed froglets. ''H. elioti'' (previously included in ''H. fasciatus'') are bred and maintained by the El Valle Amphibian Conservation Center in Panama. Although they have proven relatively difficult to keep in captivity, keeping and breeding has been achieved at levels where its conservation through such measures is considered possible, although (as of 2015 where the total captive population was 36 individuals) additional founders are necessary.


Species

The following species are recognized in the genus ''Hemiphractus ''. * ''
Hemiphractus bubalus ''Hemiphractus bubalus'', or the Ecuador horned tree frog, is a species of frog in the family Hemiphractidae. It is found in the upper Amazon Basin and lower Amazonian slopes of the Andes in northern Peru, Ecuador, and southern Colombia ( Cordill ...
'' (Jiménez de la Espada, 1871) * ''
Hemiphractus elioti ''Hemiphractus'' is a genus of frogs, the horned treefrogs, in the family Hemiphractidae. These overall brownish frogs have a pointed snout and a casque on the head. They are nocturnal, relatively rare and native to humid lowland and highland for ...
'' Hill, Martin, Stanley, and Mendelson, 2018 * '' Hemiphractus fasciatus'' Peters, 1862 * ''
Hemiphractus helioi ''Hemiphractus helioi'' is a species of frog in the family Hemiphractidae. It is found in the upper Amazon basin and lower Amazonian slopes of the Andes from Ecuador to southern Peru and into adjacent Brazil (Acre). There is also a disputed recor ...
'' Sheil & Mendelson, 2001 * ''
Hemiphractus johnsoni ''Hemiphractus johnsoni'', or the Johnson's horned treefrog, is a species of frog in the family Hemiphractidae. It is endemic to Colombia and known from the Cordillera Central in Antioquia and Caldas Departments as well as from the Cordillera ...
'' (Noble, 1917) * ''
Hemiphractus kaylockae ''Hemiphractus'' is a genus of frogs, the horned treefrogs, in the family Hemiphractidae. These overall brownish frogs have a pointed snout and a casque on the head. They are nocturnal, relatively rare and native to humid lowland and highland for ...
'' Hill, Martin, Stanley, and Mendelson, 2018 * '' Hemiphractu panamensis'' (Stejneger, 1917) * ''
Hemiphractus proboscideus ''Hemiphractus proboscideus'', or the Sumaco horned treefrog, is a species of frog in the family Hemiphractidae. It is found in the upper Amazon basin in extreme southwestern Colombia ( Amazonas and Caquetá Departments), Ecuador, and northern Pe ...
'' (Jiménez de la Espada, 1871) * ''
Hemiphractus scutatus ''Hemiphractus scutatus'', also known as Spix's horned treefrog, is a species of frog in the family Hemiphractidae. It is found in the upper Amazon basin in Bolivia, western Brazil, Peru, Ecuador, and southern Colombia. Its natural habitat is t ...
'' (Spix, 1824) Until 2001, populations now recognized as ''H. helioi'' were typically included in ''H. johnsoni'', and until 2018 populations now recognized as ''H. elioti'', ''H. kaylockae'' and ''H. panamensis'' were typically included in ''H. fasciatus''. Colombian "''H. fasciatus''" may represent a currently unrecognized species (thus limiting true ''H. fasciatus'' to Ecuador), and it is possible that ''H. scutatus'' is a species complex.


References

{{Taxonbar, from=Q510491 Hemiphractidae Amphibians of South America Amphibians of Central America Amphibian genera Taxa named by Johann Georg Wagler