Harpymimus Steveoc
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''Harpymimus'' is a basal ornithomimosaurian
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the Early Cretaceous
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
of what is now
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
. Unlike later, more derived ornithomimosaurs, ''Harpymimus'' still possessed teeth, although they appear to have been restricted to the
dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
of the lower jaw.


Discovery and naming

In 1981, a Soviet-Mongolian expedition uncovered a theropod skeleton in the Gobi Desert. In 1984 this was named and shortly described by
Rinchen Barsbold , Rinchyengiin Barsbold, born December 21, 1935 in Ulaanbaatar) is a Mongolian paleontologist and geologist. He works with the Institute of Geology, at Ulaanbaatar, Mongolia. He is an expert in vertebrate paleontology and Mesozoic stratigraphy. Bar ...
and Altangerel Perle as the type and only
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of the new
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Harpymimus'': ''Harpymimus okladnikovi''. The generic name ''Harpymimus'' is a reference to the fearsome Harpy of
Greek mythology A major branch of classical mythology, Greek mythology is the body of myths originally told by the ancient Greeks, and a genre of Ancient Greek folklore. These stories concern the origin and nature of the world, the lives and activities ...
and derived from Greek ἅρπυια (''harpyia''), "Harpy", and μῖμος (''mimos''), "mimic". The specific name honours the late Soviet
archeologist Archaeology or archeology is the scientific study of human activity through the recovery and analysis of material culture. The archaeological record consists of artifacts, architecture, biofacts or ecofacts, sites, and cultural landscap ...
Alexey Pavlovich Okladnikov. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
specimen IGM 100/29 (Mongolian Academy of Sciences,
Ulan Bator Ulaanbaatar (; mn, Улаанбаатар, , "Red Hero"), previously anglicized as Ulan Bator, is the capital and most populous city of Mongolia. It is the coldest capital city in the world, on average. The municipality is located in north ce ...
, Mongolia) consists of an almost complete and articulated but compressed skeleton, lacking only portions of the pectoral girdle, pelvic girdle, and hindlimbs. It was recovered in the Dundgovi Aimag (Eastern
Gobi The Gobi Desert ( Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world. Geography The Gobi measures from southwest to northeast ...
Province), from an exposure of the Shinekhudag Formation now part of the Khuren Dukh Formation which dates to the Mid-Late
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0 ...
Barsbold, R. and Perle, A. (1984). n first new find of a primitive orithomimosaur from the Cretaceous of the MPR ''Paleontologicheskii zhurnal'', 2: 121-123 Other
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s collected from the Shinekhudug Formation in Dundgovi include the ceratopsian '' Psittacosaurus mongoliensis''.


Description

''Harpymimus'' was for the first time extensively described in a dissertation by
Yoshitsugu Kobayashi {{Short pages monitor