''Hanssuesia'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

pachycephalosaurid Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachycephal ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

s from the late

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

period. It lived in what is now

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

and

Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...

, and contains the single species ''Hanssuesia sternbergi''.

''Hanssuesia'' is based on a skull dome originally named ''

Troodon ''Troodon'' ( ; ''Troödon'' in older sources) is a wastebasket taxon and a dubious genus of relatively small, bird-like dinosaurs known definitively from the Campanian age of the Late Cretaceous period (about 77 mya). It includes at least ...

sternbergi'' by

Barnum Brown Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. Named after the circus showman P. T. Barnum, he discovered the first documented remains of ''Tyrannosaurus'' during a career ...

and

Erich Maren Schlaikjer Erich Maren Schlaikjer (; November 22, 1905 in Newtown, Ohio – November 5, 1972) was an American geologist and dinosaur hunter. Assisting Barnum Brown, he co-described ''Pachycephalosaurus'' and what is now ''Montanoceratops''. Other discoveries ...

in 1943. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

honoured

Charles Mortram Sternberg Charles Mortram Sternberg (1885–1981) was an American-Canadian fossil collector and paleontologist, son of Charles Hazelius Sternberg. Late in his career, he collected and described ''Pachyrhinosaurus'', ''Brachylophosaurus'', ''Parksosaurus ...

who found the dome in 1928 near Steveville in south Alberta. In 1945, it was transferred to ''Stegoceras'' by C.M. Sternberg himself, as a ''Stegoceras sternbergi''. The genus ''Hanssuesia'' was first named by Robert M. Sullivan in

2003 File:2003 Events Collage.png, From top left, clockwise: The crew of STS-107 perished when the Space Shuttle Columbia disintegrated during reentry into Earth's atmosphere; SARS became an epidemic in China, and was a precursor to SARS-CoV-2; A des ...

. It honours paleontologist

Hans-Dieter Sues Hans-Dieter Sues (born January 13, 1956) is a German-born American paleontologist who is Senior Scientist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC. He receiv ...

. In the publication also the spelling variant "Hanssuessia" appeared; the same year Sullivan chose for ''Hanssuesia'' as the valid name. Its

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Troodon sternbergi'', the ''

combinatio nova ''Combinatio nova'', abbreviated ''comb. nov.'' (sometimes ''n. comb.''), is Latin for "new combination". It is used in taxonomic biology literature when a new name is introduced based on a pre-existing name. The term should not to be confused wi ...

'' is ''Hanssuesia sternbergi''. ''H. sternbergi'' is known from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

NMC 8817 and six referred specimens (mainly frontoparietals) which were collected from two formations of the

Belly River Group The Belly River Group is a stratigraphical unit of Late Cretaceous age in the Western Canadian Sedimentary Basin. It takes the name from the Belly River, a tributary of the Oldman River in southern Alberta, and was first described in outcrop o ...

of Alberta, Canada, the

Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...

(late

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

, 76.5–75 ma) in the

Dinosaur Provincial Park Dinosaur Provincial Park is a UNESCO World Heritage Site situated a two hour drive east of Calgary, Alberta, Canada; or , about a half-hour drive northeast of Brooks. The park is situated in the Red Deer River valley, which is noted for its strik ...

, and the

Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...

(middle Campanian, 77.5-76.5 ma), as well as from the upper

Judith River Formation The Judith River Formation is a fossil-bearing geologic formation in Montana, and is part of the Judith River Group. It dates to the Late Cretaceous, between 79 and 75.3 million years ago, corresponding to the "Judithian" land vertebrate age. It ...

of Montana, United States (late

, 76–75 ma). Like other pachycephalosaurs, ''Hanssuesia'' had a thick

skull roof The skull roof, or the roofing bones of the skull, are a set of bones covering the brain, eyes and nostrils in bony fishes and all land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In comparati ...

. However, ''Hanssuesia'' is distinguished from other pachycephalosaurs by having a depressed parietal region, a frontoparietal dome that is wide in front as well as in the rear, a broad nasal boss on the frontals, reduced but swollen prefrontal lobes, and a reduced parietosquamosal shelf at the dome rear. A 2020 CSVP abstract found ''

Gravitholus ''Gravitholus'' (meaning 'heavy dome') was a genus of pachycephalosaurid dinosaur from the late Cretaceous period (Campanian stage, around 75 million years ago). It was a pachycephalosaur, and like other pachycephalosaurids the skull roof formed ...

'' and ''Hanssuesia'' to be synonymous with ''Stegoceras.''

References

Late Cretaceous dinosaurs of North America Fossil taxa described in 2003 Pachycephalosaurs Paleontology in Alberta Paleontology in Montana Campanian genus first appearances Campanian genus extinctions Ornithischian genera {{pachycephalosaur-stub

See also

References