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Hair is a protein filament that grows from follicles found in the
dermis The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided i ...
. Hair is one of the defining characteristics of
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s. The
human body The human body is the structure of a Human, human being. It is composed of many different types of Cell (biology), cells that together create Tissue (biology), tissues and subsequently organ systems. They ensure homeostasis and the life, viabi ...
, apart from areas of glabrous skin, is covered in follicles which produce thick
terminal Terminal may refer to: Computing Hardware * Terminal (electronics), a device for joining electrical circuits together * Terminal (telecommunication), a device communicating over a line * Computer terminal, a set of primary input and output devic ...
and fine vellus hair. Most common interest in hair is focused on hair growth, hair types, and
hair care Hair care is an overall term for hygiene and cosmetology involving the hair which grows from the human scalp, and to a lesser extent facial, pubic and other body hair. Hair care routines differ according to an individual's culture and the physic ...
, but hair is also an important biomaterial primarily composed of protein, notably alpha-keratin. Attitudes towards different forms of hair, such as hairstyles and hair removal, vary widely across different cultures and historical periods, but it is often used to indicate a person's personal beliefs or social position, such as their age, sex, or religion.


Overview

The word "hair" usually refers to two distinct structures: #the part beneath the skin, called the
hair follicle The hair follicle is an organ found in mammalian skin. It resides in the dermal layer of the skin and is made up of 20 different cell types, each with distinct functions. The hair follicle regulates hair growth via a complex interaction between h ...
, or, when pulled from the skin, the bulb or root. This organ is located in the
dermis The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided i ...
and maintains
stem cell In multicellular organisms, stem cells are undifferentiated or partially differentiated cells that can differentiate into various types of cells and proliferate indefinitely to produce more of the same stem cell. They are the earliest type o ...
s, which not only re-grow the hair after it falls out, but also are recruited to regrow skin after a
wound A wound is a rapid onset of injury that involves laceration, lacerated or puncture wound, punctured skin (an ''open'' wound), or a bruise, contusion (a ''closed'' wound) from blunt force physical trauma, trauma or compression. In pathology, a '' ...
. # the hair shaft, which is the hard filamentous part that extends above the skin surface. A cross section of the hair shaft may be divided roughly into three zones. Hair fibers have a structure consisting of several layers, starting from the outside: # the
cuticle A cuticle (), or cuticula, is any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticle" are non- homologous, differing in their origin, structu ...
, which consists of several layers of flat, thin cells laid out overlapping one another as roof shingles # the cortex, which contains the keratin bundles in cell structures that remain roughly rod-like # the medulla, a disorganized and open area at the fiber's center


Description

Each strand of hair is made up of the medulla, cortex, and
cuticle A cuticle (), or cuticula, is any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticle" are non- homologous, differing in their origin, structu ...
.Hair Structure and Hair Life Cycle
follicle.com
The innermost region, the medulla, is an open and unstructured region which is not always present. The highly structural and organized cortex, or second of three layers of the hair, is the primary source of mechanical strength and water uptake. The cortex contains melanin, which colors the fiber based on the number, distribution and types of melanin granules. The shape of the follicle determines the shape of the cortex, and the shape of the fiber is related to how straight or curly the hair is. People with straight hair have round hair fibers. Oval and other shaped fibers are generally more wavy or curly. The cuticle is the outer covering. Its complex structure slides as the hair swells and is covered with a single molecular layer of lipid that makes the hair repel water. The diameter of human hair varies from . There are roughly two million small, tubular glands and
sweat gland Sweat glands, also known as sudoriferous or sudoriparous glands, , are small tubular structures of the skin that produce sweat. Sweat glands are a type of exocrine gland, which are glands that produce and secrete substances onto an epithelial sur ...
s that produce watery fluids that cool the body by evaporation. The glands at the opening of the hair produce a fatty secretion that lubricates the hair. Hair growth begins inside the
hair follicle The hair follicle is an organ found in mammalian skin. It resides in the dermal layer of the skin and is made up of 20 different cell types, each with distinct functions. The hair follicle regulates hair growth via a complex interaction between h ...
. The only "living" portion of the hair is found in the follicle. The hair that is visible is the hair shaft, which exhibits no biochemical activity and is considered "dead". The base of a hair's root (the "bulb") contains the cells that produce the hair shaft. Other structures of the hair follicle include the oil producing sebaceous gland which lubricates the hair and the
arrector pili The arrector pili muscles, also known as hair erector muscles, are small muscles attached to hair follicles in mammals. Contraction of these muscles causes the hairs to stand on end, known colloquially as goose bumps (piloerection). Structure E ...
muscles, which are responsible for causing hairs to stand up. In humans with little body hair, the effect results in goose bumps.


Root of the hair

The ''root of the hair'' ends in an enlargement, the ''hair bulb'', which is whiter in color and softer in texture than the shaft and is lodged in a follicular involution of the
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water rele ...
called the
hair follicle The hair follicle is an organ found in mammalian skin. It resides in the dermal layer of the skin and is made up of 20 different cell types, each with distinct functions. The hair follicle regulates hair growth via a complex interaction between h ...
. The bulb of hair consists of fibrous connective tissue, glassy membrane, external root sheath, internal root sheath composed of epithelium stratum (
Henle's layer Henle's layer is the third and the outermost layer of the inner root sheath of the hair follicle, consisting of a single layer of cubical cells with clear flattened nuclei. It is named after Germany, German physician, pathologist and anatomist Frie ...
) and granular stratum (
Huxley's layer Huxley's layer is the second layer of the inner root sheath of the hair and consists of one or two layers of horny, flattened, nucleated cells. It lies between Henle's layer and the cuticle. The layer is named after English biologist Thomas Henry ...
), cuticle, cortex and medulla.


Natural color

All natural hair colors are the result of two types of hair pigments. Both of these pigments are melanin types, produced inside the hair follicle and packed into granules found in the fibers.
Eumelanin Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amin ...
is the dominant pigment in brown hair and black hair, while pheomelanin is dominant in
red hair Red hair (also known as orange hair and ginger hair) is a hair color found in one to two percent of the human population, appearing with greater frequency (two to six percent) among people of Northern or Northwestern European ancestry and ...
. Blond hair is the result of having little pigmentation in the hair strand. Gray hair occurs when melanin production decreases or stops, while poliosis is white hair (and often the skin to which the hair is attached), typically in spots that never possessed melanin at all, or ceased for natural reasons, generally genetic, in the first years of life.


Human hair growth

Hair grows everywhere on the external body except for mucus membranes and glabrous skin, such as that found on the palms of the hands, soles of the feet, and lips. Hair follows a specific growth cycle with three distinct and concurrent phases:
anagen The hair follicle is an organ found in mammalian skin. It resides in the dermal layer of the skin and is made up of 20 different cell types, each with distinct functions. The hair follicle regulates hair growth via a complex interaction between h ...
,
catagen The growth of human hair occurs everywhere on the body except for the soles of the feet, the inside of the mouth, the lips, the backs of the ears, the palms of the hands, some external genital areas, the navel, scar tissue, and, apart from eyelas ...
, and
telogen The hair follicle is an organ found in mammalian skin. It resides in the dermal layer of the skin and is made up of 20 different cell types, each with distinct functions. The hair follicle regulates hair growth via a complex interaction between ho ...
phases; all three occur simultaneously throughout the body. Each has specific characteristics that determine the length of the hair. The body has different types of hair, including vellus hair and androgenic hair, each with its own type of cellular construction. The different construction gives the hair unique characteristics, serving specific purposes, mainly, warmth and protection.


Texture

Hair exists in a variety of textures. Three main aspects of hair texture are the curl pattern, volume, and consistency. All mammalian hair is composed of keratin, so the make-up of
hair follicles The hair follicle is an organ found in mammalian skin. It resides in the dermal layer of the skin and is made up of 20 different cell types, each with distinct functions. The hair follicle regulates hair growth via a complex interaction between h ...
is not the source of varying hair patterns. There are a range of theories pertaining to the curl patterns of hair. Scientists have come to believe that the shape of the hair shaft has an effect on the curliness of the individual's hair. A very round shaft allows for fewer disulfide bonds to be present in the hair strand. This means the bonds present are directly in line with one another, resulting in straight hair. The flatter the hair shaft becomes, the curlier hair gets, because the shape allows more
cysteine Cysteine (symbol Cys or C; ) is a semiessential proteinogenic amino acid with the formula . The thiol side chain in cysteine often participates in enzymatic reactions as a nucleophile. When present as a deprotonated catalytic residue, sometime ...
s to become compacted together resulting in a bent shape that, with every additional disulfide bond, becomes curlier in form. As the hair follicle shape determines curl pattern, the hair follicle size determines thickness. While the circumference of the hair follicle expands, so does the thickness of the hair follicle. An individual's hair volume, as a result, can be thin, normal, or thick. The consistency of hair can almost always be grouped into three categories: fine, medium, and coarse. This trait is determined by the hair follicle volume and the condition of the strand. Fine hair has the smallest circumference, coarse hair has the largest circumference, and medium hair is anywhere between the other two. Coarse hair has a more open cuticle than thin or medium hair causing it to be the most porous.


Classification systems

There are various systems that people use to classify their curl patterns. Being knowledgeable of an individual's hair type is a good start to knowing how to take care of one's hair. There is not just one method to discovering one's hair type. Additionally it is possible, and quite normal to have more than one kind of hair type, for instance having a mixture of both type 3a & 3b curls. ; The
Andre Walker Hair Typing System The Andre Walker Hair Typing System, also known as The Hair Chart is classification system for hair types created in the 1990s by Oprah Winfrey's stylist Andre Walker. It was originally created to market Walker's line of hair care products but has ...
is the most widely used system to classify hair. The system was created by the hairstylist of
Oprah Winfrey Oprah Gail Winfrey (; born Orpah Gail Winfrey; January 29, 1954), or simply Oprah, is an American talk show host, television producer, actress, author, and philanthropist. She is best known for her talk show, ''The Oprah Winfrey Show'', br ...
,
Andre Walker Andre Walker (born August 19, 1956 in Chicago, Illinois) is a hairstylist from the United States who has won seven Daytime Emmy Awards for Outstanding Achievement in Hairstyling for his work on ''The Oprah Winfrey Show''. He is best known for being ...
. According to this system there are four types of hair: straight, wavy, curly, kinky. * Type 1 is straight hair, which reflects the most sheen and also the most resilient hair of all of the hair types. It is hard to damage and immensely difficult to curl this hair texture. Because the sebum easily spreads from the scalp to the ends without curls or kinks to interrupt its path, it is the most oily hair texture of all. * Type 2 is wavy hair, whose texture and sheen ranges somewhere between straight and curly hair. Wavy hair is also more likely to become frizzy than straight hair. While type A waves can easily alternate between straight and curly styles, type B and C Wavy hair is resistant to styling. * Type 3 is
curly hair Curly is a surname, given name, nickname or stage name. It may refer to: First name, nickname or stage name * Crazy Horse (1840–1877), Oglala Sioux war chief nicknamed "Curly" * Curly (scout), nickname of Ashishishe (c. 1856–1923), Crow Ind ...
known to have an S-shape. The curl pattern may resemble a lowercase "s", uppercase "S", or sometimes an uppercase "Z" or lowercase "z". Lack of proper care causes less defined curls. * Type 4 is
kinky hair Afro-textured hair, or kinky hair, is a human hair texture originating from sub-Saharan Africa. Each strand of this hair type grows in a tiny, angle-like helix shape. The overall effect is such that, contrasted with straight, wavy, or curly ha ...
, which features a tightly coiled curl pattern (or no discernible curl pattern at all) that is often fragile with a very high density. This type of hair shrinks when wet and because it has fewer
cuticle A cuticle (), or cuticula, is any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticle" are non- homologous, differing in their origin, structu ...
layers than other hair types it is more susceptible to damage. ; This is a method which classifies the hair by curl pattern, hair-strand thickness and overall hair volume.


Functions

Many mammals have fur and other hairs that serve different functions. Hair provides thermal regulation and camouflage for many animals; for others it provides signals to other animals such as warnings, mating, or other communicative displays; and for some animals hair provides defensive functions and, rarely, even offensive protection. Hair also has a sensory function, extending the sense of touch beyond the surface of the skin. Guard hairs give warnings that may trigger a recoiling reaction.


Warmth

While humans have developed clothing and other means of keeping warm, the hair found on the head serves primarily as a source of heat
insulation Insulation may refer to: Thermal * Thermal insulation, use of materials to reduce rates of heat transfer ** List of insulation materials ** Building insulation, thermal insulation added to buildings for comfort and energy efficiency *** Insulated ...
and cooling (when sweat evaporates from soaked hair) as well as protection from ultra-violet radiation exposure. The function of hair in other locations is debated. Hats and coats are still required while doing outdoor activities in cold weather to prevent frostbite and hypothermia, but the hair on the human body does help to keep the internal temperature regulated. When the body is too cold, the
arrector pili The arrector pili muscles, also known as hair erector muscles, are small muscles attached to hair follicles in mammals. Contraction of these muscles causes the hairs to stand on end, known colloquially as goose bumps (piloerection). Structure E ...
muscles found attached to hair follicles stand up, causing the hair in these follicles to do the same. These hairs then form a heat-trapping layer above the
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water rele ...
. This process is formally called piloerection, derived from the Latin words 'pilus' ('hair') and 'erectio' ('rising up'), but is more commonly known as 'having goose bumps' in English. This is more effective in other mammals whose fur fluffs up to create air pockets between hairs that insulate the body from the cold. The opposite actions occur when the body is too warm; the arrector muscles make the hair lie flat on the skin which allows heat to leave.


Protection

In some mammals, such as hedgehogs and
porcupine Porcupines are large rodents with coats of sharp spines, or quills, that protect them against predation. The term covers two families of animals: the Old World porcupines of family Hystricidae, and the New World porcupines of family, Erethizont ...
s, the hairs have been modified into hard spines or quills. These are covered with thick plates of keratin and serve as protection against predators. Thick hair such as that of the lion's mane and grizzly bear's fur do offer some protection from physical damages such as bites and scratches.


Touch sense

Displacement and vibration of hair shafts are detected by hair follicle nerve receptors and nerve receptors within the skin. Hairs can sense movements of air as well as touch by physical objects and they provide sensory awareness of the presence of ectoparasites. Some hairs, such as eyelashes, are especially sensitive to the presence of potentially harmful matter.


Eyebrows and eyelashes

The eyebrows provide moderate protection to the
eye Eyes are organs of the visual system. They provide living organisms with vision, the ability to receive and process visual detail, as well as enabling several photo response functions that are independent of vision. Eyes detect light and conv ...
s from dirt, sweat and rain. They also play a key role in non-verbal communication by displaying emotions such as sadness, anger, surprise and excitement. In many other mammals, they contain much longer, whisker-like hairs that act as tactile sensors. The
eyelash An eyelash (also called lash) (Latin: ''Cilia'') is one of the hairs that grows at the edge of the eyelids. It grows in one layer on the edge of the upper and lower eyelids. Eyelashes protect the eye from debris, dust, and small particles and p ...
grows at the edges of the eyelid and protects the eye from dirt. The eyelash is to humans, camels, horses, ostriches etc., what
whiskers Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to touch, sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are c ...
are to cats; they are used to sense when dirt, dust, or any other potentially harmful object is too close to the eye. The eye reflexively closes as a result of this sensation.


Evolution

Hair has its origins in the common ancestor of mammals, the synapsids, about 300 million years ago. It is currently unknown at what stage the synapsids acquired mammalian characteristics such as
body hair Body hair, or androgenic hair, is the terminal hair that develops on the human body during and after puberty. It is differentiated from the head hair and less visible vellus hair, which is much finer and lighter in color. The growth of androge ...
and mammary glands, as the fossils only rarely provide direct evidence for soft tissues. Skin impression of the belly and lower tail of a pelycosaur, possibly '' Haptodus'' shows the basal synapsid stock bore transverse rows of rectangular scutes, similar to those of a modern
crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...
, so the age of acquirement of hair logically couldn't have been earlier than ~299 ma, based on the current understanding of the animal's phylogeny. An exceptionally well-preserved skull of ''
Estemmenosuchus ''Estemmenosuchus'' (meaning "crowned crocodile" in Greek) is an extinct genus of large, early omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two ...
'', a therapsid from the
Upper Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
, shows smooth, hairless skin with what appears to be glandular depressions, though as a semi-aquatic species it might not have been particularly useful to determine the integument of terrestrial species. The oldest undisputed known fossils showing unambiguous imprints of hair are the Callovian (late middle Jurassic) ''
Castorocauda ''Castorocauda'' is an extinct, semi-aquatic, superficially otter-like genus of docodont mammaliaforms with one species, ''C. lutrasimilis''. It is part of the Yanliao Biota, found in the Daohugou Beds of Inner Mongolia, China dating to the M ...
'' and several contemporary
haramiyida Haramiyida ("thief" from Arabic الحرامية (al ḥarāmiyah), "thief, bandit") is a possibly polyphyletic order of mammaliaform cynodonts or mammals of controversial taxonomic affinites. Their teeth, which are by far the most common remains ...
ns, both near-mammal
cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s, giving the age as no later than ~220 ma based on the modern phylogenetic understanding of these clades. See also the news item at More recently, studies on terminal Permian Russian coprolites may suggest that non-mammalian synapsids from that era had fur. If this is the case, these are the oldest hair remnants known, showcasing that fur occurred as far back as the latest Paleozoic. Some modern mammals have a special gland in front of each orbit used to preen the fur, called the harderian gland. Imprints of this structure are found in the skull of the small early mammals like ''
Morganucodon ''Morganucodon'' (" Glamorgan tooth") is an early mammaliaform genus that lived from the Late Triassic to the Middle Jurassic. It first appeared about 205 million years ago. Unlike many other early mammaliaforms, ''Morganucodon'' is well represe ...
'', but not in their
cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
ancestors like '' Thrinaxodon''. The hairs of the fur in modern animals are all connected to nerves, and so the fur also serves as a transmitter for sensory input. Fur could have evolved from sensory hair (whiskers). The signals from this sensory apparatus is interpreted in the
neocortex The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, sp ...
, a section of the brain that expanded markedly in animals like ''Morganucodon'' and ''
Hadrocodium ''Hadrocodium wui'' is an extinct mammaliaform that lived during the Sinemurian stage of the Early Jurassic approximately in the Lufeng Formation of the Lufeng Basin in what is now the Yunnan province in south-western China (, paleocoordinates ...
''. The more advanced therapsids could have had a combination of naked skin,
whisker Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarser ...
s, and scutes. A full pelage likely did not evolve until the therapsid-mammal transition. The more advanced, smaller therapsids could have had a combination of hair and scutes, a combination still found in some modern mammals, such as
rodents Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are nat ...
and the
opossum Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 93 species in 18 genera. Opossums originated in South America and entered North ...
. The high interspecific variability of the size, color, and microstructure of hair often enables the identification of species based on single hair filaments. In varying degrees most
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s have some skin areas without natural hair. On the human body, glabrous skin is found on the ventral portion of the fingers, palms, soles of feet and lips, which are all parts of the body most closely associated with interacting with the world around us, as are the labia minora and glans penis. There are four main types of
mechanoreceptor A mechanoreceptor, also called mechanoceptor, is a sensory receptor that responds to mechanical pressure or distortion. Mechanoreceptors are innervated by sensory neurons that convert mechanical pressure into electrical signals that, in animals, ...
s in the glabrous skin of humans:
Pacinian corpuscles Pacinian corpuscle or lamellar corpuscle or Vater-Pacini corpuscle; is one of the four major types of mechanoreceptors (specialized nerve ending with adventitious tissue for mechanical sensation) found in mammalian skin. This type of mechanorecept ...
,
Meissner's corpuscles Tactile corpuscles or Meissner's corpuscles are a type of mechanoreceptor discovered by anatomist Georg Meissner (1829–1905) and Rudolf Wagner. This corpuscle is a type of nerve ending in the skin that is responsible for sensitivity to pressu ...
, Merkel's discs, and
Ruffini corpuscles The Bulbous corpuscle or Ruffini ending or Ruffini corpuscle is a slowly adapting mechanoreceptor located in the cutaneous tissue between the dermal papillae and the hypodermis. It is named after Angelo Ruffini Angelo Ruffini (Pretare of Arquat ...
. The
naked mole-rat The naked mole-rat (''Heterocephalus glaber''), also known as the sand puppy, is a burrowing rodent native to the Horn of Africa and parts of Kenya, notably in Somali regions. It is closely related to the blesmols and is the only species in th ...
(''Heterocephalus glaber'') has evolved skin lacking in general, pelagic hair covering, yet has retained long, very sparsely scattered tactile hairs over its body. Glabrousness is a trait that may be associated with neoteny. The soft, fine hair found on many nonhuman mammals is typically called fur.


Human hairlessness


Evolutionary variation

Evolutionary biologists suggest that the genus '' Homo'' arose in
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
approximately 2.5 million years ago. They devised new hunting techniques. The higher protein diet led to the evolution of larger body and brain sizes. Jablonski postulates that increasing body size, in conjunction with intensified hunting during the day at the equator, gave rise to a greater need to rapidly expel heat. As a result, humans evolved the ability to sweat: a process which was facilitated by the loss of body hair. Another factor in human evolution that also occurred in the prehistoric past was a preferential selection for neoteny, particularly in females. The idea that adult humans exhibit certain neotenous (juvenile) features, not evinced in the other great apes, is about a century old. Louis Bolk made a long list of such traits, and
Stephen Jay Gould Stephen Jay Gould (; September 10, 1941 – May 20, 2002) was an American paleontologist, evolutionary biologist, and historian of science. He was one of the most influential and widely read authors of popular science of his generation. Gould sp ...
published a short list in '' Ontogeny and Phylogeny''. In addition, paedomorphic characteristics in women are often acknowledged as desirable by men in developed countries. For instance, vellus hair is a juvenile characteristic. However, while men develop longer, coarser, thicker, and darker terminal hair through sexual differentiation, women do not, leaving their vellus hair visible.


Texture


Curly hair

Jablonski asserts head hair was evolutionarily advantageous for pre-humans to retain because it protected the scalp as they walked upright in the intense African (equatorial)
UV light Ultraviolet (UV) is a form of electromagnetic radiation with wavelength from 10 nm (with a corresponding frequency around 30  PHz) to 400 nm (750  THz), shorter than that of visible light, but longer than X-rays. UV radiation i ...
. While some might argue that, by this logic, humans should also express hairy shoulders because these body parts would putatively be exposed to similar conditions, the protection of the head, the seat of the brain that enabled humanity to become one of the most successful species on the planet (and which also is very vulnerable at birth) was arguably a more urgent issue (axillary hair in the underarms and groin were also retained as signs of sexual maturity). Sometime during the gradual process by which ''Homo erectus'' began a transition from furry skin to the naked skin expressed by ''Homo sapiens'', hair texture putatively gradually changed from straight hair (the condition of most mammals, including humanity's closest cousins—chimpanzees) to Afro-textured hair or 'kinky' (i.e. tightly coiled). This argument assumes that curly hair better impedes the passage of UV light into the body relative to straight hair (thus curly or coiled hair would be particularly advantageous for light-skinned hominids living at the equator). It is substantiated by Iyengar's findings (1998) that UV light can enter into straight human hair roots (and thus into the body through the skin) via the hair shaft. Specifically, the results of that study suggest that this phenomenon resembles the passage of light through fiber optic tubes (which do not function as effectively when kinked or sharply curved or coiled). In this sense, when hominids (i.e. ''Homo erectus'') were gradually losing their straight body hair and thereby exposing the initially pale skin underneath their fur to the sun, straight hair would have been an adaptive liability. By inverse logic, later, as humans traveled farther from Africa and/or the equator, straight hair may have (initially) evolved to aid the entry of UV light into the body during the transition from dark, UV-protected skin to paler skin. Jablonski's assertions suggest that the adjective "woolly" in reference to Afro-hair is a
misnomer A misnomer is a name that is incorrectly or unsuitably applied. Misnomers often arise because something was named long before its correct nature was known, or because an earlier form of something has been replaced by a later form to which the name ...
in connoting the high heat insulation derivable from the true wool of sheep. Instead, the relatively sparse density of Afro-hair, combined with its springy coils actually results in an airy, almost sponge-like structure that in turn, Jablonski argues, more likely facilitates an increase in the circulation of cool air onto the scalp. Further, wet Afro-hair does not stick to the neck and scalp unless totally drenched and instead tends to retain its basic springy puffiness because it less easily responds to moisture and sweat than straight hair does. In this sense, the trait may enhance comfort levels in intense equatorial climates more than straight hair (which, on the other hand, tends to naturally fall over the ears and neck to a degree that provides slightly enhanced comfort levels in cold climates relative to tightly coiled hair). Further, it is notable that the most pervasive expression of this hair texture can be found in sub-Saharan Africa; a region of the world that abundant genetic and paleo-anthropological evidence suggests, was the relatively recent (≈200,000-year-old) point of origin for modern humanity. In fact, although genetic findings (Tishkoff, 2009) suggest that sub-Saharan Africans are the most genetically diverse continental group on Earth, Afro-textured hair approaches ubiquity in this region. This points to a strong, long-term selective pressure that, in stark contrast to most other regions of the genomes of sub-Saharan groups, left little room for genetic variation at the determining loci. Such a pattern, again, does not seem to support human sexual aesthetics as being the sole or primary cause of this distribution.


The EDAR locus

A group of studies have recently shown that genetic patterns at the EDAR locus, a region of the modern human genome that contributes to hair texture variation among most individuals of East Asian descent, support the hypothesis that (East Asian) straight hair likely developed in this branch of the modern human lineage subsequent to the original expression of tightly coiled natural afro-hair. Specifically, the relevant findings indicate that the EDAR mutation coding for the predominant East Asian 'coarse' or thick, straight hair texture arose within the past ≈65,000 years, which is a time frame that covers from the earliest of the 'Out of Africa' migrations up to now.


Disease

Ringworm is a fungal disease that targets hairy skin.
Premature greying of hair Premature greying of hair (PGH), also known as canities, can have negative effects on appearance, self-confidence, self-esteem, and social acceptance of the affected individual. Hair is said to have greyed prematurely if it occurs before the age of ...
is another condition that results in greying before the age of 20 years in Whites, before 25 years in Asians, and before 30 years in Africans.


Hair care

Hair care involves the hygiene and cosmetology of hair including hair on the scalp, facial hair ( beard and moustache),
pubic hair Pubic hair is terminal body hair that is found in the genital area of adolescent and adult humans. The hair is located on and around the sex organs and sometimes at the top of the inside of the thighs. In the pubic region around the pubis bon ...
and other body hair. Hair care routines differ according to an individual's culture and the physical characteristics of one's hair. Hair may be colored, trimmed, shaved, plucked, or otherwise removed with treatments such as waxing, sugaring, and threading.


Removal practices

Depilation is the removal of hair from the surface of the skin. This can be achieved through methods such as shaving.
Epilation Hair removal, also known as epilation or depilation, is the deliberate removal of body hair or head hair. Hair typically grows all over the human body and can vary in thickness and length across human populations. Hair can become more visible ...
is the removal of the entire hair strand, including the part of the hair that has not yet left the follicle. A popular way to epilate hair is through waxing.


Shaving

Shaving is accomplished with bladed instruments, such as razors. The blade is brought close to the skin and stroked over the hair in the desired area to cut the terminal hairs and leave the skin feeling smooth. Depending upon the rate of growth, one can begin to feel the hair growing back within hours of shaving. This is especially evident in men who develop a
five o'clock shadow Designer stubble is a facial hair style which is a short growth of beard, aimed to affect a rugged masculine or deliberately unkempt appearance. Initially made popular in the 1980s by singer George Michael and actor Don Johnson, the style la ...
after having shaved their faces. This new growth is called stubble. Stubble typically appears to grow back thicker because the shaved hairs are blunted instead of tapered off at the end, although the hair never actually grows back thicker.


Waxing

Waxing involves using a sticky wax and strip of paper or cloth to pull hair from the root. Waxing is the ideal hair removal technique to keep an area hair-free for long periods of time. It can take three to five weeks for waxed hair to begin to resurface again. Hair in areas that have been waxed consistently is known to grow back finer and thinner, especially compared to hair that has been shaved with a razor.


Laser removal

Laser hair removal is a cosmetic method where a small laser beam pulses selective heat on dark target matter in the area that causes hair growth without harming the skin tissue. This process is repeated several times over the course of many months to a couple of years with hair regrowing less frequently until it finally stops; this is used as a more permanent solution to waxing or shaving. Laser removal is practiced in many clinics along with many at-home products.


Cutting and trimming

Because the hair on one's head is normally longer than other types of body hair, it is cut with
scissors Scissors are hand-operated shearing tools. A pair of scissors consists of a pair of metal blades pivoted so that the sharpened edges slide against each other when the handles (bows) opposite to the pivot are closed. Scissors are used for cutti ...
or clippers. People with longer hair will most often use scissors to cut their hair, whereas shorter hair is maintained using a trimmer. Depending on the desired length and overall health of the hair, periods without cutting or trimming the hair can vary. Cut hair may be used in wigs. Global imports of hair in 2010 was worth $US 1.24 billion.


Social role

Hair has great social significance for
human being Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedality, bipedalism and exceptional cognitive skills due to a large and complex Human brain, brain. This has enabled the development of ad ...
s. It can grow on most external areas of the
human body The human body is the structure of a Human, human being. It is composed of many different types of Cell (biology), cells that together create Tissue (biology), tissues and subsequently organ systems. They ensure homeostasis and the life, viabi ...
, except on the palms of the
hand A hand is a prehensile, multi-fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the koala (which has two opposable thumbs on each "h ...
s and the soles of the feet (among other areas). Hair is most noticeable on most people in a small number of areas, which are also the ones that are most commonly trimmed,
plucked ''Death Laid an Egg'' ( it, La morte ha fatto l'uovo) is a 1968 ''giallo'' film directed by Giulio Questi. Written by Questi and Franco Arcalli, the film stars Ewa Aulin, Gina Lollobrigida and Jean-Louis Trintignant. Plot Married couple Anna an ...
, or shaved. These include the face, ears,
head A head is the part of an organism which usually includes the ears, brain, forehead, cheeks, chin, eyes, nose, and mouth, each of which aid in various sensory functions such as sight, hearing, smell, and taste. Some very simple animals may ...
, eyebrows, legs, and armpits, as well as the
pubic region The hypogastrium (also called the hypogastric region or suprapubic region) is a region of the abdomen located below the umbilical region. Etymology The roots of the word ''hypogastrium'' mean "below the stomach"; the roots of ''suprapubic'' mean ...
. The highly visible differences between male and female body and facial hair are a notable secondary sex characteristic. The world's longest documented hair belongs to Xie Qiuping (in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
), at 5.627 m (18 ft 5.54 in) when measured on 8 May 2004. She has been growing her hair since 1973, from the age of 13.


Indication of status

Healthy hair indicates health and youth (important in evolutionary biology). Hair color and texture can be a sign of ethnic ancestry.
Facial hair Facial hair is hair grown on the face, usually on the chin, cheeks, and upper lip region. It is typically a secondary sex characteristic of human males. Men typically start developing facial hair in the later stages of puberty or adolescence, ...
is a sign of puberty in men. White or gray hair is a sign of age or genetics, which may be concealed with
hair dye Hair coloring, or hair dyeing, is the practice of changing the hair color. The main reasons for this are cosmetic: to cover gray or white hair, to change to a color regarded as more fashionable or desirable, or to restore the original hair color ...
(not easily for some), although many prefer to assume it (especially if it is a poliosis characteristic of the person since childhood). Male pattern
baldness Hair loss, also known as alopecia or baldness, refers to a loss of hair from part of the head or body. Typically at least the head is involved. The severity of hair loss can vary from a small area to the entire body. Inflammation or scarrin ...
is a sign of age, which may be concealed with a toupee, hats, or religious and cultural adornments. Although drugs and medical procedures exist for the treatment of baldness, many balding men simply shave their heads. In early modern China, the queue was a male hairstyle worn by the Manchus from central Manchuria and the Han Chinese during the Qing dynasty; hair on the front of the head was shaved off above the temples every ten days, mimicking male-pattern baldness, and the rest of the hair braided into a long pigtail. Hairstyle may be an indicator of group membership. During the English Civil War, the followers of Oliver Cromwell decided to crop their hair close to their head, as an act of defiance to the curls and ringlets of the king's men. This led to the Parliamentary faction being nicknamed Roundheads. Recent isotopic analysis of hair is helping to shed further light on sociocultural interaction, giving information on food procurement and consumption in the 19th century. Having bobbed hair was popular among the flappers in the 1920s as a sign of rebellion against traditional roles for women. Female art students known as the "cropheads" also adopted the style, notably at the Slade School in London, England. Regional variations in hirsutism cause practices regarding hair on the arms and legs to differ. Some religious groups may follow certain rules regarding hair as part of religious observance. The rules often differ for men and women. Many subcultures have hairstyles which may indicate an unofficial membership. Many
hippie A hippie, also spelled hippy, especially in British English, is someone associated with the counterculture of the 1960s, originally a youth movement that began in the United States during the mid-1960s and spread to different countries around ...
s,
metalheads A metalhead is a member of the heavy metal subculture. Metalhead may also refer to: Music * ''Metalhead'' (album), 1999 album by Saxon * "Metalhead", a song by Blotto * "Metalhead", a song from the 2008 Miss Kittin album ''BatBox'' * Metalheadz ...
, and Indian sadhus have long hair, as well many older indie kids. Many
punks Punk or punks may refer to: Genres, subculture, and related aspects * Punk rock, a music genre originating in the 1970s associated with various subgenres * Punk subculture, a subculture associated with punk rock, or aspects of the subculture s ...
wear a hairstyle known as a mohawk or other spiked and dyed hairstyles; skinheads have short-cropped or completely shaved heads. Long stylized bangs were very common for
emo Emo is a rock music genre characterized by emotional, often confessional lyrics. It emerged as a style of and hardcore punk from the Washington D.C. hardcore punk scene, where it was known as emotional hardcore or emocore and pioneered b ...
s,
scene kids The scene subculture is a youth subculture that emerged during the early 2000s in the United States from the pre-existing emo subculture. The subculture became popular with adolescents from the mid-2000s to early 2010s. Members of the scene subc ...
and younger indie kids in the 2000s and early 2010s, among people of both genders. Heads were shaved in concentration camps, and head-shaving has been used as
punishment Punishment, commonly, is the imposition of an undesirable or unpleasant outcome upon a group or individual, meted out by an authority—in contexts ranging from child discipline to criminal law—as a response and deterrent to a particular acti ...
, especially for women with long hair. The shaven head is common in military haircuts, while Western monks are known for the tonsure. By contrast, among some Indian holy men, the hair is worn extremely long. In the time of Confucius (5th century BCE), the Chinese grew out their hair and often tied it, as a symbol of filial piety. Regular hairdressing in some cultures is considered a sign of wealth or status. The dreadlocks of the Rastafari movement were despised early in the movement's history. In some cultures, having one's hair cut can symbolize a liberation from one's past, usually after a trying time in one's life. Cutting the hair also may be a sign of mourning. Tightly coiled hair in its natural state may be worn in an Afro. This hairstyle was once worn among African Americans as a symbol of racial pride. Given that the coiled texture is the natural state of some African Americans' hair, or perceived as being more "African", this simple style is now often seen as a sign of self-acceptance and an affirmation that the beauty norms of the ( eurocentric) dominant culture are not absolute. It is important to note that African Americans as a whole have a variety of hair textures, as they are not an ethnically homogeneous group, but an ad-hoc of different racial admixtures. The film '' Easy Rider'' (1969) includes the assumption that the two main characters could have their long hairs forcibly shaved with a rusty razor when jailed, symbolizing the intolerance of some conservative groups toward members of the counterculture. At the conclusion of the Oz obscenity trials in the UK in 1971, the defendants had their heads shaved by the police, causing public outcry. During the appeal trial, they appeared in the dock wearing wigs. A case where a 14-year-old student was expelled from school in Brazil in the mid-2000s, allegedly because of his fauxhawk haircut, sparked national debate and legal action resulting in compensation.


Religious practices

Women's hair may be hidden using
headscarves A headscarf is a scarf covering most or all of the top of a person's, usually women's, hair and head, leaving the face uncovered. A headscarf is formed of a triangular cloth or a square cloth folded into a triangle, with which the head is cov ...
, a common part of the ''
hijab In modern usage, hijab ( ar, حجاب, translit=ḥijāb, ) generally refers to headcoverings worn by Muslim women. Many Muslims believe it is obligatory for every female Muslim who has reached the age of puberty to wear a head covering. While ...
'' in
Islam Islam (; ar, ۘالِإسلَام, , ) is an Abrahamic religions, Abrahamic Monotheism#Islam, monotheistic religion centred primarily around the Quran, a religious text considered by Muslims to be the direct word of God in Islam, God (or ...
and a symbol of modesty required for certain religious rituals in Eastern Orthodoxy. Russian Orthodox Church requires all married women to wear headscarves inside the church; this tradition is often extended to all women, regardless of marital status. Orthodox Judaism also commands the use of scarves and other head coverings for married women for modesty reasons. Certain
Hindu Hindus (; ) are people who religiously adhere to Hinduism.Jeffery D. Long (2007), A Vision for Hinduism, IB Tauris, , pages 35–37 Historically, the term has also been used as a geographical, cultural, and later religious identifier for ...
sects also wear head scarves for religious reasons.
Sikhs Sikhs ( or ; pa, ਸਿੱਖ, ' ) are people who adhere to Sikhism (Sikhi), a monotheistic religion that originated in the late 15th century in the Punjab region of the Indian subcontinent, based on the revelation of Guru Nanak. The term ...
have an obligation not to cut hair (a Sikh cutting hair becomes 'apostate' which means fallen from religion) and men keep it tied in a bun on the head, which is then covered appropriately using a turban. Multiple religions, both ancient and contemporary, require or advise one to allow their hair to become dreadlocks, though people also wear them for fashion. For men, Islam, Orthodox Judaism, Orthodox Christianity, Roman Catholicism, and other religious groups have at various times recommended or required the covering of the head and sections of the hair of men, and some have dictates relating to the cutting of men's facial and head hair. Some Christian sects throughout history and up to modern times have also religiously proscribed the cutting of women's hair. For some
Sunni Sunni Islam () is the largest branch of Islam, followed by 85–90% of the world's Muslims. Its name comes from the word '' Sunnah'', referring to the tradition of Muhammad. The differences between Sunni and Shia Muslims arose from a disagr ...
madhabs, the donning of a kufi or
topi ''Damaliscus lunatus jimela'' is a subspecies of topi, and is usually just called a topi. It is a highly social and fast type of antelope found in the savannas, semi-deserts, and floodplains of sub-Saharan Africa. Names The word ''tope'' or '' ...
is a form of
sunnah In Islam, , also spelled ( ar, سنة), are the traditions and practices of the Islamic prophet Muhammad that constitute a model for Muslims to follow. The sunnah is what all the Muslims of Muhammad's time evidently saw and followed and passed ...
.The War Within Our Hearts – Page 65 Sa'ad Quadri – 2013


In Arabic poetry

Since ancient times, women's long, thick, wavy hair has featured prominently in
Arabic poetry Arabic poetry ( ar, الشعر العربي ''ash-shi‘ru al-‘Arabīyyu'') is the earliest form of Arabic literature. Present knowledge of poetry in Arabic dates from the 6th century, but oral poetry is believed to predate that. Arabic poetry ...
. Pre-Islamic poets used only limited imagery to describe women's hair. For example, al-A'sha wrote a verse comparing a lover's hair to "a garden whose grapes dangle down upon me", but Bashar ibn Burd considered this unusual. One comparison used by early poets, such as
Imru al-Qays Imruʾ al-Qais Junduh bin Hujr al-Kindi ( ar, ٱمْرُؤ ٱلْقَيْس جُنْدُح ٱبْن حُجْر ٱلْكِنْدِيّ, ALA-LC: ''ʾImruʾ al-Qays Junduḥ ibn Ḥujr al-Kindīy'') was an Arab king and poet in the 6th century, an ...
, was to bunches of dates. In Abbasid times, however, the imagery for hair expanded significantly - particularly for the then-fashionable "love-locks" (''sudgh'') framing the temples, which came into style at the court of the caliph al-Amin. Hair curls were compared to hooks and chains, letters (such as '' fa'', ''
waw Waw or WAW may refer to: * Waw (letter), a letter in many Semitic abjads * Waw, the velomobile * Another spelling for the town Wau, South Sudan * Waw Township, Burma *Warsaw Chopin Airport, an international airport serving Warsaw, Poland (IATA ai ...
'', ''
lam Lam or LAM may refer to: Organizations * Laguna Art Museum, California, US * Lam Eng Rubber, a Malaysian manufacturer * Lam Research, American semiconductor equipment company * LAM Mozambique Airlines, flag carrier airline of Mozambique * Librar ...
'', and '' nun''), scorpions,
annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
s, and
polo stick Polo is a ball game played on horseback, a traditional field sport and one of the world's oldest known team sports. The game is played by two opposing teams with the objective of scoring using a long-handled wooden mallet to hit a small hard ...
s. An example was the poet Ibn al-Mu'tazz, who compared a lock of hair and a birthmark to a polo stick driving a ball.


See also

*
Body hair Body hair, or androgenic hair, is the terminal hair that develops on the human body during and after puberty. It is differentiated from the head hair and less visible vellus hair, which is much finer and lighter in color. The growth of androge ...
*
Chaetophobia The English language, English suffixes -phobia, -phobic, -phobe (from Ancient Greek, Greek φόβος ''phobos'', "fear") occur in technical usage in psychiatry to construct words that describe irrational, abnormal, unwarranted, persistent, or ...
– the fear of hair * Hair analysis (alternative medicine) * Hypertrichosis – the state of having an excess of hair on the head or body *
Hypotrichosis Hair loss, also known as alopecia or baldness, refers to a loss of hair from part of the head or body. Typically at least the head is involved. The severity of hair loss can vary from a small area to the entire body. Inflammation or scarri ...
– the state of having a less than normal amount of hair on the head or body *
Lanugo Lanugo is very thin, soft, usually unpigmented, downy hair that is sometimes found on the body of a fetus or newborn. It is the first hair to be produced by the fetal hair follicles, and it usually appears around sixteen weeks of gestation and is ...
* Seta – hair-like structures in insects *
Bristle sensilla Bristle sensilla (sometimes also called tactile hairs) are a class of mechanoreceptors found in insects and other arthropods that respond to mechanical stimuli generated by the external world. As a result, they are considered exteroceptors. Bris ...
– tactile hairs in insects *
Trichotillomania Trichotillomania (TTM), also known as hair-pulling disorder or compulsive hair pulling, is a mental disorder characterized by a long-term urge that results in the pulling out of one's own hair. A brief positive feeling may occur as hair is remov ...
– hair pulling


References


Citations


Sources

* * * *


External links

* *
How to measure the diameter of your own hair using a laser pointerInstant insight
outlining the chemistry of hair from the Royal Society of Chemistry * {{Authority control