''Gulosaurus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of basal

grippidia Ichthyopterygia ("fish flippers") was a designation introduced by Sir Richard Owen in 1840 to designate the Jurassic ichthyosaurs that were known at the time, but the term is now used more often for both true Ichthyosauria and their more primiti ...

ichthyopterygia Ichthyopterygia ("fish flippers") was a designation introduced by Sir Richard Owen in 1840 to designate the Jurassic ichthyosaurs that were known at the time, but the term is now used more often for both true Ichthyosauria and their more primitiv ...

n known from the

Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a un ...

Vega-Phroso Siltstone Member of the

Sulphur Mountain Formation The Sulphur Mountain Formation is a geologic formation of Early to Middle Triassic age. It is present on the western edge of the Western Canada Sedimentary Basin in the foothills and Rocky Mountains of western Alberta and northeastern British Col ...

of east-central

British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

. ''Gulosaurus'' was first named by Robin S. Cuthbertson, Anthony P. Russell and Jason S. Anderson in

2013 File:2013 Events Collage V2.png, From left, clockwise: Edward Snowden becomes internationally famous for leaking classified NSA wiretapping information; Typhoon Haiyan kills over 6,000 in the Philippines and Southeast Asia; The Dhaka garment fact ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Gulosaurus helmi''. The name means 'Helm's

wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for "gluttony, glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is ...

lizard' and refers to the Wolverine Nordic and Mountain Society, who maintain the area around

Wapiti Lake The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The comm ...

where it was found, and to Dr

Charles Helm Charles Daniel Helm (28 September 1844 – 14 September 1915) was a Protestant missionary and trusted confident of King Lobengula of Matabeleland who played a controversial role as an interpreter during the drafting and signing of the Rudd Concess ...

, who is a leading paleontologist around this same area. Originally ''Gulosaurus'' was thought to be either ''

Grippia longirostris ''Grippia'' is a genus of early ichthyopterygian, an extinct group of reptiles that resembled dolphins. Its only species is ''Grippia longirostris''. It was a relatively small ichthyopterygian, measuring long and weighing . Fossil remains from ...

'' or ''

Parvinatator wapitiensis ''Parvinatator'', from Latin, “parvus” little and “natator” swimmer, is an extinct genus of small ichthyopterygian marine reptile that lived during the Early to Middle Triassic. Its fossils have been found in British Columbia, Canada.Ni ...

,'' but as the fossil was incomplete it was uncertain which. In 2013, the other half of the fossil, including its skull, was discovered and this proved it to be a new species entirely, closely related to ''Grippia'' and very similar.

Features

Overall, ''Gulosaurus'' was similar to ''Grippia.'' It was the same size, around 1.5 meters long, and much of the skeleton was the same, namely the ribs and centrum discovered. Also like ''Grippia'', it had pentadactyl limbs rather than exhibiting polyphalangy, as many later ichthyopterygians did. Most of the flipper structure was very similar apart from the first metacarpal, which had a notched peripheral shaft in ''Gulosaurus'' but was lunate in ''Grippia.'' The skull of ''Gulosaurus'' was also very different to that of ''Grippia'' - it had cylindrical anterior teeth, although as no ''Grippia'' snouts have been discovered, it remains unknown if this was different. The skull also exhibited

postfrontal The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

-frontal contact at the

rostral Rostral may refer to: Anatomy * Rostral (anatomical term), situated toward the oral or nasal region * Rostral bone, in ceratopsian dinosaurs * Rostral organ, of certain fish * Rostral scale, in snakes and scaled reptiles Other uses * Rostral colu ...

limit of the

anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...

margin of the

supratemporal The supratemporal bone is a paired cranial bone present in many tetrapods and tetrapodomorph fish. It is part of the temporal region (the portion of the skull roof behind the eyes), usually lying medial (inwards) relative to the squamosal and latera ...

terrace.

Phylogeny

Below is a

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

modified from Cuthbertson ''et al.'', 2013.

References

Triassic ichthyopterygians Fossil taxa described in 2013 Fossils of British Columbia Early Triassic reptiles of North America Ichthyosauromorph genera {{Ichthyosaur-stub

Features

Phylogeny

See also

References