Great Sapphirewing
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The great sapphirewing (''Pterophanes cyanopterus'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
in the "brilliants", tribe
Heliantheini Heliantheini is one of the two tribes that make up the subfamily Lesbiinae of the hummingbird family Trochilidae. The other tribe in the subfamily is Lesbiini. The informal name "brilliants" has been proposed for this group as it includes the ge ...
in subfamily Lesbiinae. It is found in
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
, and
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved 27 May 2021


Taxonomy and systematics

The great sapphirewing is the only member of its genus. In the late 1900s it was suggested that ''Pterophanes'' be merged into genus ''Coeligena'', but later work showed that those genera are not closely related. The species has three subspecies, the nominate ''P. c. cyanopterus'', ''P. c. caeruleus'', and ''P. c. peruvianus''. The great sapphirewing has also been called Temminck's sapphirewing, páramo sapphirewing, and simply sapphirewing.Fernández, C. M., D. Luther, I. Heynen, P. F. D. Boesman, and G. M. Kirwan (2021). Great Sapphirewing (''Pterophanes cyanopterus''), version 2.0. In Birds of the World (T. S. Schulenberg and B. K. Keeney, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.gresap1.02 retrieved 3 May 2022


Description

The great sapphirewing is one of the largest hummingbirds; only the two of genus '' Topaza'' and the giant hummingbird (''Patagona gigas'') are larger. It is long including the bill. Males weigh between and females . Both sexes have a small white spot behind the eye. Males of the nominate subspecies have dark shining blue-green upperparts. The wing is mostly shining blue and the tail greenish black. The underparts are also blue-green but somewhat bluer than the upperparts. Nominate females have mostly dark metallic green upperparts with a dusky gray crown. Only the wing
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
are blue; the rest of the wing is dusky. The tail is mostly greenish black with much white on the outermost feathers. The underparts are cinnamon that is mixed with green on the sides. Males of subspecies ''P. c. caeruleus'' have even more and darker blue than the nominate. Females also have more blue, a darker crown, and less white on the outer tail feathers. Males of ''P. c. peruvianus'' have more greenish than blue upperparts than the nominate, though the wings have about the same amount of blue. The underparts are also greenish, with a buffy belly. The female's underparts are a less intense cinnamon than the nominate's and it has more white on the outer tail feathers.


Distribution and habitat

The nominate subspecies of great sapphirewing is found in Colombia's Eastern Andes, in the departments of
Norte de Santander North Santander (Spanish: Norte de Santander) () is a department of Northeastern Colombia. It is in the north of the country, bordering Venezuela. Its capital is Cúcuta, one of the country's major cities. North Santander is bordered by Venez ...
and Cundinamarca, and slightly into adjoining far western Venezuela. ''P. c. caeruleus'' is found in Colombia's Central and Western Andes. ''P. c. peruvianus'' is the most widespread; it is found from the Western Andes of Colombia south through Ecuador and Peru to Cochabamba Department in central Bolivia. The great sapphirewing inhabits the edges of humid
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
and
elfin forest Dwarf forest, elfin forest, or pygmy forest is an uncommon ecosystem featuring miniature trees, inhabited by small species of fauna such as rodents and lizards. They are usually located at high elevations, under conditions of sufficient air humi ...
and shrubby slopes with scattered trees. It also occurs well into the ''
páramo Páramo () can refer to a variety of alpine tundra ecosystems located in the Andes Mountain Range, South America. Some ecologists describe the páramo broadly as "all high, tropical, montane vegetation above the continuous timberline". A narrower ...
'', at least seasonally. In Colombia it occurs between , in Ecuador between , in Peru between , and in Bolivia mostly above .


Behavior


Movement

The great sapphirewing is mostly sedentary, but locally some move to lower elevations during the austral winter.


Feeding

The great sapphirewing feeds on the nectar of a wide variety of flowering plants, most of which are fairly small though some are larger. In Peru it seems to favor ''
Tristerix longebracteatus ''Tristerix longebracteatus'' is a species of ''Tristerix'' found in Colombia and Ecuador References External links * * longebracteatus Flora of the Andes {{Santalales-stub ...
'' and in the Colombian ''páramo'' ''
Puya clava-herculis ''Puya clava-herculis'' is a species of plants in the genus ''Puya''. This species is native to Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuad ...
''. It typically feeds at low to mid-levels of the vegetation, hovering or perching at the outside of the host plant. It usually defends patches of flowers but also forages by
trap-lining In ethology and behavioral ecology, trap-lining or traplining is a feeding strategy in which an individual visits food sources on a regular, repeatable sequence, much as trappers check their lines of traps. Traplining is usually seen in species ...
, visiting a circuit of feeding sites. In addition to feeding on nectar it captures small
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s by gleaning from foliage and by
hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist *Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Haw ...
.


Breeding

The great sapphirewing's breeding season is not well defined but includes at least February to May. The only known nest was an oddity, a three-compartment structure of moss lined with soft fibers. When it was discovered, a female sapphirewing was brooding two nestlings in one compartment and another was occupied by a female
tyrian metaltail The Tyrian metaltail (''Metallura tyrianthina'') is a species of hummingbird in the subfamily Lesbiinae, the brilliants and coquettes. It is found in Bolivia, Colombia, Ecuador, Peru, and Venezuela.HBW and BirdLife International (2020) ''Handbo ...
(''Metallura tyrianthina''). The third compartment was empty. The nest was about above ground, suspended from roots below vegetation.


Vocalization

The great sapphirewing's vocalizations include "a high, thin, liquid chatter", a "drawn-out piercing high-pitched ''zeee''", and " nagitated ''titititirrr''".


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the great sapphirewing as being of Least Concern. It has a large range and though its population size is unknown, it is believed to be stable. No specific threats have been identified. It is variously considered rare to locally common in different parts of its range. "It is sensitive to forest loss or fragmentation but, as a species that tolerates forest edges, it is less vulnerable than are many other species of montane forests."


References

{{Taxonbar, from=Q649173 Trochilinae Hummingbird species of South America Birds of the Northern Andes Birds described in 1839 Taxa named by Louis Fraser Taxonomy articles created by Polbot