Grammitid

Grammitidoideae is a subfamily of the fern family Polypodiaceae, whose members are informally known as grammitids. It comprises a clade of about 750 species. They are distributed over higher elevations in both the Old and New World. This group was previously treated as a separate family, Grammitidaceae until molecular phylogenies showed it to be nested within the Polypodiaceae. It has since been treated as an unranked clade within subfamily Polypodioideae (renamed tribe Polypodieae in one classification), and, most recently, as a separate subfamily (reducing Polypodioideae to an evolutionary grade).

Circumscription

In 2011, Christenhusz ''et al.'' placed the grammitid ferns in the subfamily Polypodioideae, within the Polypodiaceae, as an informal group without rank. In 2014, Christenhusz and Chase expanded the circumscription of both family and subfamily, placing the Polypodioideae as previously delimited, including grammitids, in tribe Polypodieae. The PPG I classification of 2016 largely reverted to the 2011 circumscriptions of these groups, but placed the grammitids in a new subfamily.

The following phylogram shows a likely relationship between grammitid ferns and the Polypodiaceae subfamilies, based on Schuettpelz & Pryer, 2008, with PPG I subfamilies shown.

Genera

The Pteridophyte Phylogeny Group classification of 2016 (PPG I) recognizes the following genera:

*''Acrosorus'' Copel. – Southeastern Asia
*''Adenophorus'' Gaudich. – Hawaii)
*''Alansmia'' M.Kessler, Moguel, Sundue & Labiak – Neotropics, Africa
*''Archigrammitis'' Parris – Malesia, Polynesia
*'' Ascogrammitis'' Sundue – Neotropics
*''Calymmodon'' C.Presl – Southeastern Asia
*''Ceradenia'' L.E.Bishop – mainly tropical Africa
*'' Chrysogrammitis'' Parris – Southeastern Asia
*''Cochlidium'' Kaulf. (including ''Xiphopteris'' Kaulf.) – Neotropics, Taiwan
*''Ctenopterella'' Parris – Southeastern Asia, Australia
*'' Dasygrammitis'' Parris – Southeastern Asia
*''Enterosora'' Baker – Neotropics, Madagascar
*'' Galactodenia'' Sundue & Labiak – Neotropics
*''Grammitis'' Sw. – Neotropics
*'' Lellingeria'' A.R.Sm. & R.C.Moran – Neotropics
*'' Leucotrichum'' Labiak – Neotropics, Madagascar
*'' Lomaphlebia'' J.Sm.
*'' Luisma'' M.T.Murillo & A.R.Sm. – Colombia
*''Melpomene'' A.R.Sm. & R.C.Moran – Neotropics
*''Micropolypodium'' Hayata – China, Japan
*'' Moranopteris'' R.Y.Hirai & J.Prado – Neotropics
*'' Mycopteris'' Sundue – Neotropics
*'' Notogrammitis'' Parris – Australia, New Zealand
*'' Oreogrammitis'' Copeland – Southeastern Asia, Australia, Pacific islands, China, Philippines
*'' Prosaptia'' C.Presl (including ''Ctenopteris'' Blume ex Kunze) – Southeastern Asia
*'' Radiogrammitis'' Parris – Southeastern Asia, Pacific islands; sunk into '' Oreogrammitis'' in 2020
*'' Scleroglossum'' Alderwerelt (including ''Nematopteris'' Alderw.) – Southeastern Asia, Pacific islands
*'' Stenogrammitis'' Labiak – Neotropics, Africa, Pacific islands
*''Terpsichore'' A.R.Sm. – Neotropics
*'' Themelium'' (T.Moore) Parris – China, Philippines, Thailand; sunk into '' Oreogrammitis'' in 2020
*'' Tomophyllum'' (E.Fournier) Parris – China, Inda, Nepal
*'' Xiphopterella'' Parris – China
*''Zygophlebia'' L.E.Bishop – Neotropics, Madagascar; sunk into ''Enterosora'' in 2019

References

{{Taxonbar, from=Q5593710

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Plant subfamilies